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THE GENUS REBUTIA<br />

1895—1981<br />

REBUTIA MUSCULA<br />

No. 2 of a series of illustrated guide books by:-<br />

Brian Fearn and Leslie Pearcy<br />

Published by Abbey Brook Cactus Nursery


Published by: Abbey Brook Cactus Nursery,<br />

Old Hackney Lane,<br />

Matlock, Derbyshire.<br />

Copyright © ABCN 1981<br />

Illustrations: All illustrations are by Brian Fearn<br />

Printed in Great Britain by:<br />

Higham Press Ltd., Shirland, Derby., DE5 6BP


A<br />

THE GENUS REBUTIA<br />

1895-1981<br />

An up to date check list of <strong>the</strong> <strong>genus</strong> Rebutia<br />

and related groups including Aylostera,<br />

Mediolobivia and Digito<strong>rebutia</strong> toge<strong>the</strong>r with<br />

photographs and descriptions of 20 of <strong>the</strong><br />

more notable species.<br />

No. 2 of a series of illustrated guide books by:-<br />

Brian Fearn and Leslie Pearcy<br />

Published by ABBEY BROOK CACTUS NURSERY<br />

1


Contents<br />

Preface . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 3<br />

Introduction . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 4<br />

Nomenclature and Classification . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 5<br />

Analytical key to <strong>the</strong> Rebutia Sections . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 7<br />

Flowers and floral structure . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 8<br />

Geographical Distribution . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 13<br />

Distribution maps and locational data . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 15<br />

Cultivation . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 23<br />

Pests and Diseases . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 26<br />

Descriptions - arranged in alphabetical order 27<br />

Check list of Genera, Subgenera and Sections . . . . . . . . . . . . . . . . . . . . . . . . 52<br />

Check list of Aylostera Speg. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 53<br />

Check list of Digito<strong>rebutia</strong> Frič & Kreuzgr. ex Buin.. . . . . . . . . . . . . . . . . . . . 54<br />

Check list of Mediolobivia Backbg. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 55<br />

Check list of Genus Rebutia including specific, varietal and name<br />

combinations which have been used in connection with <strong>the</strong> <strong>genus</strong> . . . . . 59<br />

Friedrich Ritter field numbers . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 71<br />

Alfred Lau field numbers. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 73<br />

Walter Rausch field numbers . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 74<br />

Karel Knize field numbers . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 77<br />

Bibliography . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 78<br />

Acknowledgements . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 79<br />

Glossary . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 79<br />

2


Preface<br />

‘The Genus Rebutia 1895-1981’ is <strong>the</strong> second in a planned series of publications<br />

by Abbey Brook Cactus Nursery. The first booklet ‘50 Choice Mammillarias’ has<br />

been well received and <strong>the</strong> 1st edition has sold well and is now out of print. Two<br />

years have passed since its introduction and we apologise for <strong>the</strong> delay in <strong>the</strong><br />

production of <strong>the</strong> 2nd booklet. The original intention was to produce a new booklet<br />

each year. In <strong>the</strong> meantime ano<strong>the</strong>r booklet ‘Lithops’ by Brian Fearn has been<br />

published by <strong>the</strong> National Cactus and Succulent Society and so <strong>the</strong> new booklet is<br />

really <strong>the</strong> 3rd that has been produced by Abbey Brook. We are thus still on target.<br />

There seems to be no end to <strong>the</strong> publication of new general books on Cacti and<br />

o<strong>the</strong>r Succulent plants but our intention of producing inexpensive booklets with<br />

detailed information for <strong>the</strong> serious collector is as relevant now as it was when<br />

booklet No. 1 was published.<br />

Brian Fearn and Leslie Pearcy<br />

Matlock 1981<br />

REBUTIA FLAVISTYLA<br />

Fig. 1 A superb species discovered by Friedrich Ritter and described in 1978<br />

(FR756). Beautiful light orange flowers freely produced.<br />

3


Introduction<br />

The <strong>genus</strong> Rebutia is one of <strong>the</strong> most widely cultivated groups of plants which<br />

belong to <strong>the</strong> family Cactaceae. Their ease of cultivation and free flowering nature<br />

endears <strong>the</strong>m to all cactophiles. They are plants of <strong>the</strong> Andean mountain chain in<br />

South America occurring in nor<strong>the</strong>rn Argentina and Bolivia. One of <strong>the</strong> reasons for<br />

<strong>the</strong>ir popularity is <strong>the</strong>ir ease of cultivation and adaptability to neglect. They do not<br />

exactly thrive on neglect, but never<strong>the</strong>less <strong>the</strong>y will still produce <strong>the</strong>ir beautiful<br />

flowers in <strong>the</strong> spring, provided cool, dry conditions have been given during <strong>the</strong><br />

winter months. As <strong>the</strong>se are plants from <strong>the</strong> high Andes, <strong>the</strong>y have <strong>the</strong> added ability<br />

to survive in an unheated greenhouse or porch when winter temperatures could fall<br />

to well below 0°C (32°F).<br />

Nearly all <strong>the</strong> species are small growing plants, rarely needing more than a 3½''<br />

pot, with a range of body form from globular to short cylindrical stems. The spination<br />

is very varied and often very colourful. Some plants have snow white spines, o<strong>the</strong>rs<br />

golden yellow. The flowers are often very large for <strong>the</strong> size of <strong>the</strong> plant. In <strong>the</strong><br />

majority of cases <strong>the</strong> flower colours are vivid oranges, reds, yellows and purple but<br />

occasionally bicoloured flowers are produced. There are also a few white flowered<br />

species. These plants are of great horticultural value and no collection of cacti is<br />

complete without some representatives of this <strong>genus</strong>.<br />

REBUTIA FUSCA<br />

Fig. 2 Described by Friednch Ritter in 1977 (FR940). It produces beautiful dark red<br />

flowers and is probably closely related to R. spegazziniana Backbg.<br />

4


Nomenclature and Classification<br />

It is doubtful if two people will ever agree on <strong>the</strong> taxonomy of <strong>the</strong> <strong>genus</strong> Rebutia<br />

and as a result <strong>the</strong> nomenclature is a veritable minefield. This <strong>genus</strong> has suffered<br />

more than most from <strong>the</strong> over-enthusiastic naming of new species, revisions, and<br />

changes in names and status. This has resulted in a bewildering array of possible<br />

names and combinations of names. To be quite honest, far too many Rebutia species<br />

have been described because of <strong>the</strong> demand and commercial pressures.<br />

Consequently <strong>the</strong>re is considerable confusion in this <strong>genus</strong> in trying to establish<br />

clear relationships between <strong>the</strong> many taxa. The result of all this activity has been an<br />

unwarranted cluttering of <strong>the</strong> synonymy of <strong>the</strong> group. Over 800 possible names and<br />

combinations have been found in a search of <strong>the</strong> literature and we are sure that <strong>the</strong>re<br />

are many more that we have not found. One really begins to wonder when Rebutia<br />

beryllioides has been described as a species, as a variety of 2 different species, a<br />

subspecies and finally as a form of a third species. Is it any wonder that professional<br />

taxonomists rarely work on a group of plants that are of horticultural value? With<br />

apologies to <strong>the</strong> late Professor C. D. Darlington <strong>the</strong> following misquotation springs to<br />

mind ‘cactus taxonomy is <strong>the</strong> pursuit of <strong>the</strong> impossible by <strong>the</strong> incompetent’.<br />

We believe that <strong>the</strong> main cause of this confusion is that <strong>the</strong> <strong>genus</strong> Rebutia s. l. is in<br />

an active state of evolution. This possibility does not appear to have been grasped by<br />

<strong>the</strong> many individuals who have worked on <strong>the</strong> <strong>genus</strong>. It would account for <strong>the</strong> fact that<br />

although on paper <strong>the</strong>re appear to be 4 main sections which can fur<strong>the</strong>r be<br />

subdivided into 8 or more subsections, <strong>the</strong>re are never<strong>the</strong>less some plants which<br />

cross <strong>the</strong> boundaries, having characteristics which are intermediate between two or<br />

more sections or subsections. The knowledge of <strong>the</strong> existence of <strong>the</strong>se plants is<br />

largely due to <strong>the</strong> field work of Friedrich Ritter who has found and introduced a large<br />

number of new plants, some of which have intermediate characters. This makes <strong>the</strong><br />

reduction from a multigeneric group to a single <strong>genus</strong> desirable. A fur<strong>the</strong>r<br />

complication is <strong>the</strong> fact that many species are self fertile and inbreeding, so that <strong>the</strong><br />

occurrence of local races and inbred lines is highly likely.<br />

The activities in <strong>the</strong> field by Ritter, Rausch, Lau, Knize, Vasques and o<strong>the</strong>rs have<br />

all helped in <strong>the</strong> proliferation of names. Although not all are to blame for <strong>the</strong><br />

subsequent naming of <strong>the</strong>ir introductions, <strong>the</strong>y have all collected in <strong>the</strong> same areas,<br />

often following in each o<strong>the</strong>rs footsteps. The consequence of this has been <strong>the</strong><br />

introduction of many ‘new’ species with <strong>the</strong> great possibility that <strong>the</strong>re has been<br />

considerable duplication. Every hilltop or slope seems to be inhabited by yet<br />

ano<strong>the</strong>r ‘different’ species which has been described at <strong>the</strong> earliest opportunity.<br />

Some of <strong>the</strong>se plants have never been validly described but have been distributed<br />

with field numbers and/or tentative names. What has made this situation worse has<br />

been <strong>the</strong> careless subsequent propagation of some of this material. As a result <strong>the</strong>re<br />

is a distinct possibility that <strong>the</strong> plants when distributed are frequently wrongly<br />

identified or wrongly numbered which renders <strong>the</strong>m scientifically worthless.<br />

It is with all this in mind that we have had a long hard cold look at <strong>the</strong> group as a<br />

whole and have come to certain conclusions which represent our own current<br />

thoughts on <strong>the</strong> taxonomy of <strong>the</strong> <strong>genus</strong>. It is not our intention to fur<strong>the</strong>r clutter up <strong>the</strong><br />

synonymy, and so in <strong>the</strong> main we use <strong>the</strong> original scheme proposed by Bertrand in<br />

1951 toge<strong>the</strong>r with a number of refinements. We accept <strong>the</strong> view of Donald (1975)<br />

that Sulco<strong>rebutia</strong> toge<strong>the</strong>r with Weingartia have evolved from a separate<br />

evolutionary line and are thus excluded from Rebutia.<br />

5


aran… -><br />

arran…<br />

In proposing <strong>the</strong> following arrangement we realise that whatever scheme we<br />

propose it will not be universally accepted. We have not slavishly followed any one<br />

particular system but have attempted to amalgamate <strong>the</strong> best of a number of different<br />

arrangements. One of <strong>the</strong> criteria we have used is <strong>the</strong> breeding system, i.e. whe<strong>the</strong>r a<br />

species is self sterile or self fertile. This neatly divides <strong>the</strong> Section IV Rebutia into two<br />

workable subsections. It also divides <strong>the</strong> large Backeberg <strong>genus</strong> Mediolobivia into<br />

two distinct sections, Section II Mediolobivia and Section III Digito<strong>rebutia</strong>. There will<br />

always be a problem with those plants which bridge <strong>the</strong> gap between <strong>the</strong> different<br />

sections (being nei<strong>the</strong>r wholly self fertile, nor wholly self sterile), but we believe that<br />

ours is <strong>the</strong> most workable scheme to date.<br />

The following is an attempt at producing a description for <strong>the</strong> whole <strong>genus</strong> and is<br />

based on Buxbaum (1958).<br />

Rebutia K. Sch. emend Buining & Donald.<br />

Dwarf, globular or slightly cylindrical plants usually making offshoots from <strong>the</strong> base.<br />

Tubercles often distinct, spirally arranged and sometimes formed into distinct ribs.<br />

Spines very variable in length and colour; sometimes short, stiff and bristle-like but<br />

often long and flexuous. Flowers always produced from old areoles, usually<br />

appearing crowded at <strong>the</strong> base and very large in relation to <strong>the</strong> size of <strong>the</strong> plant. The<br />

flowers are brilliantly coloured, red, yellow, purple and occasionally white,<br />

campanulate or funnel-shaped. The receptacular scales are sometimes naked but in<br />

o<strong>the</strong>rs with copious hairs and bristles. Stamens are produced from <strong>the</strong> funnel part of<br />

<strong>the</strong> flower never from a ring in <strong>the</strong> throat. Occasionally <strong>the</strong> style is united to <strong>the</strong><br />

receptacle wall. The fruit is a small capsule which opens by a circular split so that <strong>the</strong><br />

lower half which remains on <strong>the</strong> plant is like a shallow bowl in which some of <strong>the</strong><br />

seeds remain. Seeds small, black and shining but also dull and earth coloured in<br />

some species.<br />

Classification of <strong>the</strong> <strong>genus</strong> Rebutia<br />

Rebutia K. Sch. emend. Buining & Donald Type species Rebutia minuscula K. Sch.<br />

Section I Aylostera (type Rebutia deminuta (Web.) Br. & R.)<br />

Section II Mediolobivia (type Rebutia aureiflora Backbg.)<br />

Subsection I Seti<strong>rebutia</strong>e<br />

Subsection II Cylindro<strong>rebutia</strong>e<br />

Section III Digito<strong>rebutia</strong> (type Rebutia pygmaea (R. E. Fries) Br. & R.)<br />

Subsection I Pygmaeae<br />

Subsection II Euan<strong>the</strong>mae<br />

Section IV Rebutia (type Rebutia minuscula K. Sch.)<br />

Subsection I Rebutiae<br />

Subsection II Medio<strong>rebutia</strong>e<br />

Lists of <strong>the</strong> species which belong to a particular Section or Subsection, toge<strong>the</strong>r with<br />

distribution maps and locational data, are given in <strong>the</strong> chapter devoted to<br />

geographical distribution.<br />

6


companulate -><br />

campanulate<br />

Analytical key to <strong>the</strong> Rebutia Sections<br />

For reasons which have been explained, this key will not separate those species<br />

which bridge <strong>the</strong> gap between <strong>the</strong> different sections i.e. those species with<br />

intermediate characters. It should never<strong>the</strong>less help to delimit <strong>the</strong> majority of<br />

species. It is based on a key published by Krainz (1967) and amended by B. Fearn.<br />

A Body globular or cylindrical with <strong>the</strong> tubercles arranged on distinct ribs,<br />

which may be vertical or spirally arranged. Body often suffused with<br />

amethyst particularly during <strong>the</strong> winter months.<br />

B Body cylindrical, flowers broad funnel-shaped to campanulate. Flowers<br />

self sterile. Section II Mediolobivia Subsection II Cylindro<strong>rebutia</strong>e<br />

BB Body globular or short cylindrical, flowers narrow funnel shaped,<br />

lower part of receptacle usually fused with <strong>the</strong> base of style. Flowers self<br />

fertile. Section III Digito<strong>rebutia</strong><br />

C Body obviously cylindrical, spines less than 5mm long, strongly<br />

appressed, areoles arranged in vertical rows.<br />

Section III Digito<strong>rebutia</strong> Subsection I Pygmaeae<br />

CC Body globular, spines 5mm. or longer, always slightly erect never<br />

appressed, areoles arranged spirally, occasionally in vertical rows.<br />

Section III Digito<strong>rebutia</strong> Subsection II Euan<strong>the</strong>mae<br />

AA Body globular to flattened globular, with indistinct ribs. Body green. Plants<br />

usually densely spined.<br />

C Receptacular scales with hairs and bristles, seeds ± earth coloured.<br />

D Style not fused to <strong>the</strong> base of <strong>the</strong> receptacle.<br />

Section II Mediolobivia Subsection I Seti<strong>rebutia</strong>e<br />

DD Basal portion of style usually fused with <strong>the</strong> lower portion of <strong>the</strong><br />

receptacle. Section I Aylostera<br />

CC Receptacular scales naked (i.e. without hairs and bristles). Seeds black<br />

and glossy, dome shaped usually with a prominent white basal hylum<br />

area. Section IV Rebutia<br />

E Flowers self fertile. Subsection I Rebutiae<br />

EE Flowers self sterile. Subsection II Medio<strong>rebutia</strong>e<br />

It is beyond <strong>the</strong> scope of this booklet to include a workable species key.<br />

Flowers<br />

All species in <strong>the</strong> <strong>genus</strong> Rebutia are day flowering and insect pollinated. Most<br />

species are self fertile with <strong>the</strong> notable exception of 2 groups - Section II<br />

Mediolobivia and Section IV Subsection II Medio<strong>rebutia</strong>e.<br />

In Europe, Rebutias flower in late spring usually during <strong>the</strong> months of April and<br />

May. The first species to flower is Rebutia marsoneri which usually flowers at <strong>the</strong> end<br />

of February and beginning of March. It is thus one of <strong>the</strong> heralds of Spring.<br />

Occasionally it flowers much earlier than this and on one notable occasion a plant of<br />

this species was in flower at Abbey Brook on Christmas Day morning. The last plants<br />

to flower are some members of Section I Aylostera such as R. deminuta, R.<br />

pseudodeminuta and R. fiebrigii. There appears to be a distinct sequence in <strong>the</strong><br />

flowering of <strong>the</strong>se plants. This time sequence may be of great biological significance<br />

because in habitat it would maintain biological isolation even when <strong>the</strong> plants were<br />

growing in close proximity to each o<strong>the</strong>r.<br />

7<br />

hylam -><br />

hylum


In cultivation it is possible to change <strong>the</strong> flowering times of <strong>the</strong>se plants by <strong>the</strong><br />

extension of cool temperatures. This has <strong>the</strong> effect of delaying <strong>the</strong> flowering time until<br />

late May or June. Flowering time can also be altered by increasing <strong>the</strong> minimum<br />

temperature during February thus shortening <strong>the</strong> winter period. This has <strong>the</strong> effect of<br />

making <strong>the</strong> plants flower in March.<br />

Although it is possible to alter <strong>the</strong> timing of flowering, <strong>the</strong>re is one overall<br />

requirement in <strong>the</strong> successful cultivation of <strong>the</strong>se plants which appears to determine<br />

whe<strong>the</strong>r <strong>the</strong> plants actually produce or do not produce flowers. In our experience<br />

this is <strong>the</strong> winter rest period. It appears to be essential for <strong>the</strong> temperature to fall near<br />

to or below freezing for some considerable time. Having due regard to <strong>the</strong> high<br />

altitudes in <strong>the</strong> Andes of <strong>the</strong> natural habitats of <strong>the</strong>se plants 1500-4000 metres altitude<br />

(4900-13,000ft.) it is not really surprising that this is <strong>the</strong> case. At Abbey Brook we now<br />

keep all our Rebutia plants in a cold unheated glasshouse during <strong>the</strong> winter toge<strong>the</strong>r<br />

with two o<strong>the</strong>r genera Lobivia and Oreocereus.<br />

The cold requirement may well account for <strong>the</strong> poor flowering of <strong>the</strong>se plants in<br />

Sou<strong>the</strong>rn California and <strong>the</strong> total lack of flower production in <strong>the</strong> tropics. We have<br />

already suggested to our friends in <strong>the</strong> tropics that <strong>the</strong>se plants should be wrapped in<br />

newspaper and placed in a refrigerator (not a deep freeze) for some weeks. We<br />

have not yet received <strong>the</strong> results of <strong>the</strong>se experiments.<br />

Floral Structure<br />

The floral structure of <strong>the</strong> <strong>genus</strong> Rebutia has in <strong>the</strong> past been used to justify<br />

splitting <strong>the</strong> single <strong>genus</strong> Rebutia into a number of separate genera. It is now<br />

appreciated that such a split cannot be justified on <strong>the</strong> basis of floral structure alone,<br />

as it has been observed that <strong>the</strong>re are plants with a floral structure which is<br />

intermediate between <strong>the</strong> different sections.<br />

For example, <strong>the</strong> <strong>genus</strong> Aylostera was separated from <strong>the</strong> rest of Rebutia on <strong>the</strong><br />

observation that <strong>the</strong> base of <strong>the</strong> style was fused to <strong>the</strong> base of <strong>the</strong> corolla. In o<strong>the</strong>r<br />

groups for example Mediolobivia and Rebutia <strong>the</strong> style is completely free from <strong>the</strong><br />

base of <strong>the</strong> corolla. Digito<strong>rebutia</strong> was an intermediate group in which <strong>the</strong> style was<br />

only partially free. It is now recognised that <strong>the</strong>se differences are not exhibited by all<br />

<strong>the</strong> related species of a section and cannot <strong>the</strong>refore be used to justify <strong>the</strong><br />

maintenance of a number of separate genera.<br />

Similarly <strong>the</strong> presence or absence of hairs and bristles in <strong>the</strong> axils of <strong>the</strong><br />

receptacle scales is again not a good universal character for splitting up <strong>the</strong> <strong>genus</strong>. It<br />

has been found that <strong>the</strong>re are a number of species which bridge <strong>the</strong> gap between <strong>the</strong><br />

previously recognised generic units. A number of fine illustrations of <strong>the</strong> floral<br />

characters of <strong>the</strong> <strong>genus</strong> have been published Krainz (1967), and Brederoo (Donald &<br />

Brederoo, 1978). The drawings in this booklet are by Brian Fearn and have not been<br />

published previously.<br />

8


B<br />

Rebutia senilis f. stuemeri nat. size<br />

Rebutia minuscula f. grandiflora nat size<br />

9


Rebutia pygmaea nat. size<br />

Rebutia marsoneri nat. size<br />

10


Rebutia einsteinii f. schmiedcheniana nat. size<br />

Rebutia muscula nat. size<br />

11


REBUTIA CHRYSACANTHA<br />

Fig. 3 Beautiful large flowered species closely related to R. senilis.<br />

12


occuring -><br />

occurring<br />

The Geographical Distribution of <strong>the</strong> <strong>genus</strong> Rebutia<br />

The geographical distribution of <strong>the</strong> <strong>genus</strong> Rebutia occurs in North West<br />

Argentina and <strong>the</strong> sou<strong>the</strong>rn provinces of Bolivia on <strong>the</strong> eastern side of <strong>the</strong> Andes. The<br />

altitude varies between 1500-4000 metres (4,900-13,000ft. above sea level). The<br />

<strong>genus</strong> can be split into four main taxonomic sections. Although <strong>the</strong>se sections do not<br />

fall into distinct geographical groupings, certain trends can be detected. Map No. 1<br />

shows <strong>the</strong> important place names and country boundaries and Maps 2-4 <strong>the</strong><br />

geographical distribution of Rebutia species.<br />

Section I Aylostera (Map No. 4)<br />

This section has a very extensive distribution in Argentina and Bolivia. It occurs<br />

from Tuome Province in <strong>the</strong> south, throughout Salta and Jujuy provinces in Argentina<br />

to Tarija, Potosi, Chuquisaca and Santa Cruz Departments of Bolivia.<br />

a) Rebutia deminuta Weber and allied species. Rebutia deminuta is <strong>the</strong> most<br />

sou<strong>the</strong>rly member of this group coming from Trancas, Tucuman province<br />

Argentina. The closely related form Rebutia pseudominuscula occurs over an<br />

extensive area from central Salta province through Jujuy province and into<br />

Tarija.<br />

b) Rebutia pseudodeminuta Backbg. and allied species. Rebutia pseudodeminuta<br />

is widely distributed from south Salta to central Tarija and into<br />

Chuquisaca and Potosi. Many of <strong>the</strong> recently described ‘species’ of Ritter,<br />

Rausch and Lau represent geographical forms of R. pseudodeminuta. (e.g. R.<br />

maxima FR755, R. albiareolata FR761, R. tamboensis FR1142 from Tarija and R.<br />

nitida FR769 from Chuquisaca.)<br />

c) Rebutia kupperiana Böd. and allied species. Rebutia kupperiana is restricted in<br />

habitat to an area around Narvaez in Tarija department Bolivia although several<br />

o<strong>the</strong>r related Ritter ‘species’ belonging to this group have been described as<br />

coming from south Chuquisaca. (e.g. R. tuberosa Ritt., and R. rubiginosa Ritt.)<br />

d) Rebutia fiebrigii (Guerke) Br. & R. and allied species. The Rebutia fiebrigii<br />

complex occurs over an extensive area of sou<strong>the</strong>rn Bolivia and nor<strong>the</strong>rn<br />

Argentina. The main area of distribution occurs from Tarija and Chuquisaca in<br />

Bolivia to nor<strong>the</strong>rn Salta in Argentina. There is an outlier of distribution in Santa<br />

Cruz department where Rebutia ithyacantha (Card.) Diers, R. donaldiana Lau<br />

and Rowley and R. vallegrandensis Card. occur. Rebutia spinosissima has a<br />

very extensive distribution from north Salta in Argentina through Tarija and into<br />

south Chuquisaca in Bolivia.<br />

Rebutia fiebrigii itself has by far <strong>the</strong> largest distribution area of any member of<br />

this group occurring from La Quiaca in Jujuy province in Argentina to central<br />

Chuquisaca in Bolivia.<br />

e) Rebutia spegazziniana Backbg. and allied species. This species is found in<br />

