European Journal of Taxonomy 382: 1–327
https://doi.org/10.5852/ejt.2017.382
ISSN 2118-9773
www.europeanjournaloftaxonomy.eu
2017 · Van Cakenberghe V. et al.
This work is licensed under a Creative Commons Attribution 3.0 License.
Monograph
urn:lsid:zoobank.org:pub:FA508A12-9BDB-4A2B-9B0C-98FDD161443C
The bats of the Congo and of Rwanda and Burundi revisited
(Mammalia: Chiroptera)
Victor VAN CAKENBERGHE 1,*, Guy-Crispin GEMBU TUNGALUNA 2,
Prescott MUSABA AKAWA 3, Ernest SEAMARK 4 & Erik VERHEYEN 5
1
University of Antwerp, Department of Biology, Functional Morphology,
Campus Drie Eiken, Universiteitsplein, 1, B-2610 Antwerpen (Wilrijk), Belgium.
1,4
AfricanBats NPC, 357 Botha Ave, Kloofsig, 0157, Republic of South Africa.
2,3
Faculté des Sciences, Université de Kisangani, Democratic Republic of the Congo.
4
Centre for Wildlife Management, University of Pretoria, Private Bag X20 Hatield,
Pretoria 0028, Republic of South Africa.
5
Royal Belgian Institute of Natural Sciences - OD Taxonomy and Phylogeny,
Vautierstraat 29, 1000 Brussels, Belgium.
5
University of Antwerp, Department of Biology, Evolutionary Ecology, Campus Drie Eiken,
Universiteitsplein 1, B-2610 Antwerpen (Wilrijk), Belgium.
*
Corresponding author: Victor.VanCakenberghe@uantwerpen.be
2
Email: gembuguycrispin@gmail.com
3
Email: prescottmusaba@yahoo.fr
4
Email: ernest.seamark@africanbats.org
5
Email: erik.verheyen@naturalsciences.be
1
urn:lsid:zoobank.org:author:9A0E1AF5-C248-4648-9D64-443112890346
urn:lsid:zoobank.org:author:2AE2F96F-A138-419B-B650-6DB44D535D14
3
urn:lsid:zoobank.org:author:4C40421C-1D84-4BD1-A444-360300ECBEEC
4
urn:lsid:zoobank.org:author:B60CA9A1-D288-468E-AEEB-D0F136E96F5B
5
urn:lsid:zoobank.org:author:86B40463-E3D9-4147-9ED3-D7302E0D64B6
2
Abstract. In 1966, Robert William Hayman, Xavier Misonne and Walter Verheyen published their listing
of the Congolese, Rwandan and Burundian bat specimens in the collections in the museums of Tervuren,
Brussels, Geneva, London and New York. In the ifty years that have passed since, some major changes
have been introduced in the taxonomy of the Chiroptera: new species have been discovered, species
have been split off, species have been moved to other genera, and additional material has been collected.
We re-evaluated the data presented by Hayman et al., and supplemented this with specimen records found
in the literature and in online catalogs. This resulted in 136 species, represented by 20 231 specimens
(compared to 113 species and 8567 specimens originally). When available, we also recorded additional
information such as locality, sex and age, collector, collection date and preservation type of the voucher
specimen. The distribution maps of the Congolese taxa are revised to represent the current taxonomy, and
are presented in perspective against the taxon’s Species Distribution Model to assess species distribution
on the African continent. Additionally, an updated key to the various taxa is presented.
Keywords. Chiroptera, Democratic Republic of the Congo, Rwanda, Burundi, review.
1
European Journal of Taxonomy 382: 1–327 (2017)
Van Cakenberghe V., Gembu Tungaluna G.-C., Musaba Akawa P., Seamark E., & Verheyen E. 2017. The bats of
the Congo and of Rwanda and Burundi revisited (Mammalia: Chiroptera). European Journal of Taxonomy 382:
1–327. https://doi.org/10.5852/ejt.2017.382
Table of Contents
Introduction …………………………………………………………………………………………… 2
Material and methods ………………………………………………………………………………… 7
Results and Discussion ……………………………………………………………………………… 11
Annotated species list …………………………………………………………………………… 11
Nominal list of bats of the Congo, Rwanda and Burundi (CRB) region ……………………… 107
Key to the bat species of the CRB region ……………………………………………………… 112
Catalogue of voucher specimens ……………………………………………………………… 126
Summary table of museum specimens ………………………………………………………… 246
Collecting localities of the bats from the CRB region ………………………………………… 260
Acknowledgements ………………………………………………………………………………… 303
References ………………………………………………………………………………………… 303
Introduction
In December 1966, issue number 154 of the “Koninklijk Museum voor Midden-Afrika - Annalen - Reeks
In-8 - Zoologische Wetenschappen / Musée Royal de l’Afrique Central - Annales - Série In-8 - Sciences
Zoologiques” appeared. In that issue, R.W. Hayman, X. Misonne and W. Verheyen reported on the bats
occurring in the Congo, Rwanda and Burundi, primarily based on material in the collections of the three
major Belgian natural history museums: the Royal Museum for Central Africa (RMCA) in Tervuren
(also known by its Dutch and French names and acronyms: Koninklijk Museum voor Midden-Afrika
(KMMA) / Musée Royal de l’Afrique Central (MRAC) and its historic name of “the Congo Museum”),
the Royal Belgian Institute of Natural Sciences (RBINS) in Brussels (also: Koninklijk Belgisch Instituut
voor Natuurwetenschappen (KBIN) / Institut Royal des Sciences Naturelles de Belgique (IRSN)), and
the Institute of the National Parks of Belgian Congo (INPBC) (Instituut der Nationale Parken van
Belgisch Congo (INPBC) / Institut des Parcs Nationaux du Congo (IPNC)), with additions from the
British Museum (Natural History), London (BMNH) and the American Museum of Natural History,
New York (AMNH) and an addenda section by V. Aellen, covering the bats in the collection of the
Muséum d’Histoire Naturelle, Geneva (MHNG).
Today, 50 years after the appearance of “The Bats of the Congo and of Rwanda and Burundi” by Hayman
et al. (1966), the current authors deemed it timely to revisit this publication and bring it up to date, as this
last half a century has brought a lot of new techniques, which yielded important changes to our views
into the systematics and the taxonomic status of bats (order Chiroptera). Ten of the species recognized
in the current study were not known to science in 1966. Some taxa were recognized as subspecies or
synonym and were raised to full species status. In a few other cases, the taxon’s content remained the
same, but the name needed to be changed due to priority rules.
As correctly remarked by Bates et al. (2013: 313), the title of Hayman et al. (1966) might have been
a bit misleading as “The Congo” did not cover the territory of “The Republic of Congo” [= Congo
(Brazzaville)], but only the area that used to be a Belgian colony or mandate area: The Democratic
Republic of the Congo (DRC), Rwanda and Burundi. Further on in the text, the acronym CRB will be
used to refer to this area.
As mentioned by Hayman et al. (1966: 1), the irst records of bats from the CRB were reported by
Johnston (1884) from the area along the Congo River from its mouth to Bolobo. The river formed (and
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VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
still forms) an important route through Congo, as is also illustrated by more recent scientiic expeditions,
e.g., the 1974–1975 Zaire River Expedition (Gallagher & Harrison 1977; 212 bats collected) and the
2009 and 2010 Congo River expeditions (in 2009 between Lisala and Kisangani and the “Boyekoli Ebale
Congo” expedition in 2010 between Bumba and Kisangani; 594 bats; Gembu Tungaluna and Musaba,
unpublished). The latter two were organized by the Congo2010 consortium, that consisted of the RMCA
(Tervuren), the RBINS (Brussels), the Botanic Garden (Meise) and the University of Kisangani, with
funding from the Belgian Federal Directorate general for Development and the Belgian Science Policy.
A second, enormously important contribution to the knowledge of bats in the CRB was provided by the
“American Museum Congo Expedition”, which started at Boma on 22 June 1909 and which took Herbert
Lang and James P. Chapin to the northeastern part of the DRC, where they made collections in Poko
(Bas-Uélé Province); Niangara, Faradje, Medje and Aba, in or near the Garamba National Park (HautUélé Province); Avakubi, near the Ituri River, Panga and Kisangani (Tshopo Province); and then back
over Kinshasa, Boma and Matadi to Banana, where the expedition inally ended on 14 September 1915
(Osborn, 1919). They collected almost 800 bats and their data formed the basis for Allen et al.‘s “The
American Museum Congo Expedition collection of Bats” (Allen et al. 1917). Allen (1917) recognized
68 forms, covering 65 different species, which were reduced to 56 by Koopman (1965).
A further increase occurred during the 1930s to late 1950s, when major collections were made for
the Institute of the National Parks of Belgian Congo in the National Parks of the eastern part of the
CRB: Albert National Park (currently the Virunga National Park; ca 480 bats), the Garamba National
Park (ca 600 bats), the Upemba National Park (ca 220 bats) and the Kagera National Park (currently
the Akagera National Park in Rwanda; ca 170 bats). The results of these expeditions were reported by
Schouteden (1934, 1935), Frechkop (1938, 1943, 1944, 1954) and Verschuren (1957). Henri Schouteden
also published a number of papers on the mammals of the entire CRB area (Schouteden 1943, 1947,
1948).
Although being an entomologist, Narcisse Leleup collected over 260 bats between 1948 and 1958 in
the DRC provinces of Katanga, Kongo Central, Maniema and Kivu, primarily for their insect parasites
(Jobling 1954; Leleup 1956).
From the mid-1950s to the early 1960s, Anciaux de Faveaux made extensive collections in the Katanga
area (currently provinces of Haut-Katanga, Haut-Lomami, Lualaba, Tanganyika; at least 1000 bats),
which were followed in the mid-1960s by a collection from Rwanda (at least 50 bats). These resulted in a
large number of publications, primarily on reproduction and parasites (Anciaux de Faveaux 1958, 1959–
1960, 1960a, 1960b, 1964, 1965, 1972, 1977, 1978a, 1978b, 1983; Benoit 1958a, 1958b; Fain 1972;
Krampitz & Anciaux de Faveaux 1960).
In the 1960s, the independence of the Congo, Rwanda and Burundi led to a diminished presence of
Belgian biologists in the area, or at least of Belgian led expeditions, although individual collectors
still sent material to museums abroad (with number of bats in brackets), e.g., Toon De Roo (443),
P. Lootens (664 - from the 1950s until the 1990s), M.J. Celis and M. Lejeune (70 combined, 300 Lejeune
only), but other nationals also became involved, e.g., P. Kunkel (for the Senckenberg Museum Frankfurt
[SMF]; 48). In the 1970s, major collections were made by Lynn and Brian Robbins (Carnegie Museum
[CM], United States National Museum [USNM]; 162), Michael Gallagher (Harrison Institute [formerly
Harrison Zoological Museum - HZM]; 214), Frieder Meier (MHNG; 119), but also the irst African
names started to appear: Tshinyenye (RMCA; 118) and Betu Nsankulu (RMCA; 128). In the 1990s there
were major collections by Julian Kerbis Peterhans (Field Museum of Natural History [FMNH]; 50) and
Bob Swanepoel (Ditsong National Museum of Natural History [formerly Transvaal Museum - TM];
515). In the irst decade of the 21st century, a large collection was made by Alexandre Hassanin and
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European Journal of Taxonomy 382: 1–327 (2017)
colleagues for the Muséum national d’Histoire naturelle (MNHN; 150 bats). Additionally, there was also
the startup of the collection at the University of Kisangani (UNIKIS) by prof. Séraphin Ifuta (presently
University of Kinshasa [UNIKIN]), which is currently still being expanded by material collected during
ield trips under the direction of the Congolese co-authors. At UNIKIS, a Centre de Surveillance de
la Biodiversité (CSB, http://centresurveillancebiodiversite.org/) was created under the directorship of
prof. Dudu Akaibe to ensure the long-term surveillance and management of how the lora and fauna of
the Congo River rainforest evolves. As head of the department of terrestrial vertebrates, Prof. Gembu
Tungaluna is responsible for the bat-related projects.
Of special interest are also the collections from Tandala and Yalosemba (1979; 215 bats - Breman
et al. 1999) and Kikwit (1995; 538 bats - Van Cakenberghe et al. 2000) where, in the aftermath of
the Ebola epidemics, expeditions were set up in search for the reservoir species for this virus, and of
which the specimens were primarily deposited in the collections of the CM, MHNG, MSB (Museum of
Southwestern Biology), RMCA and USNM. As indicated in Fig. 1, the material included in the current
study covers almost the entire DRC, although there are still a few areas that are underrepresented, e.g.,
Fig. 1. Capture localities for all bats. The symbols represent: yellow (H): records included in Hayman
et al. (1966), and brown (V): newly added records. Other coloured pins represent type specimens: red
(H): holotype, blue (P): paratype or paralectotype (for further details, see text).