Salta province near to <strong>the</strong> border with Bolivia and into Tarija department from<br />

Padcaya to Tarija. The closely related ‘species’ R. fusca Ritt., R. vulpina Ritt., and<br />

R. tarvitaensis Ritt., are all found in Chuquisaca department, Bolivia.<br />

f) Rebutia heliosa Rausch. This species has a limited distribution area near<br />

Narvaez in Tarija department, Bolivia. There are a number of ‘species’ closely<br />

related to R. heliosa apart from var. cajasensis and var. conderensis, <strong>the</strong>se are R.<br />

perplexa and R. espinosa. Both of <strong>the</strong>se are found in Tarija near to R. heliosa<br />

and at Sou<strong>the</strong>rn Cinti R. albopectinata occurs.<br />

13


Section II Mediolobivia<br />

This series has a very limited distribution occurring only in Argentina in an area<br />

of north Salta and south Jujuy provinces.<br />

Subsection I Seti<strong>rebutia</strong>e. Plants in this section all occur near to <strong>the</strong> Salta - Jujuy<br />

border around Quebrada del Toro. Rebutia aureiflora var. elegans and related forms<br />

have <strong>the</strong> most sou<strong>the</strong>rly distribution of this group occurring around Quebrada<br />

Escoipe.<br />

Subsection II Cylindro<strong>rebutia</strong>e. There is probably only a single species which<br />

belongs in this section - R. einsteinii. The myriad of different names appear to refer to<br />

geographical variants. These plants have been found in ei<strong>the</strong>r Salta or Jujuy<br />

provinces.<br />

Section in Digito<strong>rebutia</strong>. (Map No. 2)<br />

This section has a very extensive distribution from Salta and Jujuy provinces in<br />

Argentina in <strong>the</strong> south to <strong>the</strong> Bolivian departments of Tarija, Chuquisaca, Potosi and<br />

Oruro in <strong>the</strong> north.<br />

There appear to be several separate centres of distribution where a number of<br />

named species occur in close proximity. It is here suggested that <strong>the</strong>se ‘species’<br />

should be subjected to an intensive ecological investigation to determine whe<strong>the</strong>r<br />

<strong>the</strong>se populations are biologically isolated, by pollination vectors and flowering<br />

times. Admittedly members of this series are all self fertile, but <strong>the</strong>ir highly coloured<br />

flowers would attract pollinating insects. Gene flow between populations would<br />

occur if <strong>the</strong> populations which were in close proximity flowered at <strong>the</strong> same time.<br />

We have determined that <strong>the</strong>re are 5 separate areas of distribution for <strong>the</strong><br />

majority of members of this Section.<br />

a) Argentina - Bolivia border areas, Jujuy province.<br />

e.g. R. eos Rausch, R. costata Werd., R. pygmaea (R. E. Fries) Br. & R.<br />

b) Mendez province, Tarija, Bolivia.<br />

e.g. R. brunneoradicata Ritt., R. colorea Ritt.<br />

c) Camargo - Culpina region, Cinti province, Chuquisaca, Bolivia,<br />

e.g. R. diersiana Rausch, R. mixta Ritt.<br />

d) South Chichas province, Potosi, Bolivia,<br />

e.g. R. iridescens Ritt., R. rosalbiflora Ritt.<br />

e) Environs of Potosi City, Potosi, Bolivia.<br />

e.g. R. pygmaea, R. costata f. pilifera Buin. & Donald.<br />

The species found in Oruro also belong here. The proliferation of names which<br />

is particularly true of section III, are at best, geographical variants of very few<br />

species.<br />

Section IV Rebutia (Map No. 3)<br />

Subsection I Rebutiae.<br />

This subsection contains three species complexes. These are R. minuscula, R.<br />

senilis and R. xanthocarpa. The R. minuscula complex includes R. grandiflora<br />

Backbg., R. violaciflora Backbg. and R. kariusiana. The group is totally Argentinian in<br />

distribution ranging from <strong>the</strong> north Tucuman to Salta and Jujuy provinces. The area is<br />

bounded by Tucuman City, Quebrada Escoipe, Cachipampa and Quebrada del<br />

Toro. R. minuscula occurs throughout this region apparently remaining<br />

14


morphologically constant until reaching <strong>the</strong> nor<strong>the</strong>rn limit of its distribution where it<br />

appears to intergrade with R. violaciflora.<br />

Subsection II Medio<strong>rebutia</strong>e.<br />

This subsection contains two species complexes. These are R. wessneriana and<br />

R. marsoneri. It appears that this subsection has a much wider distribution than<br />

Subsection I Rebutiae, ranging from south Salta province in Argentina e.g. R.<br />

wessneriana Bew. to Huari-Huari, Potosi department, Bolivia e.g. R. binnewaldiana<br />

W. Heinr. and R. fiebigiana W. Heinr. Some doubt has been raised as to <strong>the</strong><br />

au<strong>the</strong>nticity of <strong>the</strong> last two species. The habitat data may be incorrect. Seed<br />

distributed from <strong>the</strong>se habitats has produced plants that resemble species of Series I<br />

Aylostera (Donald, Ashingtonia 2 67). If this is correct <strong>the</strong>n <strong>the</strong> distribution is<br />

restricted to Jujuy and Salta provinces of Argentina and into Tarija department<br />

Bolivia where R. wessneriana is also found. The R. marsoneri complex is distributed<br />

throughout nor<strong>the</strong>rn Jujuy province, Argentina.<br />

Distribution maps and locational data<br />

The following is a list of <strong>the</strong> species toge<strong>the</strong>r with locational data which have<br />

been used in preparing <strong>the</strong> distribution maps. It is interesting to note that often<br />

species belonging to <strong>the</strong> same Rebutia section (i.e. closely related) have often been<br />

collected at <strong>the</strong> same place. The altitude at <strong>the</strong> different sites may vary, but <strong>the</strong> vital<br />

information on genetic isolation is not available. Without this, toge<strong>the</strong>r with ecological<br />

and experimental taxonomic data <strong>the</strong> suspicion is that <strong>the</strong>se different ‘species’ are<br />

identical merely representing <strong>the</strong> range of species variation. For example <strong>the</strong><br />

following species numbers 18, 20, 22, 24, 40, 41, 44, 45 have <strong>the</strong> same location - Mal<br />

Paso, Sth. Chichas, Potosi, Bolivia. Also species numbers 4, 12, 16, 26, 36, 46 occur at -<br />

Camargo - Culpina, Chuquisaca, Bolivia.<br />

Rebutia Section III Digito<strong>rebutia</strong> (See Map No. 2)<br />

1 brachyantha (Wessn.) Buin. & Donald. Tarija, Tarija, Bolivia.<br />

2 brunescens Rausch. Huari-Huari, Potosi, Bolivia.<br />

3 brunneoradicata Ritter. San Antonio, Mendez, Tarija, Bolivia.<br />

4 canacruzensis Rausch. Camargo-Culpina, Chuquisaca, Bolivia.<br />

5 carmeniana Rausch. Caspala, Jujuy, Argentina.<br />

6 cincinnata Rausch. Potosi, Potosi, Bolivia.<br />

7 colorea Ritter. San Antonio, Mendez, Tarija, Bolivia.<br />

8 costata (Werd.) Krainz. La Quiaca, Jujuy, Argentina.<br />

9 costata f. eucalyptana (Backbg.) Buin. & Donald. Eucaliptos, La Paz, Bolivia.<br />

10 costata f. pilifera Buin. & Donald Potosi, Potosi, Bolivia.<br />

11 densispectinata Ritter nomen nudum. Jucanas, Tarija, Bolivia.<br />

12 diersiana Rausch. Camargo-Culpina, Chuquisaca, Bolivia.<br />

13 eos Rausch. Tafna, Jujuy, Argentina.<br />

15<br />

Quaica -><br />

Quiaca


14 euan<strong>the</strong>ma (Backbg.) Buin. & Donald. Quebrada de Humahuaca, Jujuy,<br />

Argentina.<br />

15 euan<strong>the</strong>ma f. oculata (Werd.) Buin. & Donald. Humahuaca, Jujuy, Argentina.<br />

16 friedrichiana Rausch. Camargo-Culpina, Chuquisaca, Bolivia.<br />

17 graciliflora Backbg. Chichas, Potosi, Bolivia.<br />

18 gracilispina Ritter. Mal Paso, Sth. Chichas, Potosi, Bolivia.<br />

19 hirsutissima Card. Bueto, Villaserrano, Chuquisaca, Bolivia.<br />

20 iridescens Ritter. Mal Paso, Sth. Chichas, Potosi, Bolivia.<br />

21 iscayachensis Rausch. Iscayachi, Tarija, Bolivia.<br />

22 lanosiflora Ritter. Mal Paso, Sth. Chichas, Potosi, Bolivia.<br />

23 minutissima Ritter. Tarija, Tarija, Bolivia.<br />

24 mixta Ritter. Mal Paso, Sth. Chichas, Potosi, Bolivia.<br />

25 mixticolor Ritter. San Antonio, Mendez, Tarija, Bolivia.<br />

26 mudanensis Rausch. Camargo-Culpina, Chuquisaca, Bolivia.<br />

27 odontopetala Ritter. Cueva, Sth. Cinti, Chuquisaca, Bolivia.<br />

28 patericalyx Ritter. Cueva, Sth. Cinti, Chuquisaca, Bolivia.<br />

29 pauciareolata Ritter. San Antonio, Mendez, Tarija, Bolivia.<br />

30 paucicostata Ritter. San Antonio, Mendez, Tarija, Bolivia.<br />

31 poecilantha Ritter. North Cinti, Chuquisaca, Bolivia.<br />

32 potosina Ritter. Potosi, Potosi, Bolivia.<br />

33 pseudopygmaea Ritter nomen nudum. San Antonio, Mendez, Tarija, Bolivia.<br />

34 pygmaea (R. E. Fries) Br. & R. Yavi, Jujuy, Argentina., Potosi, Potosi, Bolivia.<br />

35 pygmaea f. haefneriana (Cullm.) Buin. & Donald. Oruro, Oruro, Bolivia, Iturbe,<br />

Argentina, Chichas, Potosi, Bolivia.<br />

36 raulii Rausch Camargo-Culpina, Chuquisaca, Bolivia.<br />

37 rauschii Zecher. Huari-Huari, Potosi, Bolivia.<br />

38 ritteri (Wessn.) Buin. & Donald, Iscayachi, Tarija, Bolivia.<br />

39 ritteri f. nigricans (Wessn.) Buin. & Donald. Quebrada del Toro, Salta,<br />

Argentina.<br />

40 rosalbiflora Ritter. Mal Paso, Sth. Chichas Province, Bolivia.<br />

41 rutiliflora Ritter. Mal Paso, Sth. Chichas Province, Bolivia.<br />

42 salpingantha Ritter. Villazon, Potosi, Bolivia.<br />

43 tarvitaensis Ritter. Tarvita, Azurduy, Chuquisaca, Bolivia.<br />

44 torquata Ritter. Mal Paso, Sth. Chichas Province, Bolivia.<br />

45 tropaeoliptica Ritter. Mal Paso, Sth. Chichas Province, Bolivia.<br />

46 violascens Ritter. Camargo, Cinti, Chuquisaca, Bolivia.<br />

Rebutia Section II Mediolobivia Subsection I Seti<strong>rebutia</strong>e (See Map No. 2)<br />

55 R. aureiflora Backbg. Quebrada del Toro, Salta, Argentina.<br />

56 R. aureiflora f. rubelliflora (Backbg.) Buin. & Donald. Quebrada del Toro,<br />

Salta, Argentina.<br />

57 R. aureiflora f. duursmaiana (Backbg.) Buin. & Donald. Yacones, Salta,<br />

Argentina.<br />

58 R. aureiflora f. boedeckeriana (Backbg.) Buin. & Donald. Nr. Chani Volcano,<br />

Jujuy, Argentina.<br />

59 R. aureiflora v. elegans (Backbg.) Buin. & Donald. Quebrada Escoipe, Salta,<br />

Argentina.<br />

16<br />

Chuquisiana -><br />

Chuquisaca<br />

mudadensis -><br />

mudanensis<br />

poelcilantha -><br />

poecilantha


Rebutia Section IV Rebutia Subsection I Rebutiae (See Map No. 3)<br />

60 fabrisii Rausch. Valle Grande, Salta, Argentina.<br />

61 minuscula K. Sch. Tucuman, Tucuman Prov., Argentina.<br />

62 minuscula var. grandiflora (Backbg.) Buin. & Donald. Quebrada Escoipe,<br />

Salta, Argentina.<br />

63 minuscula var. violaciflora (Backbg.) Buin. & Donald. Quebrada Escoipe,<br />

Salta, Argentina.<br />

64 margarethae Rausch. Iruya, Santa Victoria, Salta, Argentina.<br />

65 padcayensis Rausch. Padcaya, Tarija, Bolivia.<br />

66 senilis Backbg. Quebrada Escoipe, Salta, Argentina.<br />

67 xanthocarpa Backbg. Quebrada del Toro, Salta, Argentina.<br />

Rebutia Section IV Rebutia Subsection II Medio<strong>rebutia</strong>e (See Map No. 3)<br />

68 binnewaldiana W. Heinr. Huari-Huari, Potosi, Bolivia.<br />

69 fiebigiana W. Heinr. Huari-Huari, Potosi, Bolivia.<br />

70 krainziana Kess. Yavi, Jujuy, Argentina.<br />

71 marsoneri Werd. North Jujuy, Argentina.<br />

72 marsoneri f. sieperdaiana (Buin.) Buin. & Donald. Salta, Salta, Argentina.<br />

74 wessneriana Bewg. Quebrada de Humahuaca, Jujuy, Argentina; Salta,<br />

Salta, Argentina; Tarija, Bolivia and Guachipas, Escoipe, Salta, Argentina.<br />

Rebutia Section I Aylostera (See Map No. 4)<br />

76 albiflora Ritter & Buin. Rio Pilaya, North East Tarija, Bolivia.<br />

77 albipilosa Ritter. Narvaez, Tarija, Bolivia.<br />

78 albiareolata Ritter. Padcaya, Arce, Tarija, Bolivia.<br />

79 albopectinata Rausch. Culpina, Chuquisaca, Bolivia.<br />

80 aureispina Knize nomen nudum. Jucanas, Tarija, Bolivia.<br />

81 buiningiana Rausch. Nr. Iruya, Salta, Argentina.<br />

82 camargoensis Rausch. Camargo-Culpina, Chuquisaca, Bolivia.<br />

83 cajasensis Ritter. Tarija, Tarija, Bolivia.<br />

84 cintiensis Ritter. Camargo, Sth. Cinti, Chuquisaca, Bolivia.<br />

85 deminuta (Weber.) Br. & R. Trancas, Tucuman, Argentina.<br />

86 donaldiana Lau & Rowley. Pucara, Vallegrande, Santa Cruz, Bolivia.<br />

87 fiebrigii (Guerke) Br. & R. Iscayachi, Tarija, Bolivia.<br />

88 flavistyla Ritter. Tarija, Tarija, Bolivia.<br />

89 froehlichiana Rausch. Yuquina, Cinti, Chuquisaca, Bolivia.<br />

90 fulviseta Rausch. Padcaya, Arce, Tarija, Bolivia.<br />

91 fusca Ritter. Cinti Chuquisaca, Bolivia.<br />

92 heliosa Rausch. Road to Narvaez, Tarija, Bolivia.<br />

93 hoffmannii Rausch. Santa Victoria, Salta, Argentina.<br />

94 huasiensis Rausch. Inca Huasi, Cinti, Chuquisaca, Bolivia.<br />

95 ithyacantha (Card.) Diers. North Comarapa, Valle Grande, Santa Cruz, Bolivia.<br />

96 jujuyana Rausch. Quebrada de Humahuaca, Jujuy, Argentina.<br />

97 jujuyensis Lau. Santa Victoria, Salta, Argentina.<br />

98 kieslingii Rausch. Caspala, Jujuy, Argentina.<br />

99 kupperiana Böd. Narvaez, Tarija, Bolivia.<br />

C<br />

17<br />

fabrisi -><br />

fabrisii<br />

sieperdaiana -><br />

sieperdaiana<br />

Humuhuaca -><br />

Humahuaca


100 lauii Donald. Cajas Pass, Tarija, Bolivia.<br />

101 leucan<strong>the</strong>ma Rausch. Camargo-Culpina, Chuquisaca, Bolivia.<br />

102 mamillosa Rausch. Camargo, Chuquisaca, Bolivia.<br />

103 muscula Ritter & Thiele. Narvaez, Tarija, Bolivia.<br />

104 napina Ritter. South Cinti, Bolivia.<br />

105 nitida Ritter. Tarvita Azurduy, Chuquisaca, Bolivia.<br />

106 nogalensis Ritter. Tarvita Azurduy, Chuquisaca, Bolivia.<br />

107 pseudodeminuta Backbg. South Salta, Argentina - to - Chuquisaca, Bolivia.<br />

108 deminuta f. pseudominuscula (Speg.) Buin. & Donald. Cachi, Salta, Argentina.<br />

109 pulchella Rausch. North of Padilla, Chuquisaca, Bolivia.<br />

110 pulvinosa Ritter & Buin. Narvaez, Tarija, Bolivia.<br />

111 robustispina Ritter. Tarija, Tarija, Bolivia.<br />

112 rubiginosa Ritter. South Cinti, Chuquisaca, Bolivia.<br />

113 schatzliana Rausch. Camargo-Culpina, Chuquisaca, Bolivia.<br />

114 spegazziniana Backbg. Salta, Victoria - to - Tarija, Bolivia.<br />

115 sphaerica Ritter. Tarija, Tarija, Bolivia.<br />

116 spinosissima Backbg. Santa Victoria, Salta, Argentina.<br />

117 supthutiana Rausch. Camargo-Culpina, Chuquisaca, Bolivia.<br />

119 tarijensis Rausch. Tarija, Tarija, Bolivia.<br />

120 tarvitaensis Ritter. Tarvita, Azurduy, Chuquisaca, Bolivia.<br />

121 tuberosa Ritter. South Cinti, Chuquisaca, Bolivia.<br />

122 vallegrandensis Card. Candellaria, Valle Grande, Santa Cruz, Bolivia.<br />

123 vulpina Ritter. Cinti, Chuquisaca, Bolivia.<br />

124 zecheri Rausch. Iscayachi, Tarija, Bolivia.<br />

Rebutia Section II Mediolobivia Subsection II Cylindro<strong>rebutia</strong>e (See Map No. 4)<br />

125 einsteinii Frič ex Backbg. La Quiaca, Jujuy, Argentina.<br />

127 einsteinii var. gonjianii (Kies.) Donald. Tilcara, Jujuy, Argentina.<br />

128 einsteinii f. schmiedcheniana (Kohl.) Buin. & Donald. Chani Volcano, Jujuy,<br />

Argentina.<br />

129 einsteinii f. rubriviridis (Frič & Kreuzgr. ex. Backbg.) Donald.<br />

Quebrada del Toro, Salta, Argentina.<br />

130 einsteinii f. conoidea (Wessn.) Buin. & Donald. Cham Volcano, Jujuy, Argentina.<br />

18<br />

pulchell -><br />

pulchella


Cultivation<br />

Cultivation of <strong>the</strong> majority of species should present no real problems. In <strong>the</strong><br />

growing season (March-October) <strong>the</strong>y need a considerable amount of water. This<br />

may mean watering 2 or 3 times a week in hot sunny wea<strong>the</strong>r but normally once a<br />

week should be sufficient. Try not to get <strong>the</strong> soil soaking wet which leads to bloated<br />

plants which are very susceptible to soft rot. Wet soil also encourages Sciara Fly<br />

which can also lead to loss of plants. In <strong>the</strong> winter (October-March) <strong>the</strong> watering<br />

requirements are completely different as <strong>the</strong> plants are not growing. For successful<br />

flower production it appears to be essential for <strong>the</strong> temperature to fall near to or<br />

below freezing point for some considerable time. During this period <strong>the</strong> plants must<br />

be kept absolutely dry and you will <strong>the</strong>n be rewarded by <strong>the</strong> production in early<br />

spring of <strong>the</strong> superb, brightly coloured flowers produced by all members of this<br />

<strong>genus</strong>.<br />

All Rebutia species need plenty of light and a good sunny position is essential.<br />

Although <strong>the</strong>se plants can be successfully grown on <strong>the</strong> window sill, a greenhouse or<br />

conservatory is a better place especially if you wish <strong>the</strong> plants to flower. We have<br />

found from long experience that plants that are watered and kept warm 7°C (45°F) or<br />

above in <strong>the</strong> winter are very shy flowering.<br />

We can recommend <strong>the</strong> following compost although Rebutias can be grown<br />

successfully in virtually any mixture:- 3 or 4 parts of Fisons Levington Potting<br />

Compost and 1 part of coarse lime-free sand.<br />

We grow <strong>the</strong> majority of our plants in this mixture. It has also been found<br />

beneficial to feed <strong>the</strong> plants during <strong>the</strong> growing season with a low nitrogen, high<br />

potash fertiliser such as Fisons Tomorite. We normally feed <strong>the</strong> plants 3 times during<br />

<strong>the</strong> growing season i.e. once every 6 weeks using <strong>the</strong> recommended strength<br />

fertilizer. Repotting is only necessary when <strong>the</strong> plants have outgrown <strong>the</strong> container<br />

<strong>the</strong>y are growing in. Repotting is usually done at <strong>the</strong> beginning of <strong>the</strong> growing<br />

season. If regular feeding is given as recommended, repotting once in 2 or 3 years<br />

may be all that is necessary, particularly with <strong>the</strong> slower growing species.<br />

Vegetative Propagation. Rebutias can be propagated by removing and rooting<br />

offsets but it is not a recommended practice as <strong>the</strong> plants are so easy to grow from<br />

seed. R. albiflora is <strong>the</strong> only real exception and this species is usually propagated by<br />

rooting offsets as seed is rarely available. Grafting is also sometimes used to<br />

propagate <strong>the</strong> slower growing species such as R. aureiflora, R. einsteinii and R.<br />

heliosa but it is not really necessary as <strong>the</strong> plants grow well on <strong>the</strong>ir own roots.<br />

Growing from seed. Rebutias are best propagated from seed. If you are collecting<br />

your own seed it is best to collect <strong>the</strong> fruits when <strong>the</strong>y are fully ripe at <strong>the</strong> end of <strong>the</strong><br />

growing season. Many species produce copious quantities of seed, o<strong>the</strong>rs have only<br />

a few seeds in each capsule. Carefully separate <strong>the</strong> seeds from any adhering capsule<br />

wall or debris. They can <strong>the</strong>n be stored or sown immediately.<br />

To be successful you must try to imitate <strong>the</strong> conditions in which <strong>the</strong> plants grow<br />

naturally. Cacti are nearly always found under scrub or small bushes, on rocky slopes<br />

or on level ground where <strong>the</strong> soil is porous. This is because:-<br />

(1) The slope and porous soil provide adequate drainage.<br />

(2) The scrub provides shade for <strong>the</strong> seedlings and young plants.<br />

(3) The rocks radiate warmth at night and <strong>the</strong> crevices between <strong>the</strong>m trap<br />

pockets of humid air.<br />

23


When to sow. If you do not have a heated propagator, <strong>the</strong> best time to sow is in<br />

Spring. i.e. from March until <strong>the</strong> end of May. Do not sow later than this as <strong>the</strong><br />

seedlings will not be large enough to withstand <strong>the</strong> following winter.<br />

If you have a heated propagator or a warm window sill (Do not use <strong>the</strong> airing<br />

cupboard) you can sow at any time of <strong>the</strong> year, but January and February are <strong>the</strong> best<br />

months.<br />

Compost. We can recommend <strong>the</strong> following composts.<br />

Ei<strong>the</strong>r:<br />

(1) Equal parts of Fisons Levington potting compost and coarse lime-free sand.<br />

or (2) One part John Innes No. 1 or 2 and ¼ part lime-free sand.<br />

Sowing. Fill <strong>the</strong> seed trays or shallow pots to within ¼'' of <strong>the</strong> top with compost and<br />

level <strong>the</strong> surface, removing any large lumps. Water with a watering can fitted with a<br />

fine hose. Scatter <strong>the</strong> seed thinly on <strong>the</strong> surface of <strong>the</strong> damp compost. Seeds can be<br />

spaced out and lightly pressed level with <strong>the</strong> surface using a pencil. Cactus seed<br />

need light before <strong>the</strong>y will germinate, so do not cover <strong>the</strong> seeds with compost and do<br />

not put <strong>the</strong> containers in a dark cupboard.<br />

Temperature. Temperatures of 12-22°C (55-70°F) are best for germination. Once<br />

<strong>the</strong> seedlings are 2-3 weeks old normal growing temperatures are adequate.<br />

Germination. The percentage of seeds that germinate depends on <strong>the</strong> species, age<br />

of <strong>the</strong> seed and temperature, but <strong>the</strong>re should be signs of germination within 2-3<br />

weeks. A few seeds may be slow to germinate, and seedlings may not appear until<br />