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VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
the southern part of Kwango Province (former Bandundu Province), the central and northern part of
Lualaba Province (western part of the former Katanga Province), the western part of Equateur Province,
Lomami and Sankuru Provinces (former Kasai Oriental Province), and especially the central area
covered by Tshuapa, Sankuru, Kasai and Mai Ndombe Provinces, where the Parc National de la Salonga
(North and South) remains a blank area. This seems a bit strange as most of the other National Parks
were already extensively surveyed from the 1930s to the 1950s (see above), but the Salonga National
Park was only created in 1970 and is the only park situated in the Central African Rainforest, which is
impenetrable and is only accessible along the rivers running through it. Its visibility of less than 100 m
also makes it very dificult to assess any occurrence of wildlife in the park.
Jacques Verschuren collected 48 bats in Burundi in 1976 for the RBINS (Verschuren 1980). A major
collection (327 bats) from the country was made by P.L. Niort in the late 1960s–early 1970s and deposited
in the MHNG (Niort 1970). At the end of the 1970s Heinz Stephan collected 70 bats for the SMF, and at
the beginning of the 1980s, Danny Meirte (RMCA) made another important collection of 156 bats. In the
1990s Julian Kerbis Peterhans (22), L. Davenport (33) and Jay L. Udelhoven (33) made collections for
the FMNH. Besides the already mentioned Rwandese collection from the Akagera National Park made
for the INPBC and the material collected by Anciaux de Faveaux, major contributions were provided
by Alexandre Fain (late 1940s to mid 1950s; 108 bats), but primarily by a number of expeditions by the
RMCA (Danny Meirte; 171 bats) and the University of Antwerp in the early 1980s (Baeten et al. 1984;
2102 bats).
Hayman et al. (1966: 77) reported on 8567 specimens belonging to 113 species. The additional material
that has been traced for this study augmented these numbers to 20 231 bats belonging to 136 species.
For the current study not all of the specimens were (re-)examined by the authors. This was
only done for the material in the collection of UNIKIS, most of the specimens from Rwanda
(Baeten et al. 1984), Kikwit (Van Cakenberghe et al. 2000) and for speciic groups: Epomophorus
(Claessen & De Vree 1990, 1991), Nycteris (Van Cakenberghe & De Vree 1985, 1993, 1999) and
Pipistrellus / Eptesicus / Neoromicia / Hypsugo (Van Cakenberghe, unpublished).
The current study follows the structure used in Hayman et al. (1966), with the exception of two extra
sections (Material and methods and Annotated species list) and the removal of the index section. The
order in which the various taxa are presented follows the currently recognized view on chiropteran
taxonomy. For an overview, we refer to the “Nominal list of bats of the Congo, Rwanda and Burundi
(CRB) region” section. For each of the species, we provide the current name, and in the synonyms list
we only include the original name and (preixed by an asterisk) the names as used by Hayman et al.
(1966). For a complete list of synonyms, we refer to ACR (2016).
The systematics follows Simmons (2005) and “Mammals of Africa volume IV” (Happold & Happold
2013), unless explicitly mentioned in the annonated catalog section. However, due to the dificulties in
assessing the subspeciic boundaries of the various taxa, we did not distinguish any subspecies.
Since the days of Hayman et al. (1966), taxonomic techniques have evolved, leading to the discovery and/
or recognition of new species. Where morphological comparisions and univariate statistical tests were
the state-of-the-art techniques, the rise of computers in the 1970s-1980s led to multivariate statistical
analyses. Unfortunately, these were not always capable of distinguishing very similar or cryptic species
(Baker & Bradley 2006), and, therefore, other methods, looking at more fundamental building blocks of
life (biochemical techniques), were investigated such as karyotypes, electrophoresis and in recent years
DNA sequencing. Important new techniques, such as molecular phylogenetics and phylogeographic
analyses, are also contributing to our current understanding of species and their distribution patterns.
However, these biochemical techniques cannot stand on their own (Ruedas et al. 2000), and the collection
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European Journal of Taxonomy 382: 1–327 (2017)
of tissues should be backed by continued collecting of voucher specimens to substantiate their identity
(Patterson 2002). Zachos et al. (2013: 1) discourage species splitting based on gene trees inferred from
mitochondrial DNA only, as uncritical acceptance of such new species might create an unnecessary
burden on the conservation of biodiversity.
As already pointed out by Hayman et al. (1966: 3), the central geographic position of the CRB region in
Africa and its immense size makes it a crossroad for bat species from all over the continent. Konaté &
Linsenmaier (2010: 18) marked the eastern parts of the DRC provinces of Ituri, Nord-Kivu, Sud-Kivu
and Tanganyika and the western parts of Rwanda and Burundi as one of the world’s biodiversity hotspots:
the Eastern Afromontane hotspot (see also Ceballos & Ehrlich 2006). Some of the species occurring in
the CRB area have an almost pan-African distribution (e.g., Nycteris thebaica, Pipistrellus rueppellii)
whereas others were found in a very restricted area – in a number of cases only at the type locality,
e.g., Rhinolophus hilli, R. kahuzi, R. ruwenzorii, R. willardi, Chaerephon gallagheri, Mops niangarae.
With the exception of the two molossid species, all of these are only found in the eastern DRC endemic
hotspot as deined by Herkt et al. (2015: igs 3–4). In between these extreme distribution ranges, there
are species which are common in sub-Saharan Africa (e.g., Eidolon helvum, Hipposideros caffer,
Taphozous mauritanus, Nycteris hispida, Chaerephon pumilus, Mops condylurus, Scotophilus dinganii,
Neoromicia nana), others are found primarily in western Africa (e.g., Hipposideros abae, Rhinolophus
alcyone, Myonycteris daubentonii), eastern Africa (e.g., Epomophorus minimus, Rhinolophus eloquens,
Tadarida fulminans), Southern Africa (e.g., Plerotes anchietae, Rhinolophus swinnyi, Cloeotis percivali,
Hypsugo anchietae, Laephotis botswanae) or Northern Africa (e.g., Taphozous nudiventris). Some of
the species only occur in the African forest areas (e.g., Epomops franqueti, Hypsignathus monstrosus,
Megaloglossus woermanni, Saccolaimus peli, Nycteris arge, Glauconycteris superba), and some even
only in the Central African forest (e.g., Casinycteris argynnis). Hayman et al. (1966: 3) particularly
pointed to the enormous extent of the tropical forest belt contributing to the importance of the CRB area.