2-3 months after sowing.<br />

Watering. If possible, water with a fine mist spray, or place <strong>the</strong> container in a<br />

saucer and water from below. Never let <strong>the</strong> compost dry out, particularly before<br />

germination has taken place, or <strong>the</strong> germinating seeds will be killed. Conversely, do<br />

not allow <strong>the</strong> compost to become waterlogged ei<strong>the</strong>r, else <strong>the</strong> seeds will rot. After<br />

germination keep soil moist - not too wet, not too dry. During <strong>the</strong> first winter <strong>the</strong><br />

seedlings can be kept completely dry in a minimum temperature of 2°C (35°F).<br />

Care of <strong>the</strong> young seedlings. Keep <strong>the</strong> container in an airy place to prevent<br />

‘damping off. Shade <strong>the</strong> seedlings with a single thickness of tissue or newspaper for<br />

<strong>the</strong> first two months, <strong>the</strong>n avoid direct sunlight for a fur<strong>the</strong>r 6 months. After that time<br />

treat as adult plants. Try to ensure <strong>the</strong> seedlings remain a healthy deep green (or<br />

occasionally brownish) colour - a bright red colour indicates that too much light has<br />

been given, thin spindly pale green seedlings results from too little light.<br />

Transplanting. Do not transplant seedlings until <strong>the</strong>y begin to touch one ano<strong>the</strong>r in<br />

<strong>the</strong> container - this will depend on <strong>the</strong> rate of growth of individual species. If in doubt,<br />

leave <strong>the</strong> seedlings in <strong>the</strong> same container for 1 year, and do not transplant until <strong>the</strong><br />

spring of <strong>the</strong> second year. They will not come to any harm even if <strong>the</strong>y look<br />

overcrowded.<br />

Detailed germination studies on Rebutia xanthocarpa f. salmonea have been<br />

made Fearn (1974). It was found that no seed germinated below 9°C or above 26°C It<br />

was found that this species when compared with o<strong>the</strong>r species from South America<br />

e.g. Gymnocalycium spegazzinii, Frailea pumila and Helianthocereus pasacana has<br />

a very restricted temperature range. At least 50% germination occurred between<br />

11∙5°C and 22∙8°C (52°F - 70°F) a range of 11∙3°C.<br />

24<br />

spegazinnii -><br />

spegazzinii


Fig. 4 The germination of R. xanthocarpa f. salmonea has been plotted on a day to<br />

day basis and <strong>the</strong> figures on <strong>the</strong> graph indicate <strong>the</strong> number of days<br />

incubated.<br />

Fig. 5 The curves obtained for H. pasacana and R. xanthocarpa f. salmonea are<br />

shown. On successive days <strong>the</strong> maximum and minimum temperatures at<br />

which 50% of maximum germination was obtained. The resulting U shaped<br />

curves reflect <strong>the</strong> temperature span over which 50% germination occurred,<br />

<strong>the</strong> rate of germination at different temperatures and <strong>the</strong> position of<br />

maximum and minimum temperature at which 50% of maximum germination<br />

was reached.<br />

D<br />

25


Pests and Diseases<br />

Mealy Bugs are <strong>the</strong> most common pests of cacti. They look a bit like tiny wood lice but<br />

with a white waxy coat. They usually aggregate in areas which are not affected by<br />

overhead watering, e.g. in <strong>the</strong> axis of <strong>the</strong> tubercles and around <strong>the</strong> base of <strong>the</strong> plant.<br />

Spraying with Malathion or a combination with Dimethoate will eradicate <strong>the</strong>m.<br />

Root Mealy Bugs usually remain unseen until <strong>the</strong> plants are repotted. They occur as<br />

white woolly colonies which feed on <strong>the</strong> roots. Watering with a systemic insecticide<br />

such as Dimethoate should eradicate <strong>the</strong>m. Repeat <strong>the</strong> dose after seven or ten days.<br />

Red Spider Mite can be a serious pest. Its presence may remain undetected until <strong>the</strong><br />

plants have become severely disfigured with a reddish-brown area on <strong>the</strong><br />

epidermis. It is a very tiny reddish creature just visible to <strong>the</strong> naked eye. Water or<br />

spray <strong>the</strong> plants with a contact or systemic insecticide Repeat <strong>the</strong> dose after ten<br />

days.<br />

Sciara Fly is a relative newcomer as far as pests of Cacti are concerned. The first<br />

warning may be <strong>the</strong> presence of a few tiny black flies near <strong>the</strong> plants. The real pest is<br />

<strong>the</strong> larvae which lives just under <strong>the</strong> soil surface and can cause damage and rot <strong>the</strong><br />

base of <strong>the</strong> plants. Total collapse of <strong>the</strong> plant may result. Watering with a systemic<br />

insecticide should eradicate <strong>the</strong> larvae.<br />

Aphids are not an important pest in <strong>the</strong> cactus collection. They occasionally occur on<br />

flower buds and on young growth and any standard insecticide should deal with<br />

<strong>the</strong>m.<br />

REBUTIA XANTHOCARPA f. VIOLACIFLORA<br />

A charming plant with delicate lilac-rose coloured flowers<br />

26<br />

visable -><br />

visible


Rebutia Descriptions<br />

We have selected <strong>the</strong> following 20 species as representatives of <strong>the</strong> range of<br />

variation exhibited by <strong>the</strong> <strong>genus</strong> Rebutia. It includes plants from all <strong>the</strong> Sections and<br />

many of <strong>the</strong>m are old favourites having been in cultivation for some considerable<br />

time e.g. R. kupperiana, and R. minuscula. O<strong>the</strong>r species which have been included<br />

such as R. albiflora, R. muscula and R. heliosa, are among <strong>the</strong> best of recent<br />

introductions.<br />

REBUTIA ALBIFLORA<br />

Fig. 6 A very distinctive and beautiful species with white flowers.<br />

27


REBUTIA ALBIFLORA<br />

Rebutia albiflora Ritt. & Buin. Taxon (1963) 12 29<br />

Synonyms Aylostera albiflora (Ritt. & Buin.) Backbg.<br />

Classification Section I Aylostera<br />

Body Individual heads up to 25mm. in diameter but usually much smaller<br />

than this. Plants freely offsetting from <strong>the</strong> base and eventually<br />

forming large clumps. Epidermis grey-green.<br />

Areoles tiny, more or less circular, pure white<br />

Spines differentiated into approximately 15 radials and 5 centrals, up to<br />

5mm. long, straight and glassy white.<br />

Flowers inner petals white, outer petals often with a pronounced pink<br />

mid-line, 25mm. wide and 30mm. long. The tube is pink and has<br />

short white bristles at <strong>the</strong> base.<br />

Fruit small, 3-4mm. in diameter, reddish-pink and contains few seeds.<br />

Seed small, matt black.<br />

Type locality Rio Pilaya, N.E. Tarija, Bolivia. Map 4, No. 76.<br />

Discovered by Friedrich Ritter (FR766a)<br />

Notes A very distinctive and beautiful species. It is unusual in both habitat<br />

and flower for a Rebutia species. The free clumping nature of this<br />

species produces beautiful small mounds of white heads. The<br />

white flowers are small, but <strong>the</strong> unusual colour makes <strong>the</strong> plant a<br />

real gem.<br />

The majority of plants in cultivation appear to be self sterile and<br />

from <strong>the</strong> apparent lack of different clones, seed is rare. We believe<br />

that most of <strong>the</strong> plants in cultivation have been propagated<br />

vegetatively from <strong>the</strong> clone originally introduced by H. Winter of<br />

West Germany in <strong>the</strong> late 1950’s.<br />

Recently, an interesting hybrid has been produced with R. heliosa.<br />

The plants have <strong>the</strong> body form of R. heliosa but with white flowers.<br />

Kakt. u. a. Sukk. (1978) 29 184.<br />

REBUTIA ALBIPILOSA<br />

Rebutia albipilosa Ritter Taxon (1963) 12 29<br />

Synonyms Aylostera albipilosa (Ritt.) Backbg.<br />

Classification Section I Aylostera.<br />

Body up to 40mm. in diameter becoming caespitose with age. Epidermis<br />

dark green.<br />

Areoles elliptical, with white wool, raised on short tubercles and arranged<br />

on ribs.<br />

Spines c. 20 in number, strongly differentiated in centrals and radials,<br />

mostly 7-10mm. long. However, a few spines become extremely<br />

long especially at <strong>the</strong> base of <strong>the</strong> plant. These spines are often<br />

30-40mm. long. All spines are white, <strong>the</strong> longest ones having dark<br />

brown tips.<br />

28


Flowers 45mm. long and 30mm. wide, petals orange-red in colour often with<br />

a darker red midstripe. Tube and ovary with hairs and bristles.<br />

Filaments and style white with a 6 lobed cream coloured stigma.<br />

Self fertile.<br />

Fruit 5mm. in diameter, brownish-green and covered with hair and<br />

bristles.<br />

Seeds dome shaped, dull black.<br />

Type locality Narvaez, Department Tarcia, Bolivia. Map 4, No. 77.<br />

Notes This is ano<strong>the</strong>r of <strong>the</strong> Rebutia species found by Friedrich Ritter<br />

(FR754) in <strong>the</strong> early 1960’s. It is a very variable species particularly<br />

with reference to <strong>the</strong> length of <strong>the</strong> spines. Several o<strong>the</strong>r ‘species’<br />

are probably referable here for example R. vulpina (FR939) and R.<br />

robustispina (FR763). The original plant depicted by Ritter with<br />

very long and dense white spines appears to be an extreme form.<br />

Nobody has yet recollected or seen ano<strong>the</strong>r comparable plant.<br />

Donald (Ashingtonia 2 180) has speculated that Ritter’s R. flavistyla<br />

(FR756) is <strong>the</strong> true R. albipilosa. If this is correct and confirmed by<br />

Ritter it calls into question a whole series of descriptions and<br />

relationships. It is interesting to note that Ritter found and<br />

described both species, (see colour plate and fig. 1.).<br />

REBUTIA AUREIFLORA<br />

Fig. 7 An uncommon, slow growing species with very large flowers.<br />

29


REBUTIA AUREIFLORA<br />

Rebutia aureiflora Backbg. D. Kaktfrd. (1932) 124<br />

Synonyms Mediolobivia aureiflora (Backbg.) Backbg.<br />

Mediolobivia aureiflora var. albiseta Backbg.<br />

Mediolobivia aureiflora var. albilongiseta Backbg.<br />

Mediolobivia aureiflora var. albispina Backbg. nomen nudum<br />

Mediolobivia spiralisepala Schutz<br />

Rebutia aureiflora var. albilongiseta (Backbg.) Marshall & Bock,<br />

comb. nud.<br />

Rebutia aureiflora var. albiseta (Backbg.) Marshall & Bock. comb,<br />

nud.<br />

Rebutia aureiflora var. longiseta (Backbg.) Marshall & Bock. comb.<br />

nud.<br />

Rebutia aureiflora var. densispina Borg nomen nudum<br />

Rebutia aureiflora subspec. aureiflora (Backbg.) Donald<br />

Classification Section II Mediolobivia Subsection II Seti<strong>rebutia</strong>e<br />

Body 40mm. in diameter becoming caespitose from <strong>the</strong> base, eventually<br />

forming clumps 100mm. in diameter.<br />

Areoles elliptical with white wool<br />

Spines c. 20, extremely variable not strongly differentiated into centrals<br />

and radials. Seedlings and offshoots on older plants with short<br />

spines 6-8mm. long yellowish at first, becoming all white and<br />

tipped yellowish-brown. Older plants with extremely long thin<br />

flexuous spines 50mm. or more in length, white tipped yellowish-<br />

brown.<br />

Flowers very large, 50mm. long and 45mm. in diameter, orange-yellow<br />

petals, throat white, tube light brownish-orange with small<br />

greenish-brown scales long thin flexuous hair and bristles are<br />

produced up to 10mm. long. Stigma yellow, style and filaments<br />

white, self sterile.<br />

Fruit large, 8mm. in diameter brownish-orange densely covered with<br />

long hairs.<br />

Seeds dull brownish-black<br />

Type locality Quebrada del Toro, Salta, Argentina. Map 2, No. 55.<br />

Notes This species belongs to a group of slow growing plants which are<br />

self sterile. Plants are rarely offered for sale mainly because what<br />

seed is available germinates very erratically. In our experience<br />

10% germination is all that can be expected. There are fourteen<br />

described varieties and in all probability <strong>the</strong>y can all be included<br />

within <strong>the</strong> variation of <strong>the</strong> one species. For example var.<br />

albilongiseta has long pure white bristles, var. albiseta has 10mm.<br />

long white spines, var. boedeckeriana has 11-12 thin radial spines<br />

and pale orange flowers, var. rubelliflora with orange-red flowers,<br />

var. rubriflora with red flowers., In our opinion <strong>the</strong>se differences do<br />

not justify varietal status.<br />

30


REBUTIACOSTATA<br />

Fig. 8 A rare and slow growing species that is shy flowering. The plant in <strong>the</strong><br />

photograph has been in cultivation at Abbey Brook for over 20 years.<br />

REBUTIA COSTATA<br />

Rebutia costata Werd. Notizbl. Bot. Gart. u. Mus. Berlin (1934) 12 25<br />

Synonyms Lobivia costata (Werd.) Wessn.<br />

Digito<strong>rebutia</strong> costata (Werd.) Buin.<br />

Mediolobivia costata (Werd.) Krainz.<br />

Classification Section III Digito<strong>rebutia</strong> Sub Section II Euan<strong>the</strong>mae<br />

Body up to 25mm. in diameter simple at first but becoming caespitose<br />

with age. Epidermis deep bluish-green.<br />

Areoles elliptical with brownish wool, raised on tubercles arranged in 8-10<br />

distinct ribs.<br />

31


Spines centrals 0 (old areoles may have 1-2), radials 9-12 (14), pectinate<br />

and strongly projecting up to 7mm. long. White becoming<br />

yellowish-brown at <strong>the</strong> base.<br />

Flowers up to 35mm. long and 30mm. in diameter, deep orange, white throat<br />

buds green, tube greenish-brown with olive green scales and<br />

copiously hairy with a few long white bristles. Stigma cone shaped<br />

5 lobed, greenish-yellow style greenish filaments lavender pink.<br />

Seeds brownish.<br />

Type locality The type locality is unknown. It has recently been recollected by<br />

W. Rausch at La Quiaca, Jujuy, Argentina (R508).<br />

Notes This is a rare, slow growing species - <strong>the</strong> plant depicted in fig. 8 has<br />

been in cultivation for over 20 years and is still in a 3ins. pot. It is shy<br />

flowering and rarely sets seed.<br />

REBUTIA DEMINUTA<br />

Fig. 9 An easily flowered and widely grown species that was one of <strong>the</strong> first species<br />

to be discovered and described.<br />

32


REBUTIA DEMINUTA<br />

Rebutia deminuta (Web.) Br. & R.<br />

Synonyms Echinopsis deminuta Web. Bull Mus. Hist. Nat. Paris (1904) 386<br />

Echinocactus deminutus (Web.) Gürke<br />

echino<strong>rebutia</strong> deminuta (Web.) Frič & Krzgr.<br />

Aylostera deminuta (Web.) Backbg.<br />

Classification Section I Aylostera<br />

Body individual heads about 50-60mm. high, dark green. Plants freely<br />

offsetting from <strong>the</strong> base and eventually forming large clumps more<br />

than 120mm. in diameter.<br />

Areoles elliptical, light brownish.<br />

Spines 10-12, not strongly differentiated into centrals and radials. Spines<br />

white tipped brown and 5-6mm. long. Central when present<br />

usually all brown.<br />

Flowers very free flowering, flowers up to 40mm. long and 30mm. broad.<br />

Petals deep orange-red with pink filaments and an 8 lobed white<br />

stigma. Small greenish scales and white hairs present on flower<br />

tube. Self fertile.<br />

Fruit 5mm. in diameter, greenish-red in colour and covered with<br />

bristles.<br />

Seeds small, matt black.<br />

Type locality Trancas, Tucuman State, N. Argentina. Map 4, No. 85.<br />

Notes Easily flowered and widely grown species.<br />

f. pseudominuscula (Speg.) Buin. & Donald body grey-green, heads<br />

slightly smaller than <strong>the</strong> type, spines 7-14, longer, up to 8-9mm.,<br />

yellowish white, brown at base and tip. Flowers as in <strong>the</strong> type.<br />

REBUTIA EINSTEINII f. SCHMIEDCHENIANA<br />

Rebutia einsteinii f. schmiedcheniana (Köhler) Buin. & Donald<br />

Synonyms Lobivia schmiedcheniana Köhler Beitr. z. Skde. u. pflege, (1939) 37<br />

Cylindro<strong>rebutia</strong> schmiedcheniana (Köhler) Donald<br />

Mediolobivia schmiedcheniana (Köhler) Krainz.<br />

Rebutia schmiedickenana (Köhler) Marsh & Bock comb. nud.<br />

(should be Köhler and schmiedcheniana).<br />

Classification Section II Mediolobivia Sub Section II Cylindro<strong>rebutia</strong>e<br />

Body up to 35mm. in diameter becoming cylindrical with age. Some<br />

plants become caespitose but this is not always so. The epidermis<br />

is a dark brownish-green often tinged deep purple particularly<br />

during <strong>the</strong> winter.<br />

Areoles elliptical with sparse brownish wool raised on short tubercles<br />

spirally arranged.<br />

E<br />

33<br />

Hirt. -><br />

Hist.


Spines slightly differentiated into two series, 15 radials appressed,<br />

pectinate, 12 up to 75mm. long, greyish-white, brownish at <strong>the</strong><br />

base, upper 3 longer up to 18mm. and projecting. 1 (2) central<br />

spines to <strong>the</strong> upper three radials. Upper spines and centrals light<br />

brownish.<br />

Flowers 45mm. long and 35-40mm. in diameter, outer petals and sepals<br />

deep golden yellow. Stigma, style and filaments pale yellow tipped<br />

dark brownish yellow. Ovary and tube with copious white hair and<br />

long white bristles.<br />

Fruit large, 7mm, self fertile. Green tinged purple.<br />

Seed large, dome shaped, brownish.<br />

Type locality Cham volcano, Salta, N. Argentina. Map 4, No. 128.<br />

Notes This species has a very large underground tuberous root system<br />

and for this reason needs a gritty porous compost. Plants are most<br />

often seen grafted but this is unnecessary with careful cultivation.<br />

REBUTIA EINSTEINII f. SCHMIEDCHENIANUM<br />

Fig. 10 An uncommon species with a very large tuberous root system and large<br />

golden yellow flowers<br />

34


REBUTIA EUANTHEMA<br />

Fig. 11 An easily grown popular species with very showy flowers.<br />

REBUTIA EUANTHEMA<br />

Rebutia euan<strong>the</strong>ma (Backbg.) Buin. & Donald<br />

Synonyms Lobivia euan<strong>the</strong>ma Backbg. Blatt. f. Kakt. (1934) 2<br />

Mediolobivia euan<strong>the</strong>ma (Backbg.) Krainz.<br />

Acantholobivia euan<strong>the</strong>ma (Backbg.) Y. Ito<br />

Digito<strong>rebutia</strong> euan<strong>the</strong>ma (Backbg.) Buin.<br />

Echinolobivia euan<strong>the</strong>ma (Backbg.) Y. Ito<br />

Classification Section III Digito<strong>rebutia</strong> Subsection II Euan<strong>the</strong>mae<br />

Body heads 30mm. in diameter, grey-green often tinged amethyst, freely<br />

offsetting from <strong>the</strong> base and forming clumps 70-80mm. in diameter.<br />

Areoles elliptical, light brownish in colour.<br />

Spines often pectinate, about 10 radials, centrals absent Up to 10mm long,<br />

white tipped brown. Spines always slightly erect never appressed.<br />

This is one of <strong>the</strong> distinguishing features of Subsection II from<br />

subsection I Pygmaeae. (see analytical key to <strong>the</strong> Rebutia section)<br />

35


Flowers very free flowering, 45mm. long and 30mm. wide with orange-red<br />

petals. Flower tube very hairy with occasional bristles. Stigma and<br />

style greenish-yellow, 6-lobed, filaments lavender pink.<br />

Fruit brownish-red, densely covered with white hairs.<br />

Seeds dull brownish-black.<br />

Type locality given by Backeberg as South Bolivia - North Argentina. This<br />

species has recently been recollected by W. Rausch at Quebrada<br />

de Humahuaca Jujuy, Argentina (R214).<br />

Notes this is an easily grown popular species with very showy flowers. It<br />

is very closely related to a number of o<strong>the</strong>r species belonging to<br />

subsection II namely R. brachyantha and R. ritteri (see colour<br />

plate).<br />

REBUTIA FIEBRIGII<br />

Fig. 12 This photograph depicts Abbey Brook species No. CF2630 originally<br />

collected at Hacienda Ressini and having pure white spines and orange flowers.<br />

36


accomodate -><br />

accommodate<br />

REBUTIA FIEBRIGII<br />

Rebutia fiebrigii (Guerke) Br. & R.<br />

Synonyms Echinocactus fiebrigii Guerke: Notizbl. Bot. Gart. u. Mus. Berlin<br />

(1905)4,183<br />

Echino<strong>rebutia</strong> fiebrigii (Guerke) Frič & Kreuzgr.<br />

Aylostera fiebrigii (Guerke) Backbg.<br />

Rebutia nivosa Ritter nomen nudum<br />

Rebutia fiebrigii f. densiseta Cullm.<br />

Rebutia fiebrigii var. densiseta (Cullm.) Oeser<br />

Classification Section I Aylostera<br />

Body spherical becoming cylindrical with age, often 60mm. in diameter<br />

and 100- 120mm. high. Older plants offsetting from <strong>the</strong> base.<br />

Areoles prominent, elliptical, whitish in colour.<br />

Spines 30-40 bristle like radial white spines and 2-5 stiff centrals up to<br />

20mm. or more long, white tipped brown, sometimes pure white.<br />

Flowers 35mm. long and 30mm. broad, flame red with white stamens.<br />

Stigma 6 lobed pale yellow. Ovary and tube scaly with a few white<br />

hairs and bristles.<br />

Seed small, matt black.<br />

Type locality Iscayache, Bolivia 3600m. Map 4, No. 87.<br />

Notes Distinctive species and one of <strong>the</strong> first to be discovered and<br />

cultivated. Several recently described ‘species’ are probably<br />

referable here e.g. R. jujuyana, R. pulchella.<br />

R. fiebrigii, f. densiseta Cullm. (FR390) a form in which <strong>the</strong> spines<br />

are thinner and denser, spines tipped yellowish-brown, flowers<br />

orange. Originally collected at Hacienda Hura Khathalla ad<br />

Estancia Chujchi, Bolivia. Abbey Brook species No. CF2630, is a<br />

plant originally collected at Hacienda Ressini, Sucre by Gerhard<br />

Frank with longer spines than in <strong>the</strong> type, softer and all pure white,<br />

flowers clear orange. Attempts have been made to distinguish<br />

between <strong>the</strong> snow white populations and o<strong>the</strong>rs having brown<br />

tipped spines. As Donald has pointed out, <strong>the</strong> original description<br />

by Guerke suggests an almost wholly white spined plant, but it<br />

could accommodate both white spined and <strong>the</strong> more commonly seen<br />

brown tipped spined plants.<br />

REBUTIA HELIOSA<br />

Rebutia heliosa Rausch (1970) Kakt. u. a. Sukk. 21 30<br />

Synonyms Aylostera heliosa Rausch comb. nud.<br />

Classification Section I Aylostera<br />

Body solitary at first <strong>the</strong>n offsetting profusely, 20mm. high and 25mm. in<br />

diameter.<br />

Areoles 0-5mm. wide, 1mm. long, light brown, felty.<br />

37<br />

Notizke -><br />

Notizbl


Spines radials 24-26, 1mm. long, flattened against tubercles, diverted<br />

downwards, silvery-white, thickened and darker below. No<br />

central spines.<br />

Flowers 50mm. long and 25mm. in diameter, pale orange outer petals with<br />

purplish mid-stripe. The flowers have a very long thin tube (2mm.<br />

in diameter and 20mm. long) making <strong>the</strong> flower stand away from<br />

<strong>the</strong> plant body. Tube purplish, filaments white, stigma 6 lobed and<br />

style yellowish.<br />

Fruit tiny 3mm. in diameter with few seeds, (10-20) in each fruit, greenish<br />

later turning violet. Fruit covered with white hair like bristles.<br />

Seed black, spherical to helmet shaped with a papillate testa.<br />

Type locality Road between Tarija City and Narvaez at 2400-2500m., Bolivia.<br />

Map 4, No. 92.<br />

REBUTIA HELIOSA<br />

Fig. 13 One of <strong>the</strong> most beautiful, distinctive and unusual Rebutia species.<br />