However, commercial logging and expansion of farming has made the Congo rainforest one of the
most endangered ecosystems in the world (http://interesting-africa-facts.com/Africa-Landforms/CongoRainforest-Facts.html).
The apparent rarity of some of the species reported in this paper might be the result of small geographic
ranges, limited habitat breadth, low local population density, or simply a sampling artefact linked
to insuficient sampling effort, inadequate methodology (mist nets, canopy nets, harp traps, …), or
differential species detectability (Meyer et al. 2014: 114). In most cases, the exact reason for this is
currently unknown, and might possibly only be resolved once more specimens are collected, which
means that extensive ield work is still a necessity, even in times where people think that everything can
be solved from behind a computer screen.
Did we cover all of the known specimens/species? Probably not. Are all of the included specimens
correctly identiied? Unlikely. What we wanted to achieve was to provide an inventory of the
CRB specimens that are available in collections worldwide that can be used for further research, and we
certainly encourage other/local researchers to investigate these collections and verify the data provided
here.
The bottom line, however, is: you cannot deliver good, reliable, reproducible science if you cannot
conirm the identity of the animals you are (or have been) working on. This is especially important for
bat-related studies, as these animals have recently been recognized as potential carriers of pathogenic
and infectious agents that can pose threats to humans: e.g., rabies (Adedeji et al. 2010), ebola (Leirs et al.
1999; Leroy et al. 2005, 2009; Grard et al. 2011; Hayman et al. 2012; Leendertz 2016), see also Wibbelt
et al. (2010), Newman et al. (2011), Schneeberger & Voigt (2015), Ranjan et al. (2016). Moreover, Han
et al. (2016: 6) illustrate that the northeastern part of the DRC forms an African hotspot related to the
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VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
number of ilovirus-positive bat species. To substantiate these studies, we stress the importance of the
collection of voucher specimens and their safekeeping for future generations of scientists (Rocha et al.
2014; Tsang et al. 2015). However, as mentioned by Paul Racey (in Wibbelt et al. 2010: 440): “… every
effort must be taken to avoid a reductionist approach to newly described infectious diseases and that
all factors associated with the emergence of infectious diseases must be considered so that bats do not
simply become targets for persecution and eradication.” To which we could add that collecting bats for
the sole purpose of increasing collection sizes isn’t a good practice either, and hopefully this paper might
contribute to a more eficient use of already established collections.
Material and methods
The base for the current publication was formed by the catalog data in Hayman et al. (1966), with
additional information from the catalog records from the RMCA and RBINS. We also searched online
catalogs (e.g., http://vertnet.org/, http://arctos.database.museum/SpecimenSearch.cfm and http://www.
gbif.org/) and consulted the online catalogs of additional museums (for details see the “Summary table
of museum specimens” section below). Further information was obtained from Ammerman et al. (2013),
Anciaux de Faveaux (1972), Andersen (1912), Ansell (1960, 1974), Benda & Vallo (2012), Bergmans
(1988, 1989, 1991, 1994, 1997), Cotterill (2001b), Csorba et al. (2014), Dollman (1909, 1914),
Eger & Schlitter (2001), Fahr & Ebigbo (2003), Fain (1972), Fain & Aellen (1994), Freeman (1981),
Gallagher & Harrison (1977), Gregorin and Cirranello (2015), Gunnell et al. (2015a), Harrison (1957,
1959), Hassanin et al. (2015), Hayman (1937, 1946, 1960), Heller et al. (1994), Hill (1974a, 1977),
Hill & Harrison (1987), Hoofer & Van Den Bussche (2003), Hoofer et al. (2006), Kerbis Peterhans et al.
(2013), Kityo et al. (2009), Kock (1969, 1978, 1981a, 1981b, 1987), Kock & Howell (1988), Kock et al.
(2002), Koubínová et al. (2013), Lack et al. (2010), Laumanns et al. (2004), Monadjem et al. (2010b),
Noack (1889), Omatsu et al. (2008), Ossa Gómez (2012), Rahm (1966), Ralph et al. (2015), Reeder
et al. (2013), Robbins et al. (1985), Roehrs (2009), Roehrs et al. (2010), Soisook et al. (2015), Stanley
et al. (1996), Thorn et al. (2007), Uchikawa (1985), Uchikawa & Kock (1989), Van Cakenberghe &
De Vree (1985, 1993), Volleth & Heller (1994) and Volleth et al. (2001).
The supplementary material included with this paper consists of a zip ile which contains htm iles and
a separate directory, which contains pin symbols. The htm iles need to be placed in a directory called
“c:\CongoBatMaps” and the pins (*.png) in “c:\CongoBatMaps\pins”. The zip ile also contains some
additional htm iles for the specimens which were not identiied to the species level.
The pins on these maps (as on Fig. 1) are coded as follows: yellow (H): records included in Hayman
et al. (1966), and brown (V): newly added records. Other coloured pins represent type specimens: red
(H): holotype, red (L): lectotype, blue (P): paratype or paralectotype, and green (T): topotype. To be able
to separate multiple specimens from (almost) the same locality, the irst specimen is plotted at the correct
coordinates and subsequent specimens are moved slightly to the north.
For each of the pins, the following information is shown (when available): museum acronym and
number, age and sex, sort of type and the original name, current name, country, locality, coordinates,
altitude, collector and collection date. In the lower left hand corner of the map, there is a bat icon which
contains the name of the species and the version number.
In the “Annotated Species List”, the species name header is followed by a reference (Fig. xx) to the
distribution maps. For each species, two maps are included: one (“A, C or E”) shows the distribution
within the CRB area. Each symbol on the map represents a locality where the species has been collected.
The blue squares represent localities where the species was already reported from by Hayman et al.
(1966), the red dots are new localities.
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European Journal of Taxonomy 382: 1–327 (2017)
To place the species occurring from the CRB in context, a Species Distribution Modelled (SDM) map
(“B, D or F”) was created using the point data from ACR (2016). MaxEnt was used to estimate the PanAfrican distribution of each species (Phillips et al. 2006), based on 19 climate variables from the World
Clim (Hijmans et al. 2005). Each species output was clipped using a buffered minimum convex polygon
of 500 km (Kremen et al. 2008) resulting in the solid red areas on the map. The inal step was to draw
subjectively a suggested range extent of the species combining the point localities, SDM clipped output
and the Sayre et al. (2013) bioclimatic zones, resulting in the hatched area.