38<br />

helmit -><br />

helmet


Varieties Rebutia heliosa var. cajasensis Lau 405<br />

Rebutia heliosa var. condorensis Lau 401 (Syn. R. solisioides Knize<br />

nomen nudum)<br />

Rebutia heliosa x Rebutia albiflora. An interesting new hybrid, (see<br />

under R. albiflora)<br />

Notes One of <strong>the</strong> most beautiful and unusual Rebutia species, very<br />

distinctive. The plant seems to grow well on its own roots but plants<br />

are most often seen grafted which does tend to make <strong>the</strong> plants<br />

elongated and proliferate.<br />

Several ‘species’ are referable here, namely, supthutiana,<br />

albopectinata, perplexa, schatzliana and densipectinata. For an<br />

explanation of <strong>the</strong> confusion amongst <strong>the</strong>se species an article in<br />

Ashingtonia (1979) 3 by John Donald helps to sort out <strong>the</strong> situation.<br />

REBUTIA KRAINZIANA<br />

Fig. 14 This is a superb and distinctive plant with very large flowers.<br />

39


REBUTIA KRAINZIANA<br />

Rebutia krainziana Kesselring: Sukkulentenkunde (1948) II, 23-24<br />

Synonyms Rebutia calliantha var. densiseta Bew. nomen nudum<br />

Rebutia calliantha var. krainziana (Kess.) Buin. & Donald<br />

Rebutia wessneriana var. densiseta nomen nudum<br />

Rebutia wessneriana var. krainziana (Kess.) Buin. & Donald<br />

Classification Section IV Rebutia Subsection II Medio<strong>rebutia</strong>e<br />

Body up to 50mm. in diameter and becoming caespitose with age<br />

forming large clumps 100mm. or more in diameter.<br />

Areoles very prominent, ellipitical and with white felt.<br />

Spines pectinate, adpressed, up to 12 radials pure white, centrals absent.<br />

Flowers very large and beautiful, 40mm. long and 40mm. wide. Usually dark<br />

red but <strong>the</strong>re is a form (f. breviseta) with clear orange flowers. 5<br />

lobed stigma, style white, filaments yellow. Self sterile.<br />

Fruit large, 8mm. in diameter with small brownish red scales.<br />

Seeds black shiny, dome-shaped with white basal hylum area. Testa near<br />

hylum reticulate but becoming tuberculate at <strong>the</strong> apex.<br />

Type locality is unknown. It has recently been recollected at Yavi, Jujuy,<br />

Argentina. Map 3, No. 70<br />

Notes This is a superb and striking plant with very large beautiful flowers.<br />

When not in flower <strong>the</strong> plant is one of <strong>the</strong> most attractive species.<br />

R. krainziana f. breviseta with orange flowers may be a hybrid<br />

between R. krainziana and R. marsoneri. The following<br />

combinations also belong with this taxa.<br />

R. krainziana var. breviseta (Backbg.) Donald<br />

R. krainziana f. breviseta (Backbg.) Krainz & Haarm.<br />

R. senilis v. breviseta Backbg.<br />

R. senilis f. breviseta (Backbg.) Backbg.<br />

REBUTIA KUPPERIANA<br />

Rebutia kupperiana Böd.: Monatss. D.K.G. (1932) 276-277<br />

Synonyms Aylostera kupperiana (Böd.) Backbg.<br />

Echino<strong>rebutia</strong> kupperiana (Böd.) Frič & Krzgr.<br />

Classification Section I Aylostera<br />

Body up to 40mm. in diameter and becoming caespitose with age.<br />

Epidermis very dark brownish-green.<br />

Areoles circular with brownish wool, raised on short tubercles.<br />

Spines strongly differentiated into centrals and radials. Centrals 2, upper<br />

12-15mm. long, lower 20mm. long at first dark chocolate brown in<br />

age becoming greyish with a blackish tip. Radials 10-12, strongly<br />

projecting, 6-12mm. long, somewhat flexuous, white with dark<br />

brown tips.<br />

40


Flowers very large and beautiful 50mm. long and 40mm. in diameter, dark<br />

orange-red. Tube 15mm. long greenish with long bristles. Stigma 6<br />

lobed, filaments and style white.<br />

Fruit greenish, 5mm. in diameter.<br />

Seeds greyish black.<br />

Type locality near Tarija at 2500m. altitude, Bolivia. Map 4, No. 99.<br />

Notes This is yet ano<strong>the</strong>r magnificent species discovered by Friedrich<br />

Ritter in 1931.<br />

Varieties Rebutia kupperiana var. spiniflora Ritter (FR762b)<br />

REBUTIA KUPPERIANA<br />

Fig. 15 This is yet ano<strong>the</strong>r magnificent species discovered by Friedrich Ritter. It<br />

has dark brown spines and very large dark orange-red flowers.<br />

41<br />

7626 —><br />

762b


REBUTIA MARSONERI<br />

Fig. 16 Ano<strong>the</strong>r superb species with golden yellow flowers<br />

REBUTIA MARSONERI<br />

Rebutia marsoneri Werdermann: Kakteenkunde (1937) 2<br />

Synonyms Rebutia marsoneri var. albescens hort. nomen nudum<br />

Rebutia marsoneri var. breviseta hort. nomen nudum<br />

Rebutia marsoneri var. brevispina Donald nomen nudum<br />

Rebutia marsoneri var. grandiflora Donald nomen nudum<br />

Rebutia marsoneri var. spathulata Donald nomen nudum<br />

Rebutia marsoneri f. spathulata Donald nomen nudum<br />

Rebutia marsoneri var. vatteri Donald nomen nudum<br />

Rebutia peruviana hort. nomen nudum<br />

Classification Section IV Rebutia Subsection II Medio<strong>rebutia</strong>e<br />

Body up to 70mm. in diameter becoming caespitose with age.<br />

42


Areoles ellipitical with dense white wool.<br />

Spines very variable in size and number, c. 10 undifferentiated into<br />

centrals and radials, up to 15mm. long, whitish or tinged yellow. In f.<br />

brevispina all <strong>the</strong> spines are pectinate, white and less than 5mm.<br />

long.<br />

Flowers large, 40mm. long and 35mm. in diameter. Usually golden yellow<br />

but sometimes lighter yellow, sometimes orange-yellow. Stigma<br />

5-10 lobed, style and filaments golden yellow. The buds<br />

are variable in colour sometimes red and sometimes strikingly<br />

bright green.<br />

Fruit large, 8mm. in diameter.<br />

Seeds black, shiny, dome-shaped with white basal hylum area.<br />

Type locality North Jujuy, Argentina. Map 3, No. 71.<br />

Notes A superb species named after Oreste Marsoner, an Italian who has<br />

collected extensively in Argentina. This species is relatively rare<br />

in cultivation because it is self sterile and seed is not as readily<br />

available as self fertile species. There are a number of varieties<br />

and forms most of which are insufficiently different to warrant<br />

separate classification, (see colour plate).<br />

REBUTIA MINUSCULA<br />

Fig. 17 This is one of <strong>the</strong> easiest and most widely grown of all species of Cacti.<br />

43<br />

ospecies<br />

deleted


REBUTIA MINUSCULA<br />

Rebutia minuscula K. Sch.: Monats. f. Kakt. (1895) 5, 102<br />

Synonyms Echinopsis minuscula Web.<br />

Echinocactus minuscula Web.<br />

Rebutia minuscula v. multiflora Frič nomen nudum<br />

Rebutia petersonii Hort. nomen nudum<br />

Classification Section IV Rebutia Subsection I Rebutiae<br />

Body up to 50mm. in diameter and becoming strongly caespitose with<br />

age soon forming large clumps 150mm. or more in diameter.<br />

Epidermis deep green.<br />

Areoles circular with sparse white wool.<br />

Spines numerous, c. 40, undifferentiated into radials and centrals and<br />

arranged like needles on a pin cushion. Spines up to 5mm. long,<br />

whitish, sometimes becoming brownish at <strong>the</strong> base.<br />

Flowers large, up to 40mm. long and 35mm. in diameter, bright red. 6 lobed<br />

stigma, style pinkish, filaments golden yellow.<br />

Fruit 6mm. in diameter, orange-red.<br />

Seeds black, shining.<br />

Type locality Tucuman, N. Argentina. Map 3, No. 61.<br />

Notes One of <strong>the</strong> easiest and most widely grown of all species of Cacti. It<br />

can be flowered when only 1 year old from seed. It was <strong>the</strong> first<br />

species of Rebutia to be described.<br />

There is a form in cultivation R. minuscula f. grandiflora which has a<br />

very elongated flower tube making <strong>the</strong> flower 70mm. long.<br />

Consult <strong>the</strong> section on flowers and floral structure.<br />

REBUTIA MUSCULA<br />

Rebutia muscula Ritter & Thiele: Taxon (1963) 12,29<br />

Synonyms Aylostera muscula (Ritter & Thiele) Backbg.<br />

Rebutia nivosa Ritter nomen nudum<br />

Rebutia muscula v. nivosa Knize comb. nud.<br />

Classification Section I Aylostera<br />

Body up to 5cm. in diameter becoming caespitose with age, individual<br />

heads becoming elongated.<br />

Areoles circular with white wool.<br />

Spines pure white, soft bristle like, centrals indistinguishable from <strong>the</strong><br />

radials. 6mm. long, 50-55 in number.<br />

44


Flowers 40mm. long and 25mm. in diameter, sepals greenish-orange, petals<br />

flame orange, throat white. Stigma very pale green, style white,<br />

filaments white. Flower tube and ovary with very small scales. Very<br />

thin flexuous white hairs up to 8mm. long produced from <strong>the</strong> axils of<br />

<strong>the</strong> scales.<br />

Seeds small, dull black, dome shaped.<br />

Type locality Narvaez, Dept. Tarija, Bolivia. Map 4, No. 103.<br />

Notes Beautiful pure white spined species with gorgeous clear orange<br />

flowers. Flowers are often produced at odd times during <strong>the</strong> year<br />

but <strong>the</strong> main flowering time is at <strong>the</strong> end of May beginning of June.<br />

Plants labelled as R. nivosa are referable here, <strong>the</strong>re is no distinct<br />

difference and it was found at <strong>the</strong> same locality.<br />

(see illustration on front cover)<br />

REBUTIA PSEUDODEMINUTA<br />

Rebutia pseudodeminuta Backbg. non Frič (1933) D.K.F. 7<br />

Synonyms Aylostera pseudodeminuta (Backbg.) Backbg.<br />

Classification Section I Aylostera<br />

Body globular up to 10cm. high, 5-6cm. wide, light green in colour,<br />

offsetting grousely at first but later forming large clumps 100mm. or<br />

more in diameter.<br />

Areoles small with sparse light brown wool.<br />

Spines radials ten or more, 3-7mm. long, glossy white. Centrals 2-3,<br />

yellowish-white at first later turning brown-tipped, up to 15mm.<br />

long.<br />

Flowers 30mm. long and 30mm. wide, petals red, buds dark red. Style and<br />

filaments white united with <strong>the</strong> tube, 5 lobed stigma.<br />

Fruit round, small with many bristles.<br />

Seed very small, matt black.<br />

Habitat Salta, N. Argentina. Map 4, No. 107.<br />

Varieties f. albiseta (Backbg.) Buin. & Donald. Central spines entirely white.<br />

f. grandiflora (Backbg.) Buin. & Donald. Flower 4cm. wide and<br />

40mm. long.<br />

f. schneideriana (Backbg.) Buin. & Donald. Central spines up to<br />

35mm. long, dark tipped.<br />

f. schumanniana (Backbg.) Buin. & Donald. Central spines golden<br />

brown, about 12mm. long.<br />

Notes Ano<strong>the</strong>r of <strong>the</strong> ‘original’ Rebutias, quite distinct with its prominent<br />

tubercles and spination.<br />

45


REBUTIA PYGMAEA<br />

Fig. 18 A distinctive species with pectinate spines. It has very large flowers when<br />

compared to <strong>the</strong> size of <strong>the</strong> plants.<br />

REBUTIA PYGMAEA<br />

Rebutia pygmaea (R. E. Fries) Br. & R.<br />

Synonyms Echinopsis pygmaea R. E. Fries Nov. Act. Soc. Sci. Upsala (1905) IV<br />

120<br />

Acantholobivia haagei (Frič & Schelle) Y. Ito.<br />

Digito<strong>rebutia</strong> digitiformis (Backbg.) Buin.<br />

Digito<strong>rebutia</strong> haagei (Frič & Schelle) Frič<br />

Digito<strong>rebutia</strong> haagei var. digitiformis (Backbg.) Donald<br />

Digito<strong>rebutia</strong> haagei var. orurensis (Backbg.) Donald<br />

Digito<strong>rebutia</strong> haagei var. pectinata (Backbg.) Donald<br />

Digito<strong>rebutia</strong> orurensis (Backbg.) Buin. oreurensis -><br />

orurensis<br />

46


Digito<strong>rebutia</strong> pectinata (Backbg.) Buin.<br />

Digito<strong>rebutia</strong> pygmaea (R. E. Fries) Donald<br />

Digito<strong>rebutia</strong> steinmannii sensu Buin.<br />

Echinolobivia haagei (Frič & Schelle) Y. Ito.<br />

Lobivia neo-haageana Backbg.<br />

Lobivia pygmaea (R. E. Fries) Backbg.<br />

Mediolobivia haagei (Frič & Schelle) Backbg.<br />

Mediolobivia haagei var. digitiformis (Backbg.) Donald<br />

Mediolobivia haagei var. orurensis (Backbg.) Donald<br />

Mediolobivia haagei var. pectinata (Backbg.) Donald<br />

Mediolobivia haageana Borg nomen nudum<br />

Mediolobivia orurensis (Backbg.) Backbg.<br />

Mediolobivia pygmaea (R. E. Fries) Backbg.<br />

Mediolobivia pectinata (Backbg.) Backbg. non Frič<br />

Mediolobivia pectinata var. digitiformis (Backbg.) Backbg.<br />

Mediolobivia pectinata var. orurensis (Backbg.) Backbg.<br />

Mediolobivia pectinata var. neosteinmannii Backbg.<br />

Pygmaeolobivia odierata Hort. nomen nudum<br />

Pygmaeolobivia pygmaea (R. E. Fries)<br />

Rebutia haagei Frič & Schelle<br />

Rebutia orurensis Backbg.<br />

Rebutia pectinata Backbg.<br />

Rebutia pectinata var. digitiformis Backbg.<br />

Rebutia pectinata var. orurensis Backbg.<br />

Classification Section III Digito<strong>rebutia</strong> Subsection I Pygmaeae<br />

Body up to 25mm. in diameter, simple at first but becoming caespitose<br />

with age. The epidermis bluish-green often tinged deep purple<br />

particularly during <strong>the</strong> winter.<br />

Areoles elliptical with sparse greyish-brown wool, raised in <strong>the</strong> tubercles.<br />

Spines arranged in 12 distinct ribs. Centrals absent, radials strongly<br />

pectinate and adpressed, up to 12 in number, 4mm. long, greyish-<br />

white, brownish at <strong>the</strong> base.<br />

Flowers very variable, pink to salmon often bicoloured, 25mm. long and<br />

20mm. in diameter with a white throat. Stigma 6 lobed green, style<br />

green, filaments white or pinkish. Tube brownish-green with<br />

green scales with copious white wool.<br />

Fruit dull brownish-black, dome shaped, testa finely pitted.<br />

Seed large, 7mm. in diameter, greenish-brown covered with white wool.<br />

Type locality Yavi, Jujuy Province, N. Argentina. Map 2, No. 35.<br />

Notes This species has a very wide geographical range and a large<br />

number of minor variations occur. Many of <strong>the</strong>se minor variations<br />

have been described as ‘new’ species or varieties. There are for<br />

example a very large number of flower colour variants and also<br />

spine colour variants. It is indeed possible that this species could<br />

include 90% of all <strong>the</strong> species ascribed to <strong>the</strong> subsection<br />

Pygmaeae!<br />

47


REBUTIA SENILIS f. STUEMERI<br />

Rebutia senilis f. stuemeri (Backbg.) Buin. & Donald<br />

Synonyms Rebutia senilis v. stuemeri Backbg. D. Kakt. frd. (1932) 131<br />

Rebutia senilis v. stuemeriana Backbg.<br />

Classification Section IV Rebutia Subsection I Rebutiae<br />

Body up to 60mm. in diameter and becoming caespitose with age.<br />

Areoles elliptical with white wool.<br />

Spines pure white, stiff bristle like, centrals hardly distinguishable from<br />

<strong>the</strong> radials. 15-30 in number, size is variable - at <strong>the</strong> top of <strong>the</strong><br />

plant spines up to 18mm. long. Older areoles at <strong>the</strong> base of <strong>the</strong> plant<br />

produce longer spines up to 25mm. long.<br />

Flowers 50mm. long and 40mm. in diameter, clear red with a yellowish<br />

throat. Stigma 5 lobed, style and filaments golden yellow. Self<br />

fertile.<br />

Fruit yellowish, 7mm. in diameter.<br />

Seed black, shiny, elongated dome-shaped with a white basal hylum<br />

area. Testa near hylum more or less reticulate but becoming<br />

tuberculate at <strong>the</strong> apex.<br />

Type locality Quebrada Escoipe, Salta, Argentina. Map 3, No. 66.<br />

Notes R. almeyeri W. Heinr. probably belongs here. In Backbg. (1977) it is<br />

described as a self fertile species with orange-red flowers with a<br />

yellowish throat and a yellow style, locality unknown.<br />

REBUTIA SPINOSISSIMA<br />

Rebutia spinosissima Backbg. (1935) Kaktus ABC., 275<br />

Synonyms Aylostera spinosissima (Backbg.) Backbg.<br />

Aylostera spinosissima var. brunispina Backbg. nomen nudum<br />

Classification Section I Aylostera<br />

Body up to 35mm. in diameter becoming caespitose with age eventually<br />

forming large clumps.<br />

Areoles circular to elliptical with greyish wool.<br />

Spines soft and bristle like, about 20, not strongly differentiated into<br />

centrals and radials. ‘Radial’ spines up to 6mm. long, all white,<br />

‘centrals’ at first yellow tipped brown later all brown.<br />

Flowers 40mm. long and 30mm. in diameter. Petals orange-red, tube<br />

greenish, 8 lobed stigma, filaments and style white.<br />

Fruit greenish brown 4mm. in diameter with white hairs and bristles.<br />

Seeds elongated dome shaped, dull brownish black.<br />

Type locality Salta Province, N. Argentina. Map 4, No. 116.<br />

Notes This is a beautiful species with soft bristle like spines and orange<br />

flowers. It is slow growing and an asset to any Rebutia collection.<br />

Closely related to R. muscula and R. fiebrigii<br />

48


REBUTIA SPINOSISSIMA<br />

Fig. 19 A slow growing species with soft bristle-like spines and orange flowers.<br />

REBUTIA WESSNERIANA<br />

Rebutia wessneriana Bew. Sukkulentenkunde Jahrb. Schweiz.<br />

Kakt. Ges. (1948)2 24<br />

Synonyms Rebutia calliantha Bew.<br />

Rebutia hyalacantha (Backbg.) Backbg.<br />

Rebutia senilis var. hyalacantha Backbg.<br />

Rebutia senilis f. hyalacantha (Backbg.) ex Simon<br />

Rebutia wessneriana var. calliantha (Bew.) Krainz<br />

Rebutia wessneriana var. permutata (Heinr.) Buin. & Donald<br />

Rebutia wessneriana subspec. wessneriana f. calliantha (Bew.)<br />

Buin. & Donald<br />

49<br />

Schwiez<br />

-> Schweiz


REBUTIA WESSNERIANA<br />

Fig. 20 A superb large growing, large flowering species<br />

Rebutia wessneriana subspec. wessneriana var. gokrausei (Heinr.)<br />

Donald<br />

Rebutia wessneriana subspec. wessneriana var. gokrausei f.<br />

permutata (Heinr.) Donald<br />

Classification Section IV Rebutia Subsection II Medio<strong>rebutia</strong>e<br />

Body large, up to 100m. in diameter, caespitose and forming large<br />

clumps with age<br />

Areoles elliptical raised up on tubercles with white wool<br />

Spines not differentiated into centrals<br />

50<br />

gokausei -><br />

gokrausei


Flowers very large, 45mm. long and 40mm. in diameter. Petals dark red,<br />

buds dark purplish-red. Stigma 6 lobed, style yellowish-white,<br />

filaments orange. Self sterile.<br />

Fruit large, 8mm. in diameter, yellowish<br />

Seed black, shiny, dome shaped with white basal hylum area.<br />

Type locality Quebrada de Humahuaca, Jujuy, Arg.? Map 3, No. 74.<br />

Notes A superb large growing large flowered species which is<br />

uncommon in collections because <strong>the</strong> plant is self sterile and seed<br />

is <strong>the</strong>refore not readily available. Originally R. wessneriana was<br />

separated from R. calliantha by flower colour, scarlet in calliantha<br />

and bluish-red in wessneriana, and on <strong>the</strong> observation that <strong>the</strong><br />

spines cover <strong>the</strong> stem apex in calliantha but did not hide <strong>the</strong> stem<br />

apex in wessneriana. Such minor differences are not sustained and<br />

are only observed in extreme plants and are not exhibited by <strong>the</strong><br />

population as a whole. (see colour plate).<br />

REBUTIA XANTHOCARPA f. SALMONEA<br />

Rebutia xanthocarpa f. salmonea (Frič ex Backbg.) Buin. & Donald<br />

Synonyms Rebutia xanthocarpa var. salmonea Frič ex Backbg. (1951) Cact.<br />

Succ. J. Amer. 23 83<br />

Rebutia salmonea Frič non Backbg. nomen nudum<br />

Rebutia senilis var. salmonea (Backbg.) Marsh.<br />

Classification Section IV Rebutia Subsection I Rebutiae<br />

Body 40mm. in diameter, very slowly becoming caespitose with age.<br />

Areoles elliptical with greyish brown felt.<br />

Spines not differentiated into centrals and radials, c. 15. Short stiff and<br />

bristle like, 2mm. long occasionally 4mm., white tinged yellow.<br />

Flowers small, 15mm. long and 15mm. in diameter. A beautiful salmon pink.<br />

Tube yellow with greenish yellow scales. Stigma 6 lobed, yellowish<br />

white. Style and filaments white.<br />

Fruit large 7mm. in diameter, golden yellow, (not pale red, Backbg.<br />

Lexicon P. 439)<br />

Seeds small, black shining with white basal hylum area.<br />

Habitat unknown, but probably Salta area, Argentina. Map 3, No. 67.<br />

Notes An easily grown and flowered plant with a very characteristic and<br />

distinctive flower colour. (see colour plate).<br />

51<br />

Humuhuaca -><br />

Humahuaca


Check list of Genera, Subgenera and Sections<br />

The following is a list of <strong>the</strong> more important genera to which members of <strong>the</strong><br />

<strong>genus</strong> Rebutia have been assigned. There are in addition to <strong>the</strong> following list a<br />

number of additional generic names which have not been published in accordance<br />

with <strong>the</strong> code of Botanical Nomenclature and which are probably best forgotten. The<br />

correct Rebutia section for <strong>the</strong> plants which were originally placed under <strong>the</strong>se<br />

names is indicated in brackets after <strong>the</strong> name. It follows <strong>the</strong> taxonomic arrangement<br />

used in this booklet. The original publication and date is given for <strong>the</strong> most important<br />

of <strong>the</strong>se generic names.<br />

Acantholobivia Y. Ito. non Backbg. (Rebutia Section III) (1957) Expl. Diagr. 130<br />

Aylostera Speg. (Rebutia Section I) (1923) An. Soc. Cient. Argent. 96 75<br />

Cylindro<strong>rebutia</strong> Frič & Kreuzgr. nomen nudum (Rebutia Section II) (1938) Succulenta<br />

20 55 & 71<br />

Cylindro<strong>rebutia</strong> Buin. & Donald pro. sect. (Rebutia Section II) (1963)<br />

Sukkulentenkunde Jb. Schweiz Kakt. Ges. 8 98<br />

Digito<strong>rebutia</strong> Frič & Kreuzgr. nomen nudum (Rebutia Section III) (1938) Succulenta<br />

20 54<br />

Digito<strong>rebutia</strong> Frič & Kreuzgr. ex. Buin. (Rebutia Section III) (1940) Succulenta 22 51<br />

Digito<strong>rebutia</strong> Buin. & Donald pro. sect. (Rebutia Section III) (1963) Sukkulentenkunde<br />

Jb. Schweiz Kakt. Ges. 8 98<br />

Echinocactus Link & Otto (Rebutia Section 1, 11 and IV) (1827) Verh. Ver. Beford.<br />

Gartenb. 3 420<br />

Echinocactus Weber, ex K. Sch. (Rebutia Section 1,11 and IV) (1898) Gesamtbeschr.<br />

Kakt. 396<br />

Echinolobivia Y. Ito. (Rebutia Section III) (1957) Expl. Diagr.<br />

Echinopsis Zucc. (Rebutia Section 1, 11 and IV) (1837) Abh. Bayer. Akad. Wiss.<br />

München 2 675<br />

Echino<strong>rebutia</strong> Frič & Kreuzgr. nomen nudum (Rebutia Section I) (1935) Verzeichnis<br />

26<br />

Lobivia Br. & R. (Rebutia Section 11 and III) (1922) The Cactaceae III 49<br />

Lobivia subgen. Pygmaeolobivia Backbg. nomen nudum (Rebutia Section II) (1934)<br />

Blatt. f. Kakteenf.<br />

Mediolobivia Backbg. (Rebutia Section I, II and III) (1934) Blatt. f. Kakteenf.<br />

Mediolobivia subgen. Eumediolobivia Backbg. (Rebutia Section II) (1942) Cact. Jb.<br />

D.K.G. 1135<br />

Mediolobivia subgen. Pygmaeolobivia Backbg. (Rebutia Section III) (1941) Cact. Jb.<br />

D.K.G.<br />

Medio<strong>rebutia</strong> hort. nomen nudum (Rebutia Section IV)<br />

Medio<strong>rebutia</strong> Buin. & Donald pro, sect. (Rebutia Section IV)<br />

Rebulobivia Frič & Kreuzgr. (Rebutia Section II) (1936) Succulenta 18 104<br />

Rebutia K. Sch. (1895) Monatsschr. f. Kakteenfreunde 5 102<br />

Rebutia K. Sch. emend Buin. & Donald (1963) Sukkulentenkunde 7 96<br />

Rebutia subgen. Eu<strong>rebutia</strong> Backbg. non Frič. (Rebutia Section IV) (1934) Blatt. f.<br />

Kakteenf.<br />

Rebutia subgen. Neo<strong>rebutia</strong> Bew. (Rebutia Section IV) (1949) Sukkulentenkunde Jb.<br />

Schweiz Kakt. Ges. 3 54<br />

Seti<strong>rebutia</strong> Frič & Kreuzgr. nomen nudum (Rebutia Section II) (1935) Verzeichnis 26<br />

52<br />

Coc -> Soc<br />

Gesamtbexhr -><br />

Gesamtbeschr<br />

Beyer -><br />

Bayer<br />

Monatschr -><br />

Monatsschr


Seti<strong>rebutia</strong> Frič & Kreuzgr. (Rebutia Section II) (1938) Succulenta 20 53<br />

Seti<strong>rebutia</strong> Buin. & Donald pro. sect. (Rebutia Section II) (1963) Sukkulentenkunde Jb.<br />

Schweiz Kakt. Ges. 7 98<br />

Check list of Genus Aylostera Speg.<br />

albiareolata Ritter comb. nud. = R. albiareolata<br />

albiflora (Ritt. & Buin.) Backbg. = R. albiflora<br />

albipilosa (Ritt.) Backbg. = R. albipilosa<br />

albopectinata Rausch comb. nud. = R. albopectinata<br />

buiningiana Rausch comb, nud. = R. buiningiana<br />

buiningiana Ritter nomen nudum (not R. buiningiana Rausch) = R. spinosissima<br />

caespitosa Frič nomen nudum = ?<br />

camarguensis Rausch comb. nud. = R. camarguensis<br />

deminuta = R. deminuta<br />

deminuta var. pseudominuscula = R. deminuta f. pseudominuscula<br />

densipectinata Ritter comb. nud. = R. densipectinata<br />

diersiana Rausch comb. nud. = R. diersiana<br />

fiebrigii (Guerke) Backbg. = R. fiebrigii<br />

fiebrigii var. castanea Rausch nomen nudum= R. fiebrigii<br />

fiebrigii var. densiseta Cullm. = R. fiebrigii f. densiseta<br />

fiebrigii var. spinosior nomen nudum = R. fiebrigii<br />

flavistyla Ritter comb. nud. = R. flavistyla<br />

fulviseta Rausch comb. nud. = R. fulviseta<br />

fulviseta var. albispina Rausch nomen nudum<br />

graciliflora (Backbg.) Rausch comb. nud. = R. graciliflora<br />

grandiflora Frič nomen nudum = ? R. pseudodeminuta f. grandiflora<br />

heliosa Rausch nomen comb. = R. heliosa<br />

hoffmanii Diers. et Rausch comb. nud. = R. hoffmannii<br />

huasiensis Rausch comb. nud. = R. huasiensis<br />

ithyacantha Card. = R. ithyacantha<br />

jujuyana Rausch comb. nud. = R. jujuyana<br />

kieslingii Rausch comb. nud. = R. kieslingii<br />

kruegerii Card. = Sulco<strong>rebutia</strong> kruegerii<br />

kupperiana (Böd.) Backbg. = R. kupperiana<br />

lampromeliana Rausch nomen nudum = R. pulchella<br />

leucan<strong>the</strong>ma Rausch comb. nud. = R. leucan<strong>the</strong>ma<br />

mamillosa Rausch comb. nud. = R. mamillosa<br />

maxima Ritter nomen nudum = ?<br />

muscula (Ritt. & Thiele) Backbg. = R. muscula<br />

narvaecense Card. = R. narvaecense<br />

odontopetala Ritter comb. nud. = R. odontopetala<br />

padcayensis Rausch comb. nud. = R. padcayensis<br />

patericalyx Ritter comb. nud. = R. patericalyx<br />

pectinata Frič nomen nudum = R. pygmaea<br />

pseudodeminuta (Backbg.) Backbg. = R. pseudodeminuta<br />

pseudodeminuta var. albiseta Backbg. = R. pseudodeminuta f. albiseta<br />

pseudodeminuta var. grandiflora Backbg. = R. pseudodeminuta f. grandiflora<br />

53


pseudodeminuta f. rubrifilamentosa Buin. & Donald comb. nud. =<br />

R. pseudodeminuta f. rubrifilamentosa<br />

pseudodeminuta var. schneideriana Backbg. = R. pseudodeminuta f. schneideriana<br />

pseudodeminuta var. schumanniana (Backbg.) Backbg. =<br />

R. pseudodeminuta f. schumanniana<br />

pseudominuscula (Speg.) Speg. = R. deminuta f. pseudominuscula<br />

pulchella Rausch comb. nud. = R. pulchella<br />

pulvinosa (Ritt. & Buin.) Backbg. = R. pulvinosa<br />

robustiflora Frič = ?<br />

robustispina (Ritt.) Backbg. = R. robustispina<br />

rubiginosa (Ritt.) Backbg. = R. rubiginosa<br />

sanguinea Ritter comb. nud. = R. sanguinea<br />

schatzliana Rausch comb. nud. = R. schatzliana<br />

sordida Rausch nomen nudum. = ?<br />

spegazziniana (Backbg.) Backbg. = R. spegazziniana<br />

spegazziniana var. atroviridis Backbg. = R. spegazziniana var. atroviridis<br />

spegazziniana var. wal<strong>the</strong>riana Backbg. nomen nudum = R. spegazziniana<br />

spinosissima (Backbg.) Backbg. = R. spinosissima<br />

spinosissima var. aurea Rausch nomen nudum = R. spinosissima<br />

spinosissima var. brunispina = R. spinosissima<br />

spinosissima var. buiningiana Ritter nomen nudum = R. spinosissima<br />

steinmannii (Solms-Laub.) Backbg. = R. steinmannii<br />

[not R. pygmaea (R. E. Fries) Br. & R.!]<br />

supthutiana Rausch comb. nud. = R. supthutiana<br />

tarijensis Rausch comb. nud. = R. tarijensis<br />

tuberosa (Ritter) Backbg. = R. tuberosa<br />

umbraculiforma Rausch nomen nudum = R. camarguensis<br />

vallegrandensis Card. = R. vallegrandensis<br />

wal<strong>the</strong>riana (Backbg.) Y. Ito. = ? R. spegazziniana<br />

zavaletae Card. = Sulco<strong>rebutia</strong> zavaletae (Card.) Backbg.<br />

Check list of Genus Digito<strong>rebutia</strong> Frič. Kreuzgr. Buin.<br />

albopectinata Rausch comb. nud. = R. albopectinata<br />

atrovirens (Backbg.) = R. pygmaea var. atrovirens<br />

brachyantha (Wessn.) Buin. = R. brachyantha<br />

brachyantha var. ritteri (Wessn.) Donald = R. ritteri<br />

brunescens Rausch comb. nud. = R. brunescens<br />

canacruzensis Rausch comb. nud. = R. canacruzensis<br />

christinae Rausch comb. nud. = R. christinae<br />

cincinnata Rausch comb. nud. = R. cincinnata<br />

costata (Werd.) Buin. = R. costata<br />

costata var. eucalyptana (Backbg.) Donald = R. costata f. eucalyptana<br />

diersiana Rausch comb. nud. = R. diersiana<br />

diersiana var. atrovirens Rausch comb. nud. = R. diersiana v. atrovirens<br />

digitiformis (Backbg.) Buin. = R. pygmaea<br />

donaldiana Rausch nomen nudum = R. steinmannii v. cincinnata<br />

Rausch nomen nudum (R300)<br />

54<br />

steinmanii -><br />

steinmannii


eucalyptana (Backbg.) Buin. = R. costata f. eucalyptana<br />

euan<strong>the</strong>ma (Backbg.) Buin. = R. euan<strong>the</strong>ma<br />

euan<strong>the</strong>ma var. friciana Donald & Cullm. comb. nud. = R. euan<strong>the</strong>ma f. fricii<br />

euan<strong>the</strong>ma var. longispina (Backbg.) Donald comb. nud. = R. euan<strong>the</strong>ma<br />

euan<strong>the</strong>ma var. oculata (Werd.) Donald = R. euan<strong>the</strong>ma f. oculata<br />

haagei (Frič & Schelle) Frič = R. pygmaea<br />

haagei var. digitiformis (Backbg.) Donald = R. pygmaea<br />

haagei var. atrovirens (Backbg.) Donald = R. pygmaea v. atrovirens<br />

haagei var. orurensis (Backbg.) Donald = R. pygmaea<br />

haagei var. pectinata (Backbg.) Donald = R. pygmaea<br />

haefneriana (Cullm.) Donald = R. pygmaea f. haefneriana<br />

mudanensis Rausch comb. nud. = R. mudanensis<br />

nazarenoensis Rausch (1979) Succulenta 58 185.<br />

nigricans (Wessn.) Buin. = R. ritteri var. nigricans<br />

orurensis (Backbg.) Buin. = R. pygmaea<br />

oculata (Werd.) Buin. = R. euan<strong>the</strong>ma f. oculata<br />

oculata var. friciana Donald & Cullm. = R. euan<strong>the</strong>ma f. fricii<br />

pectinata (Backbg.) Buin. = R. pygmaea<br />

peterseimii (Frič & Kreuzgr.) Donald comb. nud. =<br />

R. ritteri var. nigricans f. peterseimii<br />

pygmaea (R. E. Fries) Donald = R. pygmaea<br />

rauschii Zecher comb. nud. = R. rauschii<br />

ritteri (Wessn.) Buin. = R. ritteri<br />

steinmannii sensu Buin. = R. pygmaea<br />

c.v. ‘A. V. Frič’ Donald & Cullm. = R. euan<strong>the</strong>ma f. fricii<br />

Check list of Genus Mediolobivia Backbg.<br />

albilongiseta nomen nudum = R. aureiflora<br />

albinidulans Frič nomen nudum = ?<br />

albiscoparia Frič nomen nudum = ?<br />

albiseta Frič nomen nudum = R. aureiflora<br />

albiseta var. aureiflora Frič nomen nudum = R. aureiflora<br />

albiseta var. rubriflora Frič nomen nudum = R. aureiflora f. rubriflora<br />

albispina Frič nomen nudum = ? R. aureiflora<br />

albopectinata Rausch = R. albopectinata<br />

antarctica Hort. nomen nudum = ?<br />

apricusioides Frič nomen nudum = ?<br />

auranitida (Wessn.) Krainz = R. auranitida<br />

auranitida var. flaviflora Backbg. = R. auranitida<br />

auranitida var. gracilis (Wessn.) Backbg. = R. auranitida f. gracilis<br />

aureiflora (Backbg.) Backbg. = R. aureiflora<br />

aureiflora var. albiseta Backbg. = R. aureiflora<br />

aureiflora var. albi-longiseta Backbg. = R. aureiflora<br />

aureiflora var. albispina Backbg. nomen nudum = R. aureiflora<br />

aureiflora var. boedeckeriana (Backbg.) Backbg. = R. aureiflora f. boedeckeriana<br />

aureiflora var. brevispina Backbg. nomen nudum = R. aureiflora<br />

55<br />

Kreuzr -><br />

Kreuzgr


aureiflora var. duursmaiana (Backbg.) Backbg. = R. aureiflora f. duursmaiana<br />

aureiflora var. haertlingiana hort. nomen nudum = R. aureiflora f. rubriflora<br />

aureiflora var. leucolutea Backbg. nomen nudum = R. aureiflora<br />

aureiflora var. lilacinostoma Backbg. nomen nudum = R. aureiflora<br />

aureiflora var. longiseta Backbg. nomen nudum = R. aureiflora<br />

aureiflora var. longispina Wenzel nomen nudum = R. aureiflora<br />

aureiflora var. rubelliflora (Backbg.) Backbg. = R. aureiflora f. rubelliflora<br />

aureiflora var. robustior Backbg. nomen nudum = R. aureiflora<br />

aureiflora var. rubriflora (Backbg.) Backbg. = R. aureiflora f. rubriflora<br />

aureiflora var. rubriflora f. kesselringiana (Cullm.) Köhler) = R. aureiflora f. rubriflora<br />

aureiflora var. sarothroides (Werd.) Backbg. = R. aureiflora f. sarothroides<br />

aureiflora var. variegata Backbg. nomen nudum = R. aureiflora<br />

aureiflora subvar. leucolutea (Backbg.) Backbg. = R. aureiflora<br />

aureiflora subvar. lilacinostoma (Backbg.) Backbg. = R. aureiflora<br />

atrovirens (Backbg.) Backbg. = R. pygmaea f. atrovirens<br />

atroviridis hort. (catalogue 1974 Uhlig) U1396 = ?<br />

blossfeldii (Werd.) Buin. = R. aureiflora f. rubriflora<br />

blossfeldii var. campactiflora Wessn. = R. aureiflora f. rubriflora<br />

blossfeldii var. nigrilongiseta Wessn. = R. aureiflora f. rubriflora<br />

boedeckeriana Backbg. = R. aureiflora f. boedeckeriana<br />

brachyantha (Wessn.) Krainz = R. brachyantha<br />

breviseta Frič nomen nudum = ?<br />

brunescens (Rausch) Backbg. comb. nud. = R. brunescens<br />

bruneoradicata Ritter comb. nud. = R. bruneoradicata<br />

brunicentra Frič nomen nudum = ?<br />

brunicentra var. roseiflora Frič nomen nudum = ?<br />

bruniscoparia Frič nomen nudum = ?<br />

bruniscoperia -> bruniscoparia<br />

bruniscoparia var. chata Frič nomen nudum = ?<br />

bulbispina hort. (catalogue 1974 Uhlig) U2250 = ?<br />

cachiensis Frič nomen nudum = ?<br />

cajasensis (Ritter) Backbg. comb. nud. = R. cajasensis<br />

calenduliflora Frič nomen nudum = ?<br />

calva Frič nomen nudum = ?<br />

calva var. similis Frič nomen nudum = ?<br />

carminata Backbg. ex Krainz nomen nudum = R. aureiflora f. rubriflora<br />

citriaurata Frič nomen nudum = ?<br />

columnaris (Wessn.) Krainz. = R. einsteinii f. columnaris<br />

columnaris Frič nomen nudum = ? R. einsteinii f. columnaris<br />

conoidea (Wessn.) Krainz. = R. einsteinii f. conoidea<br />

conoidea var. columnaris (Wessn.) Backbg. = R. einsteinii f. columnaris<br />

costata (Werd.) Krainz = R. costata<br />

costata var. brachyantha (Wessn.) Donald nomen nudum = R. brachyantha<br />

costata var. eucalyptana (Backbg.) Donald comb. nud. = R. costata f. eucalyptana<br />

cupreata Frič nomen nudum = ?<br />

diersiana Rausch comb. nud. = R. diersiana<br />

digitiformis (Backbg.) Backbg. = R. pygmaea<br />

disciformis Frič nomen nudum = ?<br />

duursmaiana Backbg. = R. aureiflora f. duursmaiana<br />

56


einsteinii = R. einsteinii<br />

einsteinii var. atrovirens = ? R. einsteinii f. rubriviridis<br />

elegans Backbg. = R. aureiflora v. elegans<br />

elegans var. gracilis Backbg. nomen nudum = R. aureiflora v. elegans<br />

eos Rausch comb. nud. = R. eos<br />

eos var. roseiflora Rausch comb. nud. = R. eos<br />

erythrantha Backbg. nomen nudum = R. aureiflora f. rubriflora<br />

escayachensis Knize nomen nudum KK1686<br />

euan<strong>the</strong>ma (Backbg.) Krainz = R. euan<strong>the</strong>ma<br />

euan<strong>the</strong>ma var. fricii (Hort.) Backbg. = R. euan<strong>the</strong>ma f. fricii<br />

euan<strong>the</strong>ma var. pygmaea (Backbg.) Donald comb. nud. =<br />

R. euan<strong>the</strong>ma f. neopygmaea<br />

euan<strong>the</strong>ma var. ritteri (Wessn.) Donald comb. nud. = R. ritteri<br />

eucalyptana (Backbg.) Krainz = R. costata f. eucalyptana<br />

fricii hort. = R. euan<strong>the</strong>ma f. fricii<br />

friedrichiana Rausch comb. nud. = R. friedrichiana<br />

fuauxiana Backbg. = R. fuauxiana<br />

grandiflora Frič nomen nudum = ?<br />

haageana Borg nomen nudum = ? R. pygmaea<br />

haagei (Frič & Schelle) Backbg. = R. pygmaea<br />

haagei var. atrovirens (Backbg.) Donald comb. nud. = R. pygmaea f. atrovirens<br />

haagei var. chamaeleon Frič nomen nudum = R. pygmaea f. flavovirens<br />

haagei var. digitiformis (Backbg.) Donald comb. nud. = R. pygmaea<br />

haagei var. flavovirens (Backbg.) Backbg. = R. pygmaea f. flavovirens<br />

haagei var. flavovirens Frič nomen nudum = R. pygmaea f. flavovirens<br />

haagei var. orurensis (Backbg.) Donald comb. nud. = R. pygmaea<br />

haagei var. pectinata (Backbg.) Donald comb. nud. = R. pygmaea<br />

haagei var. salmonea Frič nomen nudum = R. pygmaea f. flavovirens<br />

haagei var. tricolor Frič nomen nudum = R. pygmaea f. flavovirens<br />

haefneriana Cullm. = R. pygmaea var. haefneriana<br />

hartlingiana Borg = ?<br />

hirsutissima Card. = R. hirsutissima<br />

iscayachensis Rausch comb. nud. = R. iscayachensis<br />

ithyacantha Card. = R. ithyacantha<br />

kesselringiana Cullm. = R. aureiflora f. rubriflora<br />

longiflora Frič nomen nudum = ?<br />

longiseta Backbg. nomen nudum = R. aureiflora<br />

mammillata Frič nomen nudum = ?<br />

mammilosa = ?<br />

melanotricha Frič nomen nudum = ?<br />

miniflora Frič nomen nudum = ?<br />

minimiflora Frič nomen nudum = ?<br />

morenoensis Rausch nomen nudum = ?<br />

mudanensis Rausch comb. nud. = R. mudanensis<br />

multiflora Frič nomen nudum = ?<br />

neopygmaea Backbg. = R. euan<strong>the</strong>ma f. neopygmaea<br />

neosteinmannii (catalogue 1974 Uhlig) U1845 = R. pygmaea v. neosteinmannii<br />

nidulans Frič = ?<br />

57<br />

fricci -><br />

fricii<br />

fricci -><br />

fricii<br />

trivolor -><br />

tricolor


nigra Frič nomen nudum = ?<br />

nigricans (Wessn.) Krainz. = R. ritteri var. nigricans<br />

nigricans var. peterseimii Frič = R. ritteri f. peterseimii<br />

nigriticentra Frič nomen nudum = ?<br />

orurensis (Backbg.) Backbg. = R. pygmaea<br />

padcayensis Rausch comb. nud. = R. pygmaea<br />

patericalyx Ritter comb. nud. = R. patericalyx<br />

patericalyx var. odontopetala Ritter comb. nud. = R. patericalyx<br />

paucipetala Frič nomen nudum = ?<br />

pectinata (Backbg.) Backbg. non Frič = R. pygmaea<br />

pectinata var. atrovirens (Backbg.) Backbg. = R. pygmaea v. atrovirens<br />

pectinata var. digitiformis (Backbg.) Backbg. = R. pygmaea<br />

pectinata var. gracilis Knize nomen nudum (KK970)<br />

pectinata var. neosteinmannii Backbg. = R. pygmaea<br />

pectinata var. paznaensis Knize nomen nudum (KK969)<br />

pectinata var. orurensis (Backbg.) Backbg. = R. pygmaea<br />

permutata Knize nomen nudum KK1046<br />

pygmaea (R. E. Fries) Krainz = R. pygmaea<br />

pygmaea var. flavovirens (Backbg.) Backbg. = R. pygmaea f. flavovirens<br />

pygmaea var. longispina nomen nudum<br />

puchella Frič nomen nudum = ?<br />

pumilla Frič nomen nudum = ?<br />

pyrimidalis Frič nomen nudum = ?<br />

rauschii Zecher comb. nud. = R. rauschii<br />

reichii Borg = Neoporteria reichii<br />

raulii Rausch comb. nud. = R. raulii<br />

ritteri (Wessn.) Krainz = R. ritteri<br />

ritteri var. pilifera (Frič) Backbg. = R. ritteri<br />

robusta Hort. nomen nudum = ?<br />

rosalbiflora Ritter comb. nud. = R. rosalbiflora<br />

roseiaurata Frič nomen nudum = ?<br />

rubelliflora Backbg. = reddish-orange flowering plant, may be a natural hybrid<br />

between <strong>the</strong> red flowered R. aureiflora f. rubriflora and <strong>the</strong> yellow flowered<br />

R. aureiflora = R. aureiflora f. rubelliflora<br />

rubriflora Backbg. = R. aureiflora f. rubriflora<br />

rubriflora var. blossfeldii (Werd.) Krainz = R. aureiflora f. rubriflora<br />

rubriflora var. nigrilongiseta (Wessn.) Krainz = R. aureiflora f. rubriflora<br />

sarothroides Werd. = R. aureiflora f. sarothroides<br />

schmiedcheniana (Kohl.) Krainz. = R. einsteinii f. schmiedcheniana<br />

schmiedcheniana var. einsteinii (Frič) Backbg. = R. einsteinii<br />

schmiedcheniana var. karreri (Frič) Backbg. = R. einsteinii<br />

schmiedcheniana var. rubriviridis Frič & Kreuzgr. ex Backbg. =<br />

R. einsteinii f. rubriviridis<br />

schmiedcheniana var. steineckei Frič & Kreuzgr. ex Backbg. =<br />

scoparia Frič nomen nudum = ?<br />

scoparia var. cylindrica Frič nomen nudum = ?<br />

semicolumnaris Frič nomen nudum = ?<br />

spiralisepala Schutz = cultivated form of R. aureiflora<br />

58<br />

steineckeri -> steineckei<br />

R. einsteinii f. steineckei<br />

pygmae -><br />

pygmaea<br />

paznensis -><br />

paznaensis<br />

pygmae -><br />

pygmaea<br />

rubiflora<br />

-> rubriflora


spiralisepala Jajo ex Brederoo nomen nudum<br />

steinmannii (Solms-Laub.) Krainz = R. steinmannii<br />

steinmannii var. complanata Rausch nomen nudum<br />

steinmannii var. rotundiseta Rausch nomen nudum<br />

steinmannii var. rubriflora Rausch nomen nudum<br />

stellata Frič nomen nudum = ?<br />

struwelpeter Frič nomen nudum = ?<br />

sulcomammillata Frič nomen nudum = ?<br />

tarvitaensis Ritter comb. nud. = R. tarvitaensis<br />

tarvitensis = R. tarvitaensis<br />

turbiniformis Frič nomen nudum = ?<br />

uniclaricentra Frič nomen nudum = ?<br />

uninigricentra Frič nomen nudum = ?<br />

wal<strong>the</strong>riana Borg = R. spegazziniana<br />

Check list of Rebutia species<br />

The following is a list of specific and varietal names which have been used in<br />

connection with <strong>the</strong> <strong>genus</strong> Rebutia. A large number of <strong>the</strong>se names are invalid,<br />

unfortunately many of <strong>the</strong>m are still in use and have <strong>the</strong>refore been included. Please<br />

realise that this is not a complete check list as we are sure <strong>the</strong>re are many o<strong>the</strong>r<br />

names and combinations which have appeared in obscure publications and which<br />

are not included. Nursery catalogues are a fruitful source of misnames and invalid<br />

combinations. As it is, <strong>the</strong> list already contains nearly 800 names and combinations.<br />

We do not pretend to know what are all <strong>the</strong> correct current names for many of<br />