When examining the SDM output maps, we are aware that most make sense, while others might look
doubtful. One of the major reasons for this is that one needs to have at least 10 separate localities to
generate a reliable map, which for a number of species simply is not possible (e.g., Rhinolophus kahuzi:
only known from the type locality). In a few cases, we also found that the various models we tested did
not include areas where the species was actually collected by one of the authors. We, therefore, would
like to highlight a law in some maps who use predicted species distribution models to show areas of high
species richnesses (e.g., Herkt et al. 2015, 2017; Han et al. 2016). Areas may be shown as potentially
Fig. 2. A. Vegetation map of the CRB. B. Vegetation map of Africa (Blue = Central Congo Basin
Swamp Forest or Lakes, Dark green = Forest to Open Woodland, Light green = Shrubland or Grassland,
Brown = Desert or Semi-Desert). C. Bioclimates map. D. Colour codes used in Figure 2C (all modiied
after Sayre et al. 2013).
8
VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
containing areas of high species richness and important, but no evidence of any specimens exist.
Likewise, areas of high species richness based on specimens, may similarly be underrepresented in the
richness maps where these localities do not conform to the underlying fundamental niche assumptions
for species distribution models. SDM and the species richness derived from these should be used to
prioritize where ield veriication needs to take place (test the model). Any policies derived from such
should only be created or implemented after veriication has taken place.
Holt et al. (2013) identiied 11 zoogeographic realms and 20 regions globally. Within Africa and its
associated islands four zoogeographic realms – Saharo-Arabian, Afrotropical, Madagascan and Oceania
[Reunion and islands eastwards] – were identiied. The Afrotropical realm was further divided into two
zoogreographical regions – Guineo-Congolian and Afrotropical. Africa contains ive zoogeographical
regions. The CRB area contains two of these zoogeographic regions, with the majority of the CRB
containing a signiicant portion of the Guineo-Congolian zoogeographic region, and to a lesser extent
Afrotropical region.
Figure 2 shows an overview of the various vegetation types occurring in the CRB region (A), in Africa
(B), the African bioclimatic zones (C). Note that none of these maps include human land use change – if
the true extent of forest is considered, approximately ⅓ of the area is expected to be destroyed. These
maps represent the status of about 100 to 150 years ago or a theoretical current status, if there were
no humans. The current status most probably would include a much wider ‘new’ Savanna Grassland
habitat, due to the removal of forest, either for agriculture or urbanization (charcoal industry). This
might be the reason why we may be recording more Savanna species in the CRB. Furthermore, given
the fact that we report on specimens that were collected over a timespan of more than a century, the
vegetation type indicated on the map for any given locality might be different from the one at the actual
time the specimen was collected or from the current situation. The fact that a species has occurred in a
speciic locality at a speciic moment in time does not imply that this is currently still the case.
As the African vegetation zones have been shifting over the past century, we decided to use the
Bioclimates data (Sayre et al. 2013: 6) as we suspect this to present a more stable background, although
we cannot ignore the potential inluence of global change.
Collection acronyms
AMNH
BMNH
CAS
CM
DNSM
FMNH
HNHM
HZM
INPBC
KU
MCZ
MHNG
MNHN
MSB
NMBA
NMP
NMW
NMZL
=
=
=
=
=
=
=
=
=
=
=
=
=
=
=
=
=
=
American Museum of Natural History, New York, USA
The Natural History Museum (formerly British Museum of Natural History), London, UK
California Academy of Sciences, San Francisco, CA, USA
Carnegie Museum, Pittsburgh, PA, USA
Durban Natural Science Museum, Durban, South Africa
Field Museum of Natural History, Chicago, IL, USA
Hungarian Natural History Museum, Budapest, Hungary
Harrison Institute (formerly Harrison Zoological Museum), Sevenoaks, UK
Institute of the National Parks of Belgian Congo, Brussels, Belgium
University of Kansas, Natural History Museum, Lawrence, KS, USA
Museum of Comparative Zoology, Harvard University, Cambridge, MA, USA
Muséum d’Histoire Naturelle, Genève, Switzerland
Muséum national d’Histoire naturelle, Paris, France
Museum of Southwestern Biology, Albuquerque, NM, USA
Naturhistorisches Museum, Basel, Switzerland
National Museum (Natural History), Prague, Czech Republic
Naturhistorisches Museum, Wien, Austria
National Museum of Zambia, Livingstone, Zambia
9
European Journal of Taxonomy 382: 1–327 (2017)
OSU
=
RBINS =
RMCA =
RMNH =
SMF
SMNS
TM
=
=
=
TTU
UNIKIS
USNM
WNMU
ZFMK
ZMA
ZMB
ZMH
ZMMU
ZMUC
ZMZ
=
=
=
=
=
=
=
=
=
=
=
Oklahoma State University, Stillwater, OK, USA
Royal Belgian Institute for Natural Sciences, Brussels, Belgium
Royal Museum of Central Africa, Tervuren, Belgium
Naturalis, National Natuurhistorish Museum (formerly Rijksmuseum voor Natuurlijke
Historie), Leiden, the Netherlands
Senckenberg Museum, Frankfurt, Germany
Staatliches Museum für Naturkunde, Stuttgart, Germany
Ditsong National Museum of Natural History (formerly Transvaal Museum), Pretoria,
South Africa
Texas Tech University (The Museum), Lubbock, TX, USA
Université de Kisangani, Democratic Republic of the Congo
United States National Museum, Washington D.C., DC, USA
Western New Mexico University, Silver City, NM, USA
Zoologisches Forschungsinstitut und Museum Alexander Koenig, Bonn, Germany
Zoologisch Museum, Amsterdam, the Netherlands (currently at Naturalis, Leiden)
Zoologisches Museum Berlin, Germany
Zoologisches Institut und Zoologisches Museum, Hamburg, Germany
Zoological Museum, Moscow University, Moscow, Russia
Zoologisk Museum, University of Copenhagen, Denmark
Zoologisches Museum der Universität, Zurich, Switzerland
Additional abbreviations and symbols
♂(♂) =
♀(♀) =
?
=
AC =
AL =
AS =
ad
=
CRB =
DRC =
emb =
imm =
ind(s) =
juv =
MS =
OT =
RSA =
SA =
sad =
SB
=
SC
=
SDM =
SI
=
SK =
SN =
SO =
SS
=
UN =
male(s)
female(s)
unknown (sex)
alcohol and cranium
alcohol, skull not removed
alcohol, skull missing
adult
Congo, Rwanda and Burundi
Democratic Republic of the Congo
embryo
immature
individual(s)
juvenile
mounted skin and skull
other, undeined type
Republic of South Africa
skull and alcohol
subadult
skin, skeleton and skull
skin and cranium
Species Distribution Model
skull in skin
skull only
complete skeleton
skin only
skin and skull
unknown
10
VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
Results and Discussion
Annotated Species List
Class Mammalia Linnaeus, 1758
Order Chiroptera Blumenbach, 1779
Suborder Pteropodiformi Hutcheon & Kirsch, 2006
Pteropodiformes Hutcheon & Kirsch, 2006: 9.
The classical subdivision of the order Chiroptera into two suborders, Megachiroptera and Microchiroptera,
has undergone quite an evolution over the last decennia, which resulted in a rearrangement into the
suborders Yinpterochiroptera and Yangochiroptera (see ACR 2016: 11–13 for a discussion). As the
content of these suborders has changed rather frequently, Hutcheon & Kirsch (2006: 1) suggested
two new subordinal names: Vespertilioniformes (for the group including Emballonuridae, Nycteridae,
and the ‘yangochiropterans’) and Pteropodiformes (for the taxon comprised of Craseonycteridae,
Hipposideridae, Megadermatidae, Rhinolophidae, Rhinopomatidae, and Pteropodidae). However, if
the standardization of the nomenclature suggested by Alonso-Zarazaga (2005) is followed, ‘-formes’
indicates ordinal level, while the subordinal sufix should be ‘-formi’, leading to the suborder names
Vespertilioniformi and Pteropodiformi.
Family Pteropodidae Gray, 1821
* Pteropidae Gray, 1821: 299.
For the correct formation on the family name, see Handley (1980).
The most recent suprageneric division of the Pteropodidae is the one presented by Almeida et al.
(2016: 83), who recognize two subfamilies with representatives on the African continent: Rousettinae
(subdivided into seven tribes: Rousettini: Rousettus; Eonycterini: Eonycteris; Scotonycterini:
Scotonycteris, Casinycteris; Epomophorini: Epomophorus, Epomops, Hypsignathus, Nanonycteris,
Micropteropus; Stenonycterini: Stenonycteris; Myonycterini: Myonycteris [including Lissonycteris],
Megaloglossus; Plerotini: Plerotes), and Eidolinae (Eidolon).
Subfamily Eidolinae Almeida, Giannini & Simmons, 2016
Genus Eidolon Rainesque, 1815
Eidolon helvum (Kerr, 1792)
Fig. 3A–B
Vespertilio vampyrus helvus Kerr, 1792: 91.
* Eidolon helvum (Kerr, 1792).
Hayman et al. (1966: 74) reported a total of 347 specimens from almost all over the DRC. The 560
specimens we found do not lead to a large extension of the range, but we do have some records from
Rwanda and Burundi and from the southern part of Haut Kantanga Province. On the overall distribution,
Thomas & Henry (2013a) show an almost uniform distribution south of the Sahara, with the exception of
the drier areas of NE and SW Africa. Our assement of the distribution seems to suggest a split between a
northern area and a southern/eastern area, which runs through the DRC provinces of Tanganyika, HautLomami and Lualaba. Whether this is a real gap or only a result of the lack of voucher specimens cannot
be determined at this time.
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European Journal of Taxonomy 382: 1–327 (2017)
Subfamily Rousettinae K. Andersen, 1912
Tribe Epomophorini Gray, 1866
Genus Epomophorus Bennett, 1836
Epomophorus anselli Bergmans & Van Strien, 2004
Fig. 3C–D
Epomophorus anselli Bergmans & Van Strien, 2004: 258.
Nesi (2012: 179) reported a single specimen of this species from Kitombongo (Upemba National Park,
Haut-Lomami Province). Bergmans (2013) mentions this species from two (possibly three) localities in
Malawi and one in southwestern Tanzania. Based on these data, the SDM map suggests that E. anselli might
occur from southeastern DRC over northeastern Zambia to southwestern Tanzania and northern Malawi.
Epomophorus crypturus Peters, 1852
Fig. 3E–F
* Epomophorus crypturus Peters, 1852: 26.
* Epomophorus gambianus (Ogilby, 1835): 100.
Some authors, e.g., Nesi et al. (2011: 547), consider E. crypturus and E. gambianus to differ at the subspeciic level only. However, they admit that several recent classiications suggest they differ at the species
level. Claessen & De Vree (1990: 180) indicate that E. gambianus only occurs in the tropical savanna and
woodland zones from Senegal to the Central African Republic, Sudan and Ethiopia. E. crypturus, on the
other hand, only occurs in Zambia, southern Tanzania and southern DRC. Nesi et al. (2011: 547) furthermore
add Mozambique and eastern RSA. As all of the E. gambianus specimens mentioned by Hayman et al.
(1966: 23, map 6) are from the southernmost part of the DRC, these should be assigned to E. crypturus. The
distribution map shows that all of the specimens currently assigned to E. crypturus are from the provinces
of Haut-Katanga, Haut-Lomami and Lualaba, with one exception: RMCA 31557, a subadult female
from Luluabourg (= Kananga) in Kasaï-Central Province. This specimen might need to be re-examined.
Epomophorus labiatus (Temminck, 1837)
Fig. 4A–B
Pteropus labiatus Temminck, 1837: 83.
* Epomophorus anurus Heuglin, 1864: 12.
Bergmans (1988) and Claessen & De Vree (1991) considered E. anurus to be a synonym of E. labiatus.
Bergmans (1997: 72), however, pointed out that some of the E. labiatus specimens identiied by
Claessen & De Vree (1991) might be erroneously assigned to this species, and that their material included
multiple species (e.g., E. anselli Bergmans & Van Strien, 2004). See also Happold (2013a: 245), who
indicated that even more species might be present.
The distribution map indicates that E. labiatus is primarily found along the eastern borders of the
DRC and in Rwanda and Burundi, but also that there are some records from the more central parts
of the DRC. Specimen RMCA 31127 from Léopoldville (= Kinshasa) probably needs reinvestigation.
However, the SDM map indicates that other labiatus specimens have been found in Cameroon, Nigeria
and Ghana, which might suggest that this species might also occur in the western part of the DRC.
Epomophorus minimus Claessen & De Vree, 1991
Fig. 4C–D
Epomophorus minimus Claessen & De Vree, 1991: 216.