<strong>the</strong> names in <strong>the</strong> following list for a variety of reasons.<br />

a) often living plants of many of <strong>the</strong> named ‘species’ are not in cultivation and in<br />

addition were sometimes inadequately described without photographs. This<br />

means that accurate assessment as to status or indeed relationship to a<br />

currently accepted name is extremely difficult and often impossible.<br />

b) names have sometimes been given to plants which after fur<strong>the</strong>r field work<br />

appear to have been described from extreme geographical forms. They have<br />

never been recollected and probably never will be again (e.g. R. albipilosa)<br />

c) many of <strong>the</strong> names are names in <strong>the</strong> literature described without full<br />

description and so referal to <strong>the</strong> correct name is often impossible.<br />

d) a number of <strong>the</strong> names have not been published in accordance with <strong>the</strong> code of<br />

Botanical Nomenclature and have <strong>the</strong>refore to be excluded.<br />

The complex and superfluous nature of <strong>the</strong> names in this <strong>genus</strong> is admirably<br />

illustrated by <strong>the</strong> example of Rebutia senilis, where <strong>the</strong>re are no less than 50 possible<br />

combinations.<br />

I Starred * names have accurate locational data and have been used in <strong>the</strong><br />

compilation of <strong>the</strong> distribution maps.<br />

II The accepted species toge<strong>the</strong>r with <strong>the</strong> names of plants which are reasonably<br />

distinct and identifiable are set in bold type.<br />

III All o<strong>the</strong>r names are set in italic type.<br />

59<br />

unigricentra -><br />

uninigricentra


Check list of Rebutia Sensu Lato<br />

c.v. ‘alabaster’ (hybrid R. minuscula v. violaciflora f.<br />

kariusiana x R. c.v. stirnaldel’s meisterstuck.)<br />

*albiareolata Ritter (FR761) (1977) Kakt. u. a. Sukk. 28 78<br />

*albiflora Ritter & Buin. (FR766a) (1963) Taxon 12 29<br />

*albipilosa Ritter (FR754) (1963) Taxon 12 29<br />

albiseta Graessn. nomen nudum = ? aureiflora<br />

*albopectinata Rausch (1972) Kakt. u. a. Sukk. 23 236<br />

allegraiana Borg nomen nudum = senilis var. elegans<br />

almeyeri W. Heinr. = ? senilis f. stuemeri<br />

archibuingiana Ritter (1978) Ashingtonia 3 14<br />

arenacea Card. = Sulco<strong>rebutia</strong> arenacea (Card.) Ritter<br />

auranitida (Wessn.) Buin. & Donald (1937) Kakt. u. a. Sukk. 9 130<br />

" var. flaviflora Backbg. = auranitida<br />

" var. gracilis (Wessn.) Donald = auranitida f. gracilis<br />

" f. gracilis (Wessn.) Buin. & Donald (1937) Kakt. u. a. Sukk. 9 130<br />

aurantiaca hort. (catalogue 1974 Uhlig) U1343 = ? U1343<br />

*aureiflora Backbg. (1932) Deutsch Kakteenfreunde 124<br />

" var. albilongiseta (Backbg.) Marshall & Bock. comb. nud. = aureiflora<br />

" var. albiseta (Backbg.) Marshall & Bock. comb. nud. = aureiflora<br />

" f. boedeckeriana (Backbg.) Buin. & Donald (1934) Blatt. f. Kakteenf. 2<br />

" var. densispina Borg nomen nudum = aureiflora<br />

* " var. elegans (Backbg.) Buin. & Donald (1934) Blatt. f. Kakteenf. 9<br />

* " f. duursmaiana (Backbg.) Buin. & Donald (1934) Blatt. f. Kakteenf. 9<br />

" f. kesselringiana (Cullm.) Köhler = aureiflora f. rubriflora<br />

" var. longiseta (Backbg.) Marshall & Bock. comb. nud. = aureiflora<br />

" f. rubelliflora (Backbg.) Buin. & Donald (1935) Kaktus ABC. 247<br />

" var. rubriflora (Backbg.) Buin. & Donald<br />

* " f. rubriflora (Backbg.) Buin. & Donald (1935) Kaktus ABC. 247<br />

" var. sarothroides (Werd.) Buin. & Donald (1936) Blüh. Sukk. f. 106<br />

" var. sarothroides (Werd.) Buin. & Donald (1936) Blüh. Sukk. f. 106<br />

" var. subspec. aureiflora (Backbg.) Donald = aureiflora<br />

" var. subspec. aureiflora var. Blossfeldii (Werd.) Donald =<br />

aureiflora f. rubriflora<br />

" var. subspec. elegans (Backbg.) Donald = aureiflora var. elegans<br />

" var. subspec. elegans var. sarothroides (Werd.) Donald =<br />

aureiflora f. sarothroides<br />

aurescens hort. (catalogue 1974 Uhlig) U1344 = ?<br />

aureicantha hort. (catalogue 1974 Uhlig) U1756 = ?<br />

*aureispina Knize nomen nudum KK843 (also KK1694)<br />

beryllioides Donald nomen nudum = wessneriana var. beryllioides<br />

" var. densiseta nomen nudum = wessneriana var. beryllioides<br />

" var. longiseta nomen nudum = wessneriana var. beryllioides<br />

*binnewaldiana W. Heinr. = ? wessneriana<br />

60<br />

rebelliflora -><br />

rubelliflora


inghamiana Borg nomen nudum = Lobivia binghamiana<br />

blossfeldii Werd. = aureiflora f. rubriflora<br />

blossfeldiana Werd. ex Backbg. nomen nudum = ? aureiflora f. rubriflora<br />

boedeckeriana (Backbg.) Marshall & Bock, nomen nudum =<br />

aureiflora f. boedeckeriana<br />

boliviensis hort. (catalogue 1974 Köhres) = ?<br />

*brachyantha (Wessn.) Buin. & Donald (1937) Kakt. u. a. Sukk. 207<br />

brachyantha Card. = Sulco<strong>rebutia</strong> breviflora (Card.) Backbg.<br />

breviflora Card. nomen Illeg. = Sulco<strong>rebutia</strong> breviflora (Card.) Backbg.<br />

*brunescens Rausch (R480) (1972) Kakt. u. a. Sukk. 23 235<br />

*brunneoradicata Ritter (FR1109) Kakt. u. a. Sukk. 23 98<br />

buiningiana Ritter nomen nudum = ? spinosissima<br />

*buiningiana Rausch (R511) (1972) Kakt. u. a. Sukk. 23 98<br />

cabratai Borg nomen nudum = Lobivia densispina<br />

caineana Card. = Sulco<strong>rebutia</strong> caineana (Card.) Donald<br />

cajasensis Ritter (FR1141) (1977) Succulenta 56 64<br />

calliantha Bew. nomen nudum<br />

calliantha Wessn. = wessneriana<br />

" var. breviseta Backbg. = wessneriana<br />

" var. beryllioides Buin. & Donald = wessneriana var. beryllioides<br />

" var. beryllioides f. breviseta (Backbg.) Buin. & Donald = wessneriana<br />

" f. hyalacantha Buin. & Donald = wessneriana<br />

" var. densiseta Bew. nomen nudum = krainziana<br />

" var. kariusiana Buin. & Donald = minuscula var. violaciflora f. kariusiana<br />

" var. krainziana (Kess.) Buin. & Donald = krainziana<br />

*camarguensis Rausch (R311) (1976) Succulenta 55 42<br />

*canacruzensis Rausch (1976) Kakt. u. a. Sukk. 27 49 = a form of pygmaea<br />

canaletas Knize nomen nudum KK1565<br />

candiae Card. = Sulco<strong>rebutia</strong> candiae (Card.) Buin. & Donald<br />

canigueralii Card. = Sulco<strong>rebutia</strong> canigueralii (Card.) Buin. & Donald<br />

caracarensis Card. = Sulco<strong>rebutia</strong> caracarensis (Card.) Donald.<br />

*carmeniana Rausch (1978) Kakt. u. a. Sukk. 29 105<br />

carminea Buin. = minuscula var. violaciflora<br />

christinae Rausch (R492a) (1975) Kakt. u. a. Sukk. 26 145<br />

chrysacantha Backbg. (1935) Kaktus ABC, 416<br />

" var. densispina nomen nudum<br />

" var. elegans (Backbg.) Backbg. = senilis var. elegans<br />

" var. iseliniana (Krainz) Donald comb. nud. =<br />

senilis var. iseliniana<br />

" var. kesselringiana (Bew.) Donald comb. nud. =<br />

senilis var. kesselringiana<br />

*cincinnata Rausch (R300) (1976) Kakt. u. a. Sukk. 27 4<br />

*cintiensis Ritter(FR938) (1978) Ashingtonia 2 206<br />

citricarpa Frič & Backbg. = xanthocarpa f. citricarpa<br />

" var. salmonea Frič nomen nudum = xanthocarpa f. salmonea<br />

*colorea Ritter (FR1106?) (1977) Kakt. u. a. Sukk. 28 78<br />

conoidea Wessn. = einsteinii f. conoidea<br />

" var. columnaris Wessn. = einsteinii f. columnaris<br />

corroana Card. = Sulco<strong>rebutia</strong> corroana (Card.) Don. & Bred.<br />

61<br />

spinossima -><br />

spinosissima<br />

3111 -> 311


*costata Werd. (1934) Notizbl. Bot. Gait. u. Mus. Berlin 12 25<br />

* " f. eucalyptana (Backbg.) Buin. & Donald (1935) Kaktus ABC, 414<br />

" f. pilifera Buin. & Donald (1965) Cact. Succ. J. Gt. Brit. 27 40<br />

dasyphrissa Werd. = xanthocarpa f. dasyphrissa<br />

*deminuta (Web.) Br. & R. (1904) Bull. Mus. Hist. Nat. Paris, 386<br />

* " f. pseudominuscula (Speg.) Buin. & Donald (1905) Anal Mus. Nac.<br />

Buenos Aires Ser. III, 488<br />

*densipectinata Ritter (FR758) nomen nudum = ? form of R. albopectinata<br />

*diersiana Rausch (R631) (1975) Kakt. u. a. Sukk. 26 25<br />

" var. atrovirens Rausch (R633) (1975) Kakt. u. a. Sukk. 26 26<br />

" var. minor Rausch nomen nudum<br />

donaldiana Lau & Rowley (Lau 348) (1974) Ashingtonia 1 76<br />

durispina Frič nomen nudum = ?<br />

" var. violrosea Frič nomen nudum = ?<br />

duursmaiana (Backbg.) Marshall & Bock. comb. nud. = aureiflora f. duursmaiana<br />

*einsteinii Frič ex Backbg. (1956) Descr. Cact. Nov. 30<br />

" var. atrovirens Frič = ? einsteinii f. rubriviridis<br />

" var. columnaris (Wessn.) Buin. & Donald = einsteinii f. columnaris<br />

" f. karreri (Backbg.) Buin. & Donald = einsteinii<br />

" var. columnaris f. conoidea (Wessn.) Buin. & Donald =<br />

einsteinii f. conoidea<br />

* " f. columnaris (Wessn.) Buin. & Donald (1940) Beitr. Sukk. u. pflege 4<br />

" var. conoidea (Wessn.) Buin. & Donald = einsteinii f. conoidea<br />

" f. conoidea (Wessn.) Buin. & Donald (1940) Beitr. Sukk. u. pflege 3<br />

* " var. gonjianii (Kies.) Donald (1973) Bol. Soc. Argentina Bot. 15 132<br />

" var. rubriviridis (Frič & Kreuzgr. ex Backbg.) Buin. & Donald =<br />

einsteinii f. rubriviridis<br />

" f. rubriviridis (Frič & Kreuzgr. ex Backbg.) Donald (1956)<br />

Descr. Cact. Nov. 30<br />

* " f. schmiedcheniana (Kohl.) Buin. & Donald (1939) Beitr. Sukk. u. pflege 37<br />

" var. steineckei (Frič & Kreuzgr. ex. Backbg.) Buin. & Donald =<br />

einsteinii f. steineckei<br />

" f. steineckei (Frič & Kreuzgr. ex. Backbg.) Buin. & Donald (1956)<br />

Descr. Cact. Nov. 30<br />

elegans Backbg. = aureiflora var. elegans<br />

" var. gracilis Backbg. = aureiflora<br />

*eos Rausch (R333) (1972) Succulenta 51 2<br />

" var. roseiflora Rausch nomen nudum (R333a)<br />

espinosae Knize nomen nudum KK1518 = R. narvaesensis<br />

euan<strong>the</strong>ma (Backbg.) Buin. & Donald (1956) Backbg. Descr. Cact. Nov. 30<br />

" f. fricii (Backbg.) Buin. & Donald (1956) Backbg. Descr. Cact. Nov. 30<br />

" f. neopygmaea (Backbg.) Buin. & Donald (1956) Backbg.<br />

Descr. Cact. Nov. 30<br />

" f. oculata (Werd.) Buin. & Donald (1935) Blüh. Kakt. u. a. Sukk. 99<br />

eucalyptana (Backbg.) Donald = costata f. eucalyptana (Backbg.) Buin. & Donald<br />

*fabrisii Rausch (R688) (1977) Kakt. und Sukk. 28 52<br />

" var. aureiflora Rausch (R687) (1977) Kakt. u. a. Sukk. 28 53<br />

famatimensis (Speg.) Speg. = Lobivia densispina<br />

62<br />

Hurt -> Hist<br />

steineckeri -> steineckei<br />

fametimensis -> famatimensis<br />

WRONG recombination!!!


*fiebigiana W. Heinr. = ? wessneriana<br />

*fiebrigii (Guerke) Br. & R. (1905) Notizbl. Bot. Gart. u. Mus. Berlin 4 183<br />

" var. densiseta (Cullm.) Oeser = fiebrigii f. densiseta<br />

" var. densiseta Cullm.<br />

*flavistyla Ritter (FR756) (1978) Ashingtonia 3 12<br />

friciana Donald & Cullm. nomen nudum = euan<strong>the</strong>ma f. fricii<br />

fricii (Hort.) Backbg. nomen nudum = euan<strong>the</strong>ma f. fricii<br />

*friedrichiana Rausch (R630) (R642) (1976) Succulenta 55 101<br />

*froehlichiana Rausch (R649) (1975) Succulenta 54 225<br />

fuauxiana (Backbg.) Backbg. = R. pygmaea f. fuauxiana<br />

*fulviseta Rausch (R583) (R319) (1970) Kakt. u. a. Sukk. 21 29<br />

" var. albispina Rausch nomen nudum (R495)<br />

*fusca Ritter (FR940) (1977) Kakt. u. a. Sukk. 28 78<br />

fusca Hort. nomen nudum<br />

gibbulosa Knize nomen nudum KK1563<br />

glanduliflora Card. nom. prov. = Sulco<strong>rebutia</strong> spec.?<br />

glaucescens Backbg. nomen nudum = ?<br />

glomeriseta Card. = Sulco<strong>rebutia</strong> glomeriseta (Card.) Ritter<br />

glomeriseta Card. = Sulco<strong>rebutia</strong> glomeriseta (Card.) Buin. & Donald<br />

gonjianii Kiesl. = einsteinii var. gonjianii<br />

*graciliflora Backbg. (1963) Descr. Cact. Nov. 3 13<br />

graciliflora var. borealis Ritter nomen nudum FR34la<br />

gracilis Borg nomen nudum Hybrid - intermediate form between R. senilis and<br />

R. xanthocarpa<br />

*gracilispina Ritter (FR1118) (1977) Kakt. u. a. Sukk. 28 76<br />

graessneri Frič nomen nudum = minuscula var. violaciflora<br />

grandiflora Backbg. non Frič = minuscula var. grandiflora<br />

x grandilacea Johnson hybrid (grandiflora x violaciflora)<br />

haageana Frič ex Backbg. = R. pygmaea<br />

haagei Frič & Schelle (1930) Kaktusar 88 = R. pygmaea<br />

haefneriana Cullm. = pygmaea f. haefneriana<br />

hahniana Werd. = ? ritteri var. nigricans f. hahniana<br />

hahnii -> hahnii Werd. = ?<br />

hahnii hahnii Frič. nomen nudum = ?<br />

haseltonii Card. = Sulco<strong>rebutia</strong> haseltonii (Card.) Donald<br />

heinrichiana hort. nomen nudum (catalogue 1977 Uhlig) = ?<br />

*heliosa Rausch (R314) (1970) Kakt. und Sukk. 21 30<br />

" var. cajasensis Donald (Lau401) (1979) Ashingtonia 5 144<br />

" var. condorensis Donald (Lau405) (1979) Ashingtonia 5 143<br />

heliosa x albiflora An interesting new hybrid<br />

hertrichiana comb. nud. = Lobivia hertrichiana<br />

*hirsutissima Card. (1971) Cact. Succ. J. Am. 43 244<br />

*hoffmannii Diers & Rausch (R521a) (1977) Kakt. u. a. Sukk. 28 105<br />

*huasiensis Rausch (R313) (1977) Kakt. u. a. Sukk. 28 25<br />

hyalacantha (Backbg.) Backbg. = wessneriana<br />

inflexiseta Card. = Sulco<strong>rebutia</strong> inflexiseta (Card.) Donald<br />

*iridescens Ritter (FR1434) (1977) Kakt. u. a. Sukk. 28 76<br />

*iscayachensis Rausch (R335b) (1977) Succulenta 56 1<br />

*ithyacantha (Card.) Diers. (1972) Kakt. u. a. Sukk. 23 12<br />

63<br />

densiseta -><br />

densiseta<br />

freidrichiana -><br />

friedrichiana<br />

hertricheana -><br />

hertrichiana<br />

Sukkulenta -><br />

Succulenta


*jujuyana Rausch (R220) (1973) Kakt. u. a. Sukk. 24 147<br />

kariusiana Wessn. Locality unknown - selected from 200 field collected plants of<br />

R. senilis and R. marsoneri imported by Uebelmann. (1963) Kakt. u. a. Sukk. 14 149 =<br />

minuscula v. violaciflora f. kariusiana<br />

*kieslingii Rausch (R694) (1977) Kakt. u. a. Sukk. 28 177<br />

knuthiana Backbg. (1935) Kaktus ABC 416 = minuscula var. violaciflora f. knuthiana<br />

*krainziana Kesselring (1948) Sukkulentenkunde II Jb. Schweiz Kakt. Ges. 23<br />

" var. breviseta (Backbg.) Donald (1935) = ? Hybrid<br />

krainziana x marsoneri Kaktus ABC 416<br />

" f. breviseta (Backbg.) Krainz & Haarm.<br />

" var. breviseta f. beryllioides (Buin. & Donald) Krainz & Haarm. =<br />

wessneriana var. beryllioides<br />

" var. hyalacantha (Backbg.) Buchheim = wessneriana<br />

" var. longiseta nomen nudum = wessneriana<br />

" var. wessneriana (Bew.) Krainz & Haarm. = wessneriana<br />

" var. wessneriana f. calliantha (Bew.) Krainz & Haarm. = wessneriana<br />

kruegeri (Card.) Backbg. = Sulco<strong>rebutia</strong> kruegeri (Card.) Ritter<br />

*kupperiana Bod (1932) Monats. D. K. Gesell. 276<br />

*kupperiana var. spiniflora Ritter (FR762b) (1977) Kakt. u. a. Sukk. 28 78<br />

*lanosiflora Ritter (FR1116) (1977) Kakt. u. a. Sukk. 28 77<br />

lasseniana hort. nomen nudum = ?<br />

*lauii Donald (Lau416)<br />

lateritia Knize nomen nudum KK1519<br />

*leucan<strong>the</strong>ma Rausch (R305) (1975) Kakt. u. a. Sukk. 26 125<br />

" var. coccinifera Ritter (1977) Succulenta 56 63<br />

longiflora Frič nomen nudum = ?<br />

longispina Frič nomen nudum = ?<br />

lucida Frič nomen nudum = ?<br />

*mamillosa Rausch R302 (1972) Succulenta 51 69<br />

" var. australis Rausch (1977) Kakt. u. a. Sukk. 28 77 = ? var. or form of<br />

R. fulviseta<br />

" var. orientalis Ritter (1977) Kakt. u. a. Sukk. 28 77 = ? var. or form of<br />

R. spegazziniana<br />

maresii Borg nomen nudum = Lobivia densispina<br />

*margarethae Rausch (R521) (1972) Kakt. und Sukk. 23 4<br />

*marsoneri Werd. (1937) Kakteenkunde 2<br />

" var. albescens hort. nomen nudum<br />

" var. breviseta hort. nomen nudum<br />

" var. brevispina Donald nomen nudum<br />

" var. grandiflora Donald nomen nudum<br />

" var. sieperdaiana (Buin.) Donald = marsoneri f. sieperdaiana<br />

* " f. sieperdaiana (Buin.) Buin. & Donald (1941) Succulenta 23 15<br />

" var. spathulata hort. nomen nudum<br />

" f. spathulata Donald nomen nudum<br />

" c.v. spatula Donald<br />

" var. vatteri Donald nomen nudum<br />

maximiflora Frič nomen nudum = ?<br />

c.v. meisterstuck Hort. White flowered hybrid produced by Stirnadel.<br />

Rebutia spec x Echinopsis spec (polyancistra?)<br />

64<br />

krugeri -><br />

kruegeri<br />

7626 -> 762b<br />

laterita -><br />

lateritia<br />

Sukkulenta -><br />

Succulenta<br />

Stirnaldel -><br />

Stirnadel


melachlora Ritter & Buin. nomen nudum = leucan<strong>the</strong>ma v. cocciniflora<br />

melanea Borg nomen nudum = Lobivia densispina ?<br />

*minuscula K. Schum. (1895) Monatsschte Kakteenkde. 5 102<br />

" f. bruneo-aurantiaca Simon nom. prov. = minuscula<br />

" var. citricarpa (Frič) Simon = xanthocarpa f. citricarpa<br />

" var. coerulescens (Backbg.) Simon = xanthocarpa f. coerulescens<br />

" var. grandiflora Krainz nomen nudum<br />

" var. grandiflora Marsh. nom. prov.<br />

* " var. grandiflora (Backbg.) Buin. & Donald<br />

" var. subspec. grandiflora (Backbg.) Donald = minuscula var. grandiflora<br />

" f. knuthiana (Backbg.) Buin. & Donald = minuscula var.<br />

violaciflora f. knuthiana<br />

" var. multiflora Frič nomen nudum = minuscula<br />

" f. kariusiana (Wessn.) Buin. & Donald = minuscula var.<br />

violaciflora f. kariusiana<br />

" var. salmonea (Frič) Simon = xanthocarpa f. salmonea<br />

" var. senilis (Backbg.) Simon = senilis<br />

" f. violaciflora (Backbg.) Buin. & Donald = minuscula var. violaciflora<br />

* " var. violaciflora (Backbg.) Buin. & Donald (1935) Blatt. f. Kakt. 8<br />

" var. violaciflora f. kariusiana (Wessn.) Donald (1963) u. a. Sukk. 14 149<br />

" var. violaciflora f. rosiflora nom. prov.<br />

" var. violaciflora f. knuthiana (Backbg.) Buin. & Donald (1935) Kaktus<br />

ABC 277 & 416<br />

" var. subspec. violaciflora (Backbg.) Donald = minuscula var. violaciflora<br />

" var. subspec. violaciflora f. kariusiana (Wessn.) Donald = minuscula var.<br />

violaciflora f. kariusiana<br />

*minutissima Ritter (FR1124) (1977) Kakt. u. a. Sukk. 28 78<br />

*mixta Ritter (FR1429) (1977) Kakt. u. a. Sukk. 28 76<br />

*mixticolor Ritter (FR1108) (1977) Kakt. u. a. Sukk. 27 169<br />

*mudanensis Rausch (R689) (1976) Kakt. u. a. Sukk. 27 169<br />

multicolor Ritter nomen nudum (FR1108)<br />

multiplex (Pfeiff.) v. Roeder (1938) Kakteenkunde 124 = ? Echinopsis multiplex<br />

multicostata hort. nomen nudum = ?<br />

muricata Frič nomen nudum = minuscula var. violaciflora<br />

*muscula Ritter & Thiele (FR753) (1963) Taxon 12 29<br />

" var. nivosa Knize comb. nud. KK1301<br />

*napina Ritter (FR942) nomen nudum (nom. prov.?)<br />

narvaecensis (Card.) Donald (1971) Cact. & Succ. J. Am. 43 245<br />

neopygmaea Backbg. = euan<strong>the</strong>ma f. neopygmaea<br />

nicolaii Frič nomen nudum = ? (1932) Mollers Deutsch. Gartnerzeit. 47 422<br />

nidulans Borg nomen nudum = ?<br />

nigra Frič nomen nudum = ?<br />

nigricans (Wessn.) Backbg. = ritteri var. nigricans<br />

*nitida Ritter (FR769) (1978) Ashingtonia 3 14<br />

nivea nomen nudum = ? muscula<br />

nivosa Ritter nomen nudum = ? muscula<br />

*nogalensis Ritter (FR768) (1977) Kakt. und Sukk. 28 78<br />

oculata Werd. (1935) Blüh. Kakt. u. a. Sukk. 99 = euan<strong>the</strong>ma f. oculata<br />

*odontopetala Ritter (FR757a) (1977) Kakt. u. a. Sukk. 28 76<br />

65<br />

c. -> v.<br />

coerulencens<br />

-> coerulescens<br />

1 -> 29


oderata Hort. nomen nudum = wessneriana<br />

odorata?<br />

orurensis Backbg. = pygmaea<br />

*padcayensis Rausch (R322) Succulenta 56 233<br />

pallida Rausch (1977) Succulenta 56 233<br />

*patericalyx Ritter (FR757) (1977) Kakt. u. a. Sukk. 28 74<br />

" var. odontopetala Ritter nomen nudum<br />

*pauciareolata Ritter (FR1121) (1977) Kakt. u. a. Sukk. 28 77<br />

*paucicostata Ritter (FR936) (1977) Kakt. u. a. Sukk. 28 77<br />

pectinata Backbg. = pygmaea<br />

" var. atrovirens = pygmaea var. atrovirens<br />

" var. digitiformis Backbg. = pygmaea<br />

" var. neosteinmannii Backbg. = pygmaea f. neosteinmannii<br />

" var. orurensis Backbg. = pygmaea<br />

periferia hort. nomen nudum = xanthocarpa (f. salmonea?)<br />

permutata Heinr. = wessneriana<br />

" var. gokrausei Heinr. = wessneriana<br />

peruviana hort. nomen nudum = marsoneri<br />

perplexa Donald (Lau 329a) (not narvaecense (Card. Donald) (1979)<br />

Ashingtonia 3 150<br />

petersainii Borg nomen nudum = ritteri<br />

petersonii hort. nomen nudum = minuscula<br />

pilayensis Knize nomen nudum (KK863)<br />

pilifera (Buin. & Donald) Rausch nomen nudum = costata f. pilifera<br />

pilosa Knize nomen nudum (KK858)<br />

polymorpha Card. = Sulco<strong>rebutia</strong> polymorpha (Card.) Backbg.<br />

*potosina Ritter (FR1428) (1977) Kakt. u. a. Sukk. 28 77<br />

prolifera Rausch nomen nudum = fabrisii<br />

pseudodeminuta Backbg. non Frič (1933) D.K.F. 7<br />

" f. albiseta (Backbg.) Buin. & Donald (1951) Cact. Succ. J. Amer. 23 82<br />

" var. longiseta nomen nudum<br />

" f. grandiflora (Backbg.) Buin. & Donald (1951) Cact. Succ. J. Amer. 23 82<br />

" f. rubrifilamentosa Buin. & Donald (1963) Sukkulentenkunde 7 102<br />

" f. schneideriana (Backbg.) Buin. & Donald (1951)<br />

Cact. Succ. J. Amer. 23 82<br />

" f. schumanniana (Backbg.) Buin. & Donald (1933) Der Kakteenfr. 7<br />

pseudoheliosa nomen nudum = albopectinata (Lau 401)<br />

pseudokrainziana nomen nudum<br />

pseudominuscula (Speg.) Br. & R. = deminuta f. pseudominuscula<br />

*pseudopygmaea Ritter (FR1122) nomen nudum<br />

*pulchella Rausch (1972) Kakt. u. a. Sukk. 23 340<br />

pulchera Card. = Sulco<strong>rebutia</strong> pulchera (Card.) Donald<br />

*pulvinosa Ritter & Buin. (FR766) (1963) Taxon 12 29<br />

pulvispina Knize nomen nudum KK1568<br />

pseudograessneri hort. (catalogue 1974 Uhlig) U844 = ?<br />

*pygmaea (R. E. Fries) Br. & R. (1905) Nov. Act. Soc. Sci. Upsala IV 120<br />

" f. atrovirens (Backbg.) Buin. & Donald (1935) Kaktus ABC 242<br />

" f. flavovirens (Backbg.) Buin. & Donald (1935) Kaktus ABC 243<br />

" f. fuauxiana (Backbg.) Buin. & Donald (1956) Descr. Cact. Nov. 131<br />

* " f. haefneriana (Cullm.) Buin. & Donald (1955) Kakt. u. a. Sukk. 6<br />

66


" f. neosteinmannii (Backbg.) Buin. & Donald (1956) Descr. Cact. Nov. 130<br />

" var. tupizensis Rausch nomen nudum (R676)<br />

*raulii Rausch (R485) (1980) Kakt. u. a. Sukk. 31 170<br />

*rauschii Zecher (R297) (1977) Kakt. u. a. Sukk. 28 73<br />

reichii Borg nomen nudum = ? Neoporteria reichii<br />

residua Knize nomen nudum (KK1517)<br />

rigidispina Frič nomen nudum = ?<br />

*ritteri (Wessn.) Buin. & Donald (1938) Beitr. z. Skde. u. pflege 3<br />

* " var. nigricans (Wessn.) Buin. & Donald (1938) Beitr. z. Sukk. u. Pfl. 51<br />

" var. nigricans f. hahniana Buin. & Donald (1965) Cact. Succ. J. Gt. Brit. 27 40<br />

" var. nigricans f. peterseimii Buin. & Donald (1965)<br />

Cact. Succ. J. Gt. Brit. 27 40<br />

" f. peterseimii Buin. & Donald = ritteri var. nigricans f. peterseimii<br />

" f. hahniana Buin. & Donald = ritteri var. nigricans f. hahniana<br />

ritteriana hort. (catalogue 1974 Andreae) = ? ritteri<br />

*robustispina Ritter (FR618) (1977) Succulenta 56 64<br />

" var. minor Ritter nomen nudum<br />

*rosalbiflora Ritter (FR1115) (1977) Kakt. u. a. Sukk. 28 76<br />

" var. amblypetala Ritter (FR1119) (1977) Kakt. u. a. Sukk. 28 76<br />

rubelliflora (Backbg.) Marshall & Bock. comb. nud. = aureiflora f. rubelliflora<br />

*rubiginosa Ritter (FR767) (1963) Taxon 12 29<br />

rubispina Graessner nomen nudum = ?<br />

rubrispina Backbg. = ?<br />

rubriflora (Backbg.) Marshall & Bock. comb. nud. = aureiflora f. rubriflora<br />

*rutiliflora Ritter (FR1113) (1977) Kakt. u. a. Sukk. 28 76<br />

sieperdaiana Buin. = marsoneri f. sieperdaiana<br />

*salpingantha Ritter (FR937) (1977) Kakt. u. a. Sukk. 28 77<br />

salmonea Frič non Backbg. nomen nudum catalogue name = ?<br />

sanguinea Ritter (1977) Succulenta 56 65<br />

" var. minor Ritter nomen nudum<br />

sarothroides Werd. = aureiflora f. sarothroides<br />

scarlatea Frič nomen nudum = minuscula<br />

" var. brevispina Frič nomen nudum = minuscula<br />

*schatzliana Rausch (R640) Kakt. u. a. Sukk. 26 244<br />

schmiedickeana (Kohl) Marsh. & Bock. comb. nud. = einsteinii f. schmiedcheniana<br />

schmiedcheniana var. karreri Backbg. = einsteinii<br />

" var. rubriviridis Frič = einsteinii f. rubriviridis<br />

" var. steineckei Frič = einsteinii f. steineckei<br />

steineckeri -><br />

steineckei<br />

*senilis Backbg. (1932) D. Kaktfrd. 123<br />

" f. subspec. senilis Backbg. = senilis<br />

" f. subspec. senilis f. lilacinorosea (Backbg.) Buin. & Donald =<br />

senilis f. lilacinorosea<br />

" f. subspec. senilis f. stuemeri (Backbg.) Buin. & Donald =<br />

senilis f. stuemeri<br />

" f. subspec. chrysacantha (Backbg.) Donald = chrysacantha<br />

67<br />

Ffl -> Pfl<br />

sarothioides -><br />

sarothroides<br />

scarlata -><br />

scarlatea


" f. subspec. chrysacantha f. iseliniana (Krainz) Buin. & Donald =<br />

senilis var. iseliniana<br />

" f. subspec. chrysacantha f. kesselringiana (Backbg.) Buin. & Donald =<br />

senilis var. kesselringiana<br />

" f. subspec. chrysacantha f. elegans (Backbg.) Buin. & Donald =<br />

senilis var. elegans<br />

" var. aurescens Backbg. = ? This taxon probably belongs with<br />

R. wessneriana ra<strong>the</strong>r than with R. senilis. In our experience<br />

it is self sterile and fits better with Subsection II ra<strong>the</strong>r than Subsection I.<br />

" f. blossfeldii nomen nudum = senilis<br />

" var. breviseta Backbg. = krainziana<br />

" f. breviseta Backbg. = krainziana<br />

" var. calva Frič nomen nudum = ?<br />

" var. cana Borg nomen nudum = ?<br />

" var. cana Backbg. = senilis<br />

" f. cana Backbg. ex Simon. = senilis<br />

" var. chrysacantha (Backbg.) Donald comb. nud. = chrysacantha<br />

" var. chrysacantha f. elegans (Backbg.) Buin. & Donald =<br />

senilis var. elegans<br />

" var. chrysacantha f. iseliniana (Krainz) Buin. & Donald =<br />

senilis var. iseliniana<br />

" var. chrysacantha f. kesselringiana (Bew.) Buin. & Donald =<br />

senilis var. kesselringiana<br />

" var. citricarpa (Frič) Marsh = xanthocarpa f. citricarpa<br />

" var. colinsii hort. (catalogue 1974 Uhlig) U2667 = ?<br />

" var. dasyphrissa (Werd.) Marshall & Bock. comb. nud. =<br />

xanthocarpa f. dasyphrissa<br />

" var. dasyphrissa (Werd.) Donald = xanthocarpa f. dasyphrissa<br />

" var. gracilis nomen nudum = ?<br />

" var. elegans (Backbg.) Backbg. (1951) Cact. Succ. J. Amer. 23 83<br />

" var. erecta hort. nomen nudum = ?<br />

" var. hyalacantha Backbg. (Backbg.) Donald = wessneriana<br />

" var. hyalacantha f. hyalacantha (Backbg.) ex Simon = wessneriana<br />

" var. hyalacantha f. blossfeldiana Köhler nomen nudum = ?<br />

" var. iseliniana Krainz (1946) Schweizer Garten 284<br />

" var. iseliniana f. chrysacantha (Backbg.) Buin. & Donald = chrysacantha<br />

" var. iseliniana f. elegans (Backbg.) Buin. & Donald = senilis var. elegans<br />

" var. iseliniana f. kesselringiana (Bew.) Buin. & Donald =<br />

senilis var. kesselringiana<br />

" var. kesselringiana Bew. (1947) Sukkulentenkunde 19<br />

" f. lilacinorosea (Backbg.) Buin. & Donald (1935) Kaktus ABC 416<br />

" var. lilacinorosea Backbg. = senilis f. lilacinorosea<br />

" var. longiflora nomen nudum = ?<br />

" var. longispina Frič nomen nudum = ?<br />

" var. luteirosea (Backbg.) Marsh = xanthocarpa var. luteirosea<br />

" var. pallidior (Backbg.) Donald nomen comb. subnud. = ?<br />

" var. pallidior (Backbg.) Donald comb. nud. = ? xanthocarpa<br />

" var. sieperdaiana (Buin.) Backbg. = marsoneri f. sieperdaiana<br />

68<br />

pallidor -><br />

pallidior


" var. schieliana Bew. = senilis f. schieliana<br />

" f. schieliana (Bew.) Donald (1957) Kakt. u. a. Sukk. 8 105<br />

" var. semperflorens Poind. = ?<br />

" var. stuemeri Backbg. = senilis f. stuemeri<br />

" f. stuemeri (Backbg.) Buin. & Donald (1932) Deutsch Kakt. frd. 131<br />

" var. stuemeriana Backbg. = senilis f. stuemeri<br />

" var. violaciflora Backbg. nomen nudum = ?<br />

" var. xanthocarpa (Backbg.) Marsh = xanthocarpa<br />

singularis Ritter FR1423 (1978) Ashingtonia 3 12<br />

solisioides Knize nomen nudum = heliosa var. condorensis<br />

*spegazziniana Backbg. (1934) Blätter, f. Kakteen forschg. 2<br />

" atroviridis (Backbg.) Backbg. (1951) Cact. Succ. J. Am. 23 82<br />

*sphaerica Ritter (FR1140) nomen nudum<br />

*spinosissima Backbg. (1935) Blatt. f. Kakteenf. 1935-8<br />

" var. brunispina Backbg. nomen nudum = spinosissima<br />

" var. aurea Rausch nomen nudum (R318)<br />

" var. similis Frič nomen nudum = spinosissima<br />

spiralisepala Jajo nomen nudum<br />

*steinmannii (Solms-Laub.) Br. & R. (1907) Bt. Zeit. 65 153 (Section I Aylostera!)<br />

" var. cincinnata Rausch nomen nudum (R300) = ?<br />

" var. nigricans Wessn. = ritteri var. nigricans<br />

" var. pilifera (Buin. & Donald) Rausch comb. nud. = costata f. pilifera<br />

*supthutiana Rausch (R629) (1976) Kakt. u. a. Sukk. 27 121<br />

steinbachii Werd. = Sulco<strong>rebutia</strong> steinbachii (Werd.) Backbg.<br />

" rosiflora Backbg. = Sulco<strong>rebutia</strong> steinbachii v. rosiflora<br />

" var. violaciflora Backbg. = Sulco<strong>rebutia</strong> steinbachii v. violaciflora<br />

" var. violacifera Backbg. = Sulco<strong>rebutia</strong> steinbachii v. violacifera<br />

steineckei Frič nomen nudum (1932) Mollers Deutsch Gartner Zeit. 47 422 = ?<br />

*tamboensis Ritter (FR1142) (1978) Ashingtonia 2 207<br />

taratensis Card. = Sulco<strong>rebutia</strong> taratensis (Card.) Buin. & Donald<br />

*tarijensis Rausch (R87) (1975) Kakt. u. a. Sukk. 26 195<br />

*tarvitaensis Ritter (FR773) (1977) Kakt. u. a. Sukk. 28 78<br />

tiraquensis Card. = Sulco<strong>rebutia</strong> tiraquensis (Card.) Ritter<br />

" var. longiseta Card. = Sulco<strong>rebutia</strong> tiraquensis (Card.) Donald<br />

*torquata Ritter & Buin. (FR1117) (1977) Succulenta 56 63<br />

totorensis Card. = Sulco<strong>rebutia</strong> totorensis (Card.) Ritter<br />

*tropaeolipicta Ritter (FR1114) (1977) Kakt. u. a. Sukk. 28 78<br />

tuberculato-chrysacantha Card. = Sulco<strong>rebutia</strong> tuberculato-chrysacantha<br />

(Card.) Bred. & Donald<br />

*tuberosa Ritter (FR770) (1963) Taxon 12 28<br />

tunariensis Card. = Sulco<strong>rebutia</strong> tunariensis (Card.) Buin. & Donald<br />

turbinata hort. nomen nudum Winter catalogue name = ? wessneriana<br />

*vallegrandensis Card. (1970) Cact. Succ. J. Amer. 42 35<br />

violacea Frič hort. nomen nudum = xanthocarpa var. violaciflora<br />

" var. albispina Frič nomen nudum = ?<br />

" var. densispina Frič nomen nudum = ?<br />

" var. paucipetala Frič nomen nudum = ?<br />

" var. rosea Frič nomen nudum = ?<br />

69<br />

schieleana -><br />

schieliana<br />

semperlorens -><br />

semperflorens<br />

Sols -><br />

Solms<br />

200 -> 300<br />

steineckeri -><br />

steineckei<br />

tropaeolipitica -><br />

tropaeolipicta


" var. stellata Frič nomen nudum = ?<br />

*violascens Ritter (FR352) (1977) Kakt. u. a. Sukk. 28 76<br />

violaciflora Backbg. = minuscula var. violaciflora<br />

" var. brunispina hort. (catalogue 1974 Uhlig) U1175<br />

" var. carminea (Burning) Donald = minuscula var. violaciflora<br />

" var. densispina nomen nudum = minuscula var. violaciflora<br />

" var. graessneri nomen nudum = minuscula var. violaciflora<br />

" var. knuthiana (Backbg.) Donald comb. nud. =<br />

minuscula var. violaciflora f. knuthiana<br />

" var. luteispina (Backbg.) Borg hort. nomen nudum =<br />

minuscula var. violaciflora<br />

" var. rosea hort. (catalogue 1974 Uhlig) Ul173<br />

violbrunispina Frič nomen nudum = ?<br />

violsalmonea Frič nomen nudum = ?<br />

vizcarrae Card. = Sulco<strong>rebutia</strong> vizcarrae (Card.) Donald<br />

volcanensis hort. (1974 catalogue Andraea) = ?<br />

*vulpina Ritter (FR939) (1977) Succulenta 56 66<br />

wal<strong>the</strong>riana Backbg. nomen nudum = ? spegazziniana<br />

*wessneriana Bew. (1948) Sukkulentenkunde Jahrb. Schweiz. Kakt. Ges. 2 24<br />

" var. beryllioides Buin. & Donald<br />

" var. calliantha (Bew.) Krainz = wessneriana<br />

" var. densiseta nomen nudum = krainziana<br />

" var. krainziana (Kess.) Buin. & Donald = krainziana<br />

" var. permutata (Heinr.) Buin. & Donald = wessneriana<br />

" subspec. beryllioides Buin. & Donald = wessneriana var. beryllioides<br />

" subspec. wessneriana f. calliantha (Bew.) Buin. & Donald =<br />

wessneriana<br />

" subspec. wessneriana var. gokrausei (Heinr.) Donald = wessneriana gokausei -><br />

" subspec. wessneriana var. gokrausei f. permutata (Heinr.) Donald = gokrausei<br />

wessneriana<br />

c.v. 'Ruby' Donald<br />

c.v. 'Turbine' Donald<br />

winterae hort. nomen nudum = ?<br />

winteriana hort. nomen nudum = ?<br />

*xanthocarpa Backbg. (1932)D.K.F 131<br />

" var. citricarpa Frič ex Backbg. = f. citricarpa<br />

" f. citricarpa (Frič ex Backbg.) Buin. & Donald (1951) Cact. & Succ.<br />

J Amer. 23 83<br />

" var. coerulescens (Backbg.) Buin. & Donald = f. coerulescens<br />

" f. coerulescens (Backbg.) Buin. & Donald (1956) Backbg. Descr. Cact.<br />

Nov. 31<br />

" var. dasyphrissa (Werd.) Backbg. = f. dasyphrissa<br />

" f. dasyphrissa (Werd.) Buin. & Donald (1935) Blüh. Kakt. u. a. Sukk. 103<br />

" var. elegans Backbg. = senilis var. elegans<br />

" f. graciliflora (Backbg.) Donald (1963) Backbg. Descr. Cact. Nov. 3 13<br />

" var. kesselringiana comb. nud. = senilis var. kesselringiana<br />

" var. lilacinorosea nomen nudum = f. violaciflora<br />

" var. luteirosea Backbg. (1951) Cact. Succ. J. Amer. 23 83<br />

" var. magentaflora Hort. nomen nudum = f. violaciflora<br />

70<br />

Jakrb. Schwiez -><br />

Jahrb. Schweiz


" var. pallidior Backbg. = ? xanthocarpa<br />

" var. ruficeps hort. nomen nudum = ?<br />

" var. salmonea Frič ex Backbg. = f. salmonea<br />

" f. salmonea (Frič ex Backbg.) Buin. & Donald (1951) Cact. Succ. ]. Amer.<br />

23 83<br />

" f. sanguinea nomen nudum = ?<br />

" var. violaciflora (Backbg.) Backbg. = f. violaciflora<br />

" f. violaciflora (Backbg.) Buin. & Donald (1956) Backbg. Descr. Cact.<br />

Nov. 31<br />

zavaletae Card. = Sulco<strong>rebutia</strong> zavaletae (Card.) Backbg.<br />

*zecheri Rausch (R650) (1977) Succulenta 56 29<br />

Digito<strong>rebutia</strong> nazarenoensis Rausch. This new species was described by Walter<br />

Rausch in 1979. As it has not been described as a Rebutia <strong>the</strong> following combination<br />

has been made:-<br />

Rebutia nazarenoensis (Rausch). Fearn & Pearcy comb. nov.<br />

Digito<strong>rebutia</strong> nazarenoensis Rausch (1979) Succulenta 58 185.<br />

Field Data<br />

The following lists of field numbers are those which have been assigned to<br />

Rebutia plants collected by Friedrich Ritter, Walter Rausch, Alfred Lau and Karel<br />

Knize, all of whom have collected extensively in Bolivia and Argentina.<br />

Friedrich Ritter Field Numbers<br />

(All <strong>the</strong>se field numbers are prefixed by <strong>the</strong> letters FR)<br />

57 Digito<strong>rebutia</strong> haagei<br />

57a " brachyantha (pygmaea?)<br />

57b " haagei form<br />

58 Rebutia calliantha<br />

83 fiebrigii densiseta (nivosa)<br />

84 " fiebrigii<br />

84a " fiebrigii var.<br />

84b " fiebrigii var. vulpes<br />

337 " spec. (brachyantha?)<br />

339 Mediolobivia orurensis<br />

340 " eucalyptana<br />

341 " haefneriana steinmannii<br />

341a Rebutia (mamillosa v. australis) graciliflora v. borealis<br />

341b " mamillosa v. orientalis<br />

351 " haagei steinmannii<br />

352 " (violascens) haefneriana<br />

390 " fiebrigii v. densiseta<br />

402 Mediolobivia pilifera<br />

404 Aylostera spegazzinii v. atroviridis<br />

405 Mediolobivia sarothroides<br />

406 Rebutia euan<strong>the</strong>ma<br />

407 Mediolobivia aureiflora<br />

71<br />

pallidor -><br />

pallidior


618 Aylostera robustispina<br />

622 " spec.<br />

752 " pseudodeminuta<br />

753 " muscula<br />

754 " albipilosa<br />

755 " maxima<br />

756 " flavistyla<br />

757 " patericalyx<br />

757a " var. odontopetala<br />

758 " densipectinata (dark form)<br />

759 " densipectinata (pale form)<br />

760 " sanguinea spegazziniana<br />

760a " sanguinea var. minor<br />

761 " albiareolata<br />

762 Rebutia kupperiana<br />

762a " " form<br />

762b " " var. spiniflora<br />

763 " robustispina<br />

763a " " var. minor<br />

765 " spec.<br />

766 Aylostera pulvinosa<br />

766a " albiflora<br />

767 " rubiginosa<br />

768 Rebutia nogalensis<br />

769 " nitida<br />

770 " tuberosa<br />

771 " spec.<br />

772 " spec.<br />

773 " tarvitaensis<br />

935 " melachlora<br />

936 " ritteri v. paucicostata<br />

937 " salpingantha costata<br />

938 " cintiensis<br />

939 " vulpina<br />

939a " vulpina var.<br />

940 " fusca<br />

941 " senilis<br />

941a " spec.<br />

942 Rebutia napina<br />

1101 " gracilis<br />

1106 " colorea<br />

1106a " colorea var.<br />

1107 " pygmaea<br />

1108 " mixticolor<br />

1109 " brunneoradicata<br />

1110 " mamillosa var. australis - grandiflora<br />

1113 " rutiliflora<br />

1113a " spec.<br />

1114 " tropaeolipicta<br />

72<br />

sulco<strong>rebutia</strong> verticillacantha!<br />

Hymenorabutia quinesensis!