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VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
Fig. 3. Distribution maps. A–B. Eidolon helvum (Kerr, 1792). C–D. Epomophorus anselli Bergmans &
Van Strien, 2004. E–F. Epomophorus crypturus Peters, 1852. A, C, E. Distribution in the CRB area. B,
D, F. Pan-African distribution.
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European Journal of Taxonomy 382: 1–327 (2017)
From the same locality as above (Kitombongo), Nesi (2012: 179) also reported the presence of
E. minimus. He also reported it from a second locality near by: Luena. Happold (2013b: 248) indicated
that the species is known from approximately 40 localities in northeastern Africa, in Ethiopia, Somalia,
Kenya and Uganda, which are at quite some distance from these DRC localities. Re-investigation of the
MNHN specimens might, therefore, be necessary to conirm their identity.
Epomophorus minor Dobson, 1880
Fig. 4E–F
Epomophorus minor Dobson, 1880: 715.
* Epomophorus labiatus minor (Dobson, 1880).
Happold (2013a) followed Claessen & De Vree (1991) in considering minor to be a junior synonym of
Epomophorus labiatus. She also pointed out that Bergmans (1997) was strongly opposed to this, as he
pointed out a number errors concerning the E. labiatus specimens used by Claessen & De Vree (1991).
The distribution map shows that the specimens from the eastern part from the DRC it well with the
major (eastern African) distribution area of the species. The specimen from Congo da Lemba (RMCA
408a, a juvenile male) does seem to it with a potential distribution along the Angolan coast, which,
however, is not yet supported by material from that country (ACR 2016).
Epomophorus wahlbergi (Sundevall, 1846)
Fig. 5A–B
Pteropus Wahlbergi Sundevall, 1846: 118.
* Epomophorus wahlbergi haldemani (Halowell, 1846): 52.
Bergmans (1988: 134) discussed the status of the two subspecies of E. wahlbergi and concluded that the
size variations are of a clinal nature and the recognition of subspecies does not seem to be appropriate,
a view which is also followed by Happold (2013c).
The records in Hayman et al. (1966: 24) were primarily from the southern half of the DRC, with two
specimens from Rutshuru (Nord-Kivu Province - INPBC V1346 and V1636), and one specimen from
Ibembo (Bas-Uélé Province - RMCA 20278). The additional material we report on, is also from the
southern part of the DRC, but also from the Nord-Kivu and South-Kivu Provinces, as well as from
Burundi and Rwanda. The locality of a further central Congolese specimen (RMCA 18844b from
Bokungu in Tshuapa Province) is rather unexpected, as Bergmans (1988: 131) stated that E. wahlbergi
is not found moist forests.
Genus Epomops Gray, 1870
Epomops dobsonii (Bocage, 1889)
Fig 5C–D
Epomophorus Dobsonii Bocage, 1889: 1.
* Epomops dobsoni (Bocage, 1889)
Bergmans (1989: 89) suggested that a number of cranial characters (e.g., the pterygoid wing morphology,
the post dental palate and palatal ridges) of E. dobsonii are rather atypical for an Epomops and show
more resemblance with Epomophorus, suggesting that the original generic assignment by Bocage might
be correct. He nevertheless kept the species in Epomops. However, very recently a phylogenetic study
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VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
Fig. 4. Distribution maps. A–B. Epomophorus labiatus (Temminck, 1837). C–D. Epomophorus minimus
Claessen & De Vree, 1991. E–F. Epomophorus minor Dobson, 1880. A, C, E. Distribution in the CRB
area. B, D, F. Pan-African distribution.
15
European Journal of Taxonomy 382: 1–327 (2017)
by Almeida et al. (2016: 83) showed that dobsonii is most closely related to Epomophorus wahlbergi
and these authors transferred the species to the genus Epomophorus.
Distributionwise, E. dobsonii was reported by Hayman et al. (1966: 24, map 9) from the extreme southern
part of the DRC (provinces of Haut-Katanga and Lualaba). Further material from these provinces is
reported in this paper, but also some records from Haut-Lomami Province (near the Upemba National
Park), and from Kasende (MCZ 31738; near Kananga in Kasaï-Central Province). The specimens from
Rwanda (Shangi Schad, RMCA 85.006-M-0463 and Rubona, RMCA 16037) extend the distribution
range of the species. Bergmans (1979), who already reported on the latter specimen, indicated that this
extension was not beyond the already known vegetation types: mopane and miombo woodlands, and the
Central African rain forest block.
Epomops franqueti (Tomes, 1860)
Fig. 5E–F
Epomophorus franqueti Tomes, 1860: 54.
* Epomops franqueti franqueti (Tomes, 1860).
Bergmans (1989: 113) could not ind any differences between E. f. franqueti and the West African
E. f. strepitans K. Andersen, 1910 and considered the latter as a synonym of the nominate form.
Hayman et al. (1966: 24, map 10) provided records from all over the DRC, with the exception of the
former Katanga Province. From that province, we report one specimen: RMCA 10514 from Mulongo, in
the current Tanganyika Province. The Pan-African distribution of the species indicates that is primarily
a forest species, but with some occurrences in savanna areas.
Genus Hypsignathus H. Allen, 1862
Hypsignathus monstrosus H. Allen, 1862
Fig. 6A–B
H[ypsignathus] monstrosus H. Allen, 1862 (for 1861): 157.
* Hypsignathus monstrosus H. Allen, 1862.
The distribution of this large bat is very similar to that of Epomops franqueti. Also, this species occurs
over almost the entire DRC, with the exception of the former Katanga Province. The additional specimens
we were able to track ill in the spaces between the records reported by Hayman et al. (1966: 25, map
11), but do not expand the distribution area reported by these authors.
Genus Micropteropus Matschie, 1899
Micropteropus intermedius Hayman, 1963
Fig. 6C–D
* Micropteropus intermedius Hayman, 1963: 100.
Hayman et al. (1966: 27, map 13) reported two specimens belonging to this species, one from Thysville
(= Mbanza-Ngungu) and one from Luluabourg (= Kananga). One additional specimen was reported by
Bergmans (1989: 101) from “Banana/Netonna”. Here, we add two further specimens from Luluabourg.
All of these localities are relatively close to the Angolan border, where the holotype was collected.
As indicated by Bergmans (1989: 103) a number of these specimens were originally assigned to other,
similar-sized epomophorine species (e.g., M. pusillus or Epomophorus grandis [not yet reported from
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VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
Fig. 5. Distribution maps. A–B. Epomophorus wahlbergi (Sundevall, 1846). C–D. Epomops dobsonii
(Bocage, 1889). E–F. Epomops franqueti (Tomes, 1860). A, C, E. Distribution in the CRB area. B, D,
F. Pan-African distribution.
17
European Journal of Taxonomy 382: 1–327 (2017)
CRB]) due to the careless preservation or examination of the soft palate. Therefore, we cannot exclude
that some of the specimens reported here as M. pusillus might belong to the current species.
Micropteropus pusillus (Peters, 1867)
Fig. 6E–F
Epomophorus pusillus Peters, 1867: 870.
* Micropteropus pusillus (Peters, 1867).
Compared with Hayman et al. (1966: map 14), the distribution for this species has been extended in the
eastern part of the DRC into Burundi, and in the southwestern part of the DRC, but in the northwestern
part of the country it seems to be lacking. The specimens from Boende seem a bit out of place, as
Thomas & Henry (2013b: 271) mentioned that it avoids the Congolian rainforest except around its
edges. Monfort (1992: 142) believes this species occurs in Rwanda, based on a reference to “RuandaUrundi” by Schouteden (1947) and the locality “Congo-Nil-Aka” mentioned by Hayman et al. (1966:
27 - RBINS 13522). Monfort claims this locality to refer to “Nil-Akagera”, which is in Rwanda and
not in Burundi, but Hayman et al. (1966: 81) provide coordinates for “Congo-Nil-Akka”, 04°35′ N,
29°45′ E, which put it in the Garamba National Park on the border with South Sudan.
Genus Nanonycteris Matschie, 1899
Nanonycteris veldkampii (Jentink, 1888)
Fig. 7B
Epomophorus veldkampii Jentink, 1887: 51.
Bergmans (1989: 147) referred to Frechkop (1954), who mentioned two N. veldkampii specimens from
Lusinga (Upemba National Park, Haut Lomami Province). These immature specimens were re-identiied
by Hayman et al. (1966: 28) as Plerotes anchietae (Seabra, 1900).
The Paris museum, however, has three N. veldkampii specimens collected in the DRC by Nicolas Nesi,
Blaise Kadjo and Christiane Denys in May 2010. Unfortunately, no speciic locality for these specimens
was available. The distribution map presented by Fahr (2013a: 278) indicates that this species primarily
occurs in West Africa, from Guinea to western Cameroon, but there is also a record from Bangui in the
Central African Republic. Based on this information, we can assume that N. veldkampii will occur in the
northwestern part of the DRC.
Tribe Myonycterini Lawrence & Novick, 1963
Genus Megaloglossus Pagenstecher, 1885
Megaloglossus woermanni Pagenstecher, 1885
Fig. 7C–D
* Megaloglossus woermanni Pagenstecher, 1885: 245.
* Megaloglossus woermanni prigoginei Hayman, 1966: 26.
In Hayman et al. (1966), Hayman described a separate subspecies for nine skins and skulls collected by
Dr. A. Prigogine in the former Kivu Province and one additional specimen collected by Mr. Schepens in
Lundjulu (currently in Nord-Kivu Province). This subspecies differed from the West African nominate
subspecies in larger forearm length and skull size, and darker colour.
Bergmans & Van Bree (1972) [in Bergmans 1997] found that the forearm length and the greatest length
of the skull of the holotype of M. woermanni from Gabon fell within the range given by Hayman for
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VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
Fig. 6. Distribution maps. A–B. Hypsignathus monstrosus H. Allen, 1862. C–D. Micropteropus
intermedius Hayman, 1963. E–F. Micropteropus pusillus (Peters, 1867). A, C, E. Distribution in the
CRB area. B, D, F. Pan-African distribution.
19
European Journal of Taxonomy 382: 1–327 (2017)
prigoginei, and should, therefore, be considered a synonym of the typical form, a view that has been
accepted by almost all subsequent authors.
Distributionwise, Hayman et al. (1966: 26, map 12) reported this species from the northern part of
the DRC. The additional records we report extend its presence in the southern part of the country, in
the provinces Congo Central, Kwango, Kwilu, Kasaï and Kasaï-Oriental. The southernmost record is
RMCA 33065 from the Kakanda Cave near Likasi (former Jadotville). Its occurrence so far from the
modelled distribution makes it a bit doubtful, but the record was mentioned by Bergmans (1997: 60),
who thereby conirmed its identiication and locality information.
Genus Myonycteris Matschie, 1899
Myonycteris angolensis (Bocage, 1898)
Fig. 7E–F
Cynonycteris Angolensis Bocage, 1898: 133, 138.
* Rousettus (Lissonycteris) angolensis (Bocage, 1898).
The lack of acoustical orientation (as only species with Rousettus s.l.), their different mode of locomotion,
and a number of characters which more closely resembles those of Eidolon and other pteropodids than
of Rousettus, already made Novick (1958) [in Aellen & Brosset 1968] and Lawrence & Novick (1963)
conclude that the subgeneric status of Lissonycteris should be raised to full generic status. Lawrence &
Novick (1963: 4) furthermore postulated from Myonycteris specimens stored in alcohol that the posture
of their wings in life is like that of Lissonycteris. This character, together with the slender rostrum,
made them propose to include the two genera in a myonycterine section. The narrowing of the anterior
palate (which results in the upper tooth rows bending inwards) and the relative heaviness of P4 reported
by Bergmans (1980: 179) conirmed this grouping, as did the very similar karyotype (only a different
polymorphism in pair 1) reported by Haiduk et al. (1981: 223). Genetic sequence analyses by Agnarsson
et al. (2011) and Almeida et al. (2011) both conirmed the close relationship between Lissonycteris
and Myonycteris, and led Nesi et al. (2012: 133) to conclude that Lissonycteris should be considered a
synonym of Myonycteris.
The distribution of M. angolensis across CRB is split into three areas: one in Kongo Central Province in
the extreme west, a second in the extreme southeastern part of the DRC (Haut-Katanga and Haut-Lomami
Provinces) and a third along the eastern border of the DRC, extending into Rwanda and Burundi. More
recently, the species has also been collected in the Kisangani area (Tshopo Province). RMCA 9904a,
reported from Kabalo (Tanganyika Province) by Hayman et al. (1966: 30), was actually collected in the
foothills of the Ruwenzori Mountains (Wim Wendelen, pers. comm.).
Myonycteris relicta Bergmans, 1980
Fig. 8A–B
Myonycteris relicta Bergmans, 1980: 171, 173.
One specimen, identiied as belonging to this species by Norberto Giannini, was collected in the Kasaï
Province (Beyi - Kalonji