1115 " rosalbiflora<br />

1115a " spec.<br />

1116 " lanosiflora<br />

1117 " torquata<br />

1118 " gracilispina<br />

1118a " gracilispina var.<br />

1119 " rosalbiflora v. amblypetala<br />

1120 " spec.<br />

1121 " pauciareolata<br />

1122 " pseudopygmaea<br />

1123 " ritteri<br />

1124 " minutissima<br />

1130 " spec.<br />

1138 " mamillosa v. orientalis<br />

1138a " mamillosa var.<br />

1139 " poecilantha<br />

1140 " sphaerica<br />

1141 Mediolobivia cajasensis<br />

1142 Rebutia tamboensis<br />

1142a " pseudodeminuta<br />

1142b " pseudodeminuta form<br />

1423 " singularis<br />

1428 " potosina<br />

1429 " mixta<br />

1434 " iridescens<br />

Rebutias collected by Alfred Lau<br />

(All <strong>the</strong>se field numbers are prefixed by <strong>the</strong> word LAU)<br />

323 R. fiebrigii forma nova. 2700m Mina Asientos, Cochabamba.<br />

329 R. fiebrigii var. densiseta. 2600m Mina Asientos, Cochabamba.<br />

329a R. perplexa<br />

346 R. fiebrigii form, shorter and darker centrals. 2400m Pucara-La Higuera,<br />

Santa Cruz.<br />

346a R. spinosissima var. nova. 2400m Pucara-La Higuera, Santa Cruz.<br />

348 R. donaldiana. 2500m Pucara-Vallegrande, Santa Cruz.<br />

348a R. fiebrigii. 2500m Pucara-Vallegrande, Santa Cruz.<br />

350 R. ithyacantha. 2600m Pucara-Vallegrande, Santa Cruz.<br />

351 R. vallegrandensis. 2100m Pucara-Vallegrande, Santa Cruz.<br />

353 R. vallegrandensis. 2600m Pucara-Vallegrande Pass, Santa Cruz.<br />

383 R. brunescens. 3500m Tarabuco, Chuquisaca.<br />

393 R. fiebrigii form. 2900m. Calle Calle Mts., Chuquisaca.<br />

401 R. albopectinata. 2450m Condor Pass, Tarija.<br />

402 R. rubiginosa. 1600m Narvaez, Tarija.<br />

402a R. fiebrigii form. 1600m Narvaez, Tarija.<br />

404 R. buiningiana Ritt. nomen nudum (R. spinosissima?) 2500m Cara del Diablo,<br />

Tarija.<br />

405 R. albopectinata form. 2400m Cajas Pass, Tarija.<br />

73<br />

Monvillea!<br />

paecilantha -> poecilantha<br />

tamboenisis -><br />

tamboensis<br />

narvaecensis -> perplexa


407 R. rubiginosa form. 2400m. 10km. W. Narvaez, Tarija.<br />

409 R. spec. nova. 3500m Tarija to Iscayachi.<br />

410 R. tuberosa. 3500m Tarija to Iscayachi.<br />

411 R. christinae. Tarija to Iscayachi.<br />

411a R. spegazziniana var. atroviridis 3000m Tarija to Iscayachi.<br />

412 R. spegazziniana var. nova. Road to Cajas, Tarija.<br />

413 R. padcayensis. Cara del Diablo, Tarija.<br />

415 = 405 R. albopectinata. 3200m Iscayachi, Tarija.<br />

416 R. lauii. 2400m Cajas Pass, Tarija.<br />

422a R. spec. nova. 3350m Otavi-Camargo, Chuquisaca.<br />

422b R. spec. nova. 3350m Otavi-Camargo, Chuquisaca.<br />

424 R. spec. 3300m Camargo, Chuquisaca.<br />

431 R. pygmaea. 3800m El Aguilar, Jujuy.<br />

442 R. minuscula var. 1500m Sierra Medina, Tucuman Province,<br />

476 R. einsteinii var. gonjianii 3500m Quebrada del Toro, Jujuy.<br />

477 R. einsteinii var. gonjianii 3200m Quebrada del Toro, Jujuy.<br />

519 R. haagei. 3100m Humahuaca, Jujuy.<br />

520 R. spec, (similar to R. costata and R. fuauxiana). 3000m East of Volcan, Jujuy.<br />

522 R. pseudodeminuta. 2700m East of Volcan, Jujuy.<br />

533 R. spec. nova (possibly R. auranitida var. gracilis). 2700m above Tastil, Salta.<br />

541 R. spec.(nigricans?) 3800m Tafne, Jujuy.<br />

541a R. spec. (costata?) 3600m Tafne, Jujuy.<br />

541b R. eos. 3600m Tafne, Jujuy.<br />

541c R. eos var. nova? 3600m. Tafne, Jujuy.<br />

544 R. margarethae. 2100-3000m Santa Victoria, Salta.<br />

544a R. margarethae. 2400m Santa Victoria, Salta.<br />

545 R. jujuyensis. 2300m Santa Victoria, Salta.<br />

546 R. spec. nova. 2200m Santa Victoria, Salta.<br />

546a R. spec. nova. 2200m Santa Victoria, Salta.<br />

547 R. spegazziniana v. atroviridis 2300-3000m Santa Victoria, Salta.<br />

550 R. margarethae. 2800m Road from La Quiaca, Santa Victoria Pass, Salta.<br />

550a R. pseudodeminuta form? 2800m Road from La Quiaca, Santa Victoria Pass,<br />

Salta.<br />

554 R. margarethae. 3500m Road from La Quiaca, Santa Victoria Pass, Salta.<br />

557 R. wessneriana v. beryllioides. 2400m Leon, Jujuy Province.<br />

560 R. xanthocarpa v. lilacinorosea. 10km North West of Antilla, Salta.<br />

561 R. pseudodeminuta form. 10km North West of Antilla, Salta.<br />

Walter Rausch Field Numbers<br />

(All <strong>the</strong>se field numbers are prefixed by <strong>the</strong> letter R)<br />

1 Rebutia xanthocarpa<br />

11 Aylostera pseudominuscula<br />

14 Mediolobivia nigricans var. Piedra de Molinos<br />

35 " haagei<br />

67 Aylostera ithyacantha<br />

71 Mediolobivia steinmannii var.<br />

87 " tarijensis (Ritter?)<br />

74<br />

R. spegazzinniana -><br />

Sulco. tarijensis!<br />

spegazinniana -><br />

spegazziniana<br />

Aquilar -><br />

Aguilar<br />

Quebrado -><br />

Quebrada<br />

steimannii -><br />

steinmannii


88 Aylostera robustispina<br />

89 " fiebrigii v. densiseta<br />

91 Mediolobivia spec.<br />

158 " aureiflora<br />

162 " nigricans<br />

163 " einsteinii v. atrovirens (viridis?)<br />

167 Rebutia senilis<br />

208 Mediolobivia steinmannii var.<br />

208a " haefneriana<br />

210 " orurensis<br />

214 " euan<strong>the</strong>ma<br />

220 Aylostera sordida/jujuyana<br />

234 Rebutia wessneriana<br />

274 Aylostera vallegrandensis<br />

295 Rebutia rosalbiflora Potosi<br />

295a " spec. Iscayache<br />

296 Mediolobivia steinmannii v. pilifera<br />

297 " rauschii<br />

300 " steinmannii v. cincinnata<br />

301 " pectinata (yellow)<br />

302 Rebutia mammillosa<br />

304 Mediolobivia pectinata (white)<br />

305 Rebutia leucan<strong>the</strong>ma<br />

311 Aylostera umbraculiforma = camarguensis<br />

312 Mediolobivia albopectinata<br />

313 Rebutia huasiensis<br />

314 Aylostera heliosa<br />

317 " spinosissima<br />

318 " var. aurea? var. buiningiana?<br />

319 " fulviseta<br />

320 " lampromeliana = pulchella<br />

321 " fiebrigii v. castanea<br />

322 " padcayensis<br />

324 " kupperiana<br />

333 Mediolobivia eos<br />

333a " eos var. roseiflora<br />

334 " steinmannii v. rotundipetala<br />

335 " pygmaea (Villazon)<br />

335a " pygmaea (Iscayachi)<br />

335b " iscayachensis (Corda)<br />

454 " eucalyptana<br />

480 Rebutia brunescens (Tarabuco?)<br />

484 Mediolobivia mudanensis<br />

485 " rauhii (Rio Honda)<br />

486 " steinmannii v. complanata (Rio Honda)<br />

492 Aylostera graciliflora (Iscayachi)<br />

492a Rebutia christinae (Santa Victoria)<br />

493 " brunescens var.<br />

494 Mediolobivia steinmannii v. rubriflora<br />

75<br />

camplanata -><br />

complanata<br />

Iscaychi -><br />

Iscayachi<br />

rubiflora -><br />

rubriflora


495 Rebutia fulviseta v. albispina<br />

496 Aylostera spegazziniana<br />

501 Mediolobivia haagei<br />

502 " haagei<br />

503 Aylostera fiebrigii<br />

503a " fiebrigii f. brown spine<br />

506 Mediolobivia pygmaea var.<br />

507 " haagei var. (Santa Victoria-Purmamarca)<br />

507a " haagei var.<br />

508 " costata<br />

508a " pygmaea v. longispina<br />

509 " einsteinii<br />

511 Rebutia buiningiana<br />

515 Mediolobivia haefneriana<br />

518 Rebutia margarethae<br />

520 Mediolobivia ritteri<br />

521 Rebutia margarethae<br />

521a " hoffmannii<br />

522 Mediolobivia morenoensis<br />

576 " pygmaea<br />

577 " nigricans<br />

578 Rebutia einsteinii v. gonjianii<br />

579 Mediolobivia pygmaea<br />

583 Rebutia fulviseta<br />

597 " pulchella var.<br />

629 " supthutiana<br />

630 Rebutia friedrichiana (pygmaea?)<br />

631 " diersiana<br />

632 " haefneriana<br />

633 " diersiana var. atrovirens<br />

640 " schatzliana<br />

641 " schatzliana var.<br />

642 " friedrichiana<br />

643 " spinosissima<br />

644 " steinmannii var.<br />

646 Mediolobivia friedrichiana<br />

649 Rebutia froehlichiana<br />

650 Aylostera zecheri<br />

654 " pseudominuscula var.<br />

655 " pseudominuscula<br />

660 Rebutia violascens<br />

661 " senilis<br />

676 " pygmaea v. tupizensis<br />

679 Aylostera pseudodeminuta var.<br />

680 Rebutia aureiflora var. duursmaiana<br />

681 " violaciflora<br />

687 " fabrisii var. aureiflora<br />

688 " fabrisii<br />

689 Mediolobivia mudanensis<br />

76<br />

hoffmanni -><br />

hoffmannii<br />

gonjiana -><br />

gonjianii<br />

frohlichiana -><br />

froehlichiana<br />

duirsmaiana -><br />

duursmaiana


690 Rebutia pseudominuscula (carmeniana)<br />

694 " kieslingii<br />

697 " pseudodeminuta<br />

700 " oculata<br />

Karel Knize Field Numbers<br />

(All <strong>the</strong>se numbers are prefixed by <strong>the</strong> letters KK)<br />

597 Rebutia senilis f. Huahata, 2200m, Arg.<br />

598 " pseudominuscula. Catamarca, 2800m, Arg.<br />

637 Mediolobivia pectinata. Oruro, 4000m, Bolivia.<br />

797 " haefneriana. Oruro, Capulla, 3000m, Bolivia.<br />

798 " pectinata var. Oruro, Machacamarca, 4000m, Bolivia.<br />

839 Rebutia albipilosa. Tarcia, Tarija, 2800m, Bolivia.<br />

840 " violaciflora. Salta, Escoipe, 3100m, Arg.<br />

841 " narvaecensis f. Narvaez, Tarija, 2300m, Bolivia.<br />

842 " muscula. Piedra, Larga, 2300m, Bolivia.<br />

843 " aureispina. Jucanas, Tarija, 3000m, Bolivia.<br />

844 " heliosa. Jucanas, 3000m, Bolivia.<br />

845 " senilis f. Salta, Escoipe, 3200m, Arg.<br />

846 " pseudodeminuta. Piedra Larga, Tarija, 2800m, Bolivia,<br />

847 " deminuta f. Narvaez, Tarija, 3000m, Bolivia.<br />

848 " densiseta? Jucanas, Tarija, 3000m, Bolivia.<br />

849 " densipectinata. Jucanas, Tarija, 3000m, Bolivia<br />

852 " albopectinata. Sama, Tarija, 3800m, Bolivia.<br />

853 " fiebrigii v. densiseta. Iscayache, Tarija, 3500m, Bolivia.<br />

854 Mediolobivia conoidea. Iscayache, Tarija, 3000m, Bolivia<br />

855 Rebutia fiebrigii. El Molle, Tarija, 3000m, Bolivia.<br />

856 " spegazzinii. Iscayache, 3500m, Bolivia.<br />

858 " pilosa. Jucanas, Las Cajas, 2800m, Bolivia.<br />

859 " ritteri/tuberosa. Sama, Tarija, 3500m, Bolivia.<br />

860 " buiningiana. Las Cajas, Tarija, 2800m, Bolivia.<br />

861 " rubiginosa. San Antonio, Iscayache, 3200m, Bolivia.<br />

862 " tuberosa. Sama, Tarija, 3500m, Bolivia.<br />

863 " pilayensis. El Nogal, Tarija, 2500m, Bolivia.<br />

871 Mediolobivia euan<strong>the</strong>ma. Sama, Iscayache, 4300m, Bolivia.<br />

872 " auranitida? Sama, Iscayache, 4300m, Bolivia.<br />

873 " auranitida v. gracilis. Sama, Tarija, 4200m, Bolivia.<br />

968 " orurensis. Oruro, Paria, 4100m, Bolivia.<br />

969 " pectinata. v. paznaensis. Pazna, 4100m, Bolivia.<br />

970 " pectinata. v. gracilis. Pazna, 4300m, Bolivia.<br />

971 " eucalyptana. Eucalyptos, 4100m, Bolivia.<br />

972 " pectinata v. challa. Challapata, 4000m, Bolivia.<br />

973 " euan<strong>the</strong>ma v. oculata. Challapata, 4100m, Bolivia.<br />

974 " pectinata v. Poopo, 4200m, Bolivia.<br />

978 " spec. Huari, 4000m, Bolivia.<br />

1046 " permutata?<br />

77<br />

300m -><br />

3000m<br />

narvaezensis -><br />

narvaecensis<br />

pseudominuta -><br />

pseudodeminuta<br />

300m -><br />

3000m<br />

paznensis. -><br />

paznaensis.


1052 Rebutia ithyacantha. La Cueva, Tarija, 2500m, Bolivia.<br />

1096 " violaciflora v. knuthiana. Salta, El Aguilar, 2800m, Arg.<br />

1150 " narvaecensis. Piedra Larga, 2500m, Bolivia.<br />

1151 " muscula v. nivosa. Piedra Larga, 2600m, Bolivia.<br />

1152 " spec.<br />

1232 Mediolobivia ritteri. Iscayache, 3800m, Bolivia.<br />

1242 Rebutia grandiflora f. Salta, Escoipe, 3300m, Arg.<br />

1243 " senilis f. Salta, Iscayache, 3500m, Arg.<br />

1244 " xanthocarpa. Salta, El Aguilar, 3800m, Arg.<br />

1301 " muscula v. nivosa. Narvaez, Tarija, 3000m, Bolivia.<br />

1302 " narvaecensis. Narvaez, Tarija, 2800m, Bolivia.<br />

1464 Mediolobivia elegans. Salta, 3800m, Arg.<br />

1465 " pygmaea. El Aguilar, 4000m, Arg.<br />

1466 " nigricans. Salta, 3800m, Arg.<br />

1467 " pectinata v. digitiformis. Escoipe, 3800m, Arg.<br />

1468 " auranitida var. Sierra Padjoja, 4100m, Bolivia.<br />

1469 " auranitida var. Sierra Padjoja, 4100m, Bolivia.<br />

1516 Rebutia cintiensis<br />

1517 " residua<br />

1518 " espinosae<br />

1519 " lateritia<br />

1563 " gibbulosa<br />

1565 " canaletas<br />

1568 " pulvispina<br />

1686 Mediolobivia iscayachensis<br />

1694 Rebutia aureispina<br />

1695 " potosina<br />

Bibliography<br />

The following is a list of <strong>the</strong> most important publications<br />

concerning <strong>the</strong> <strong>genus</strong> Rebutia<br />

Backeberg C. (1932) Der Kakteenfreund<br />

Backeberg C. (1951) Some Results of Twenty Years of Cactus Research.<br />

Cactus Succ. J. Amer. 23 81-83<br />

Backeberg C. (1959) Die Cactaceae Vol. II Fischer Verlag, Jena.<br />

Backeberg C. (1961) Die Cactaceae Vol. VI Fischer Verlag, Jena.<br />

Backeberg C. (1966) Kakteen Lexikon Fischer Verlag, Jena.<br />

Backeberg C. (1977) Cactus Lexicon Blandford Press, Poole.<br />

Backeberg C. & Knuth F.M. (1935) Kaktus ABC Gyldendalske Boghandel,<br />

Denmark.<br />

Berger A. (1929) Kakteen, Stuttgart<br />

Bertrand A. (1951) Genre 66 Rebutia K. Sch. emend A. Bertrand. Cactus (Paris) 28 53<br />

Brederoo A.J. (1955) Rebutiae Succulenta, 36<br />

Brederoo A.J. (1956) Aylostera Succulenta, 56<br />

Brederoo A.J. (1956) Mediolobivia Succulenta, 92<br />

Brederoo A.J. (1957) Digito<strong>rebutia</strong> Succulenta, 53<br />

Brederoo A.J. (1957) Mediolobivia spiralisepala Jajo Succulenta, 92<br />

78<br />

Slata -><br />

Salta<br />

Piedro -><br />

Piedra<br />

narvaezensis. -> narvaecensis.<br />

narvaezensis. -> narvaecensis.<br />

laterita -><br />

lateritia


Britten -><br />

Britton<br />

Brederoo A.J. (1959) Cylindro<strong>rebutia</strong> Succulenta, 121<br />

Brinkman K.H. (1976) Die Gattung Sulco<strong>rebutia</strong> Kakt. u. a. Sukk.<br />

Britton N.L. & Rose J.N. (1923) The Cactaceae 1st Ed. Carnegie Institute,<br />

Washington Publication No. 248<br />

Borg J. (1937) Cacti. Macmillan, London<br />

Buining A.F.H. & Donald J.D. (1963) Die Gattung Rebutia Sukkulentenkunde Jb.<br />

Schweiz Kakt. Ges. 97<br />

Buxbaum F. (1938) Was ist Rebutia, was Aylostera? Beitr. Sukk. u. Pflege, 69.<br />

Buxbaum F. (1958) Cactus Culture based on Biology. Blandford, London<br />

Buxbaum F. (1967) Rebutia, Krainz Die Kakteen 37<br />

Donald J.D. (1950) An introduction to some new genera and subgenera proposed<br />

by Backeberg. Natn. Cactus Succ. J. 5 67<br />

Donald J.D. (1955) A Rebutia Reference and Synonymy Brit. Sect. IOS Publ. No. 2.<br />

Donald J.D. (1956) Rebutiae, Succulenta 84<br />

Donald J.D. (1957) Guide to <strong>the</strong> Rebutiae I The Nomenclature of <strong>the</strong> Rebutiae,<br />

Natn. Cactus Succ. J. 12 3<br />

Donald J.D. (1957) Guide to <strong>the</strong> Rebutiae II Key to <strong>the</strong> Rebutiae, Natn. Cactus Succ.<br />

J. 12 9<br />

Donald J.D. (1957) Notes and observations on New Cacti, I Rebutia krainziana<br />

and its varieties, Natn. Cactus Succ. J. 12 27<br />

Donald J.D. (1957) Was ist Rebutia violaciflora und was Rebutia carminea,<br />

Kakt. u. a. Sukk. 8 24<br />

Donald J.D. (1959) New plants Nr. 2 Rebutia calliantha and Rebutia wessneriana<br />

Natn. Cactus Succ. J. 14 4<br />

Donald J.D. (1973) Rebutia albopectinata Ashingtonia 1 23<br />

Donald J.D. (1974) The Rebutias From <strong>the</strong> Lau expeditions to Bolivia and Argentina,<br />

Ashingtonia 1 64<br />

Donald J.D. (1976) Studies over Rebutia, Echinopsis en Lobivia, Succulenta 55 235<br />

Donald J.D. (1974-1978) The Classification of <strong>the</strong> Rebutias, Ashingtonia 1-3<br />

Donald J.D. (1977) Echinopsis, Lobivia, Rebutia. Where to draw <strong>the</strong> line?<br />

Cact. Succ. J. Gt. Brit. 39 11<br />

Donald J.D. (1977) Fur<strong>the</strong>r observations on <strong>the</strong> Rebutia, Sulco<strong>rebutia</strong>, Weingartia<br />

problem, IOS Bulletin 3 167<br />

Donald J.D. & Brederoo A.J. (1978) The systematics of Rebutia Pt. 7. Ashingtonia 3<br />

Fearn B. (1974) An investigation into <strong>the</strong> effect of temperature on <strong>the</strong> seed<br />

germination of nine species of Cacti using <strong>the</strong>rmal gradient bars.<br />

Cactus Succ. J. Amer. 46 215<br />

Fearn B. (1977) An investigation into <strong>the</strong> effect of age on <strong>the</strong> germination potential<br />

of seeds of 600 species of Cacti. Excelsa 7 103<br />

Fritz G. (1979) Rebutia albopectinata Rausch. Kakt. u. a. Sukk. 30 12<br />

Frits -><br />

Fritz G. (1980) Nochmals zu Rebutia spec. Lau 401/405. Kakt. u. a. Sukk. 31 81<br />

Fritz<br />

Glass C. & Foster R. (1980) Whats New. Cactus Succ. J. Am. 52 164<br />

Goemas W. Maretlmans L. & Wacker Fde. (1959) Rebutia, Succulenta 38 30 & 53<br />

Rebutian -><br />

Köhler von U. (1957) Die rosaviolett-Blühenden Rebutien, Kakt. u. a. Sukk. 8 28 Rebutien<br />

Köhler von U. (1961) Medio (Pygmaea) Lobivia pygmaea (Fries) Backbg. Kakt.<br />

u. a. Sukk. 12 105<br />

Köhler von U. (1963) Zweibrücker Hybride 1962 15 113<br />

Krainz H. (1967) Rebutia, Die Kakteen.<br />

Marshall W.T. & Bock T.M. (1941) Cactaceae. Abbey Garden Press, Pasadena.<br />

79


Marshall W.T. & Woods R.S. (1945) Glossary of Succulent Plant Terms<br />

Abbey Gdn. Press Pasadena.<br />

Rausch W. (1975) Walter Rausch comments Ashingtonia 1 128<br />

Ritter F. (1977) Nieuwe cactussen uit Zuid-Amerika Succulenta 66<br />

Stace C.A. (1980) Plant Taxonomy and Biosystematics, Contemporary Biology,<br />

Edward Arnold.<br />

Schumann K. (1897) Gesemtbeschreibung der Kakteen, Neudamm.<br />

Schumann K. & Guerke M. (1898) Blühende Kakteen, Neudamm.<br />

Acknowledgements<br />

We sincerely wish to thank all <strong>the</strong> various persons who have helped in <strong>the</strong><br />

preparation of this booklet. The descriptions are based on plants in cultivation at<br />

Abbey Brook.<br />

For technical suggestions and helpful advice to Mr. N. Wall of Higham Press and<br />

Carol and Ann at Abbey Brook for typing <strong>the</strong> original manuscript. Also to <strong>the</strong><br />

Librarian, Royal Botanic Gardens, Kew for <strong>the</strong> use of facilities, to Dr. Gillian Fearn for<br />

help in proof reading and to Thelma Stuart for photographic assistance.<br />

Glossary<br />

acicular needle shaped.<br />

apomixis a biological system which can produce viable seeds by a process<br />

o<strong>the</strong>r than by direct fertilisation of <strong>the</strong> egg cell,<br />

apomictic a species which can produce seeds by <strong>the</strong> process of apomixis.<br />

areole a restricted area from which spines and/or flowers are produced,<br />

bristle hair-like spine,<br />

caespitose growing in clumps,<br />

campanulate bell-shaped,<br />

central spine <strong>the</strong> spine or spines arising from or near <strong>the</strong> centre of an areole,<br />

usually <strong>the</strong> largest spine or spines,<br />

filament <strong>the</strong> stalk upon which is borne <strong>the</strong> an<strong>the</strong>r or pollen sac.<br />

funnel-form a flower which gradually widens from <strong>the</strong> base,<br />

globose shaped like a globe,<br />

hilum <strong>the</strong> scar at <strong>the</strong> base of <strong>the</strong> seed making <strong>the</strong> point of attachment to <strong>the</strong><br />

funicle.<br />

spine a pointed more or less rigid structure which is homologous with a<br />

leaf.<br />

stamen <strong>the</strong> male floral organ which bears <strong>the</strong> pollen grains. It consists of a<br />

stalk called <strong>the</strong> filament and an an<strong>the</strong>r or pollen sac.<br />

stigma <strong>the</strong> part of <strong>the</strong> flower that is <strong>the</strong> pollen receptor,<br />

style <strong>the</strong> stalk joining <strong>the</strong> stigma to <strong>the</strong> ovary,<br />

type locality <strong>the</strong> place where a type specimen was collected.<br />

80


The Cactus Centre of England<br />

Abbey Brook is <strong>the</strong> oldest established Cactus Nursery in <strong>the</strong> U. K. and we celebrated<br />

our 25th Anniversary in 1981. We have a vast range of plants for sale (well over 2,000<br />

species) and more than 500,000 plants on view.<br />

Plants and seeds of a wide range of Rebutia species available.<br />

Comprehensive colour illustrated plant list available, stamp appreciated.<br />

We also produce a separate seed list.<br />

Nursery Visits<br />

The nursery is open to visitors 6 days a week throughout <strong>the</strong> year including Saturdays<br />

and Sundays.<br />

2p.m. - 6p.m. (Closed all day Tuesdays)<br />

We are open on all Bank Holidays except Christmas Day, Boxing Day and New Years<br />

Day.<br />

The nursery is situated just north of Matlock on <strong>the</strong> A6 trunk road.<br />

Abbey Brook Cactus Nursery<br />

Old Hackney Lane,<br />

Matlock, Derbyshire, England.<br />

☎ National 0629 55360<br />

☎ International + 44 629 55360<br />

81


R. euan<strong>the</strong>ma ▲ ▲ R. xanthocarpa f. salmonea<br />

R. wessneriana ▼ ▼ R. albipilosa<br />

R. senilis v. elegans ▼ ▼<br />

82<br />

R. marsoneri

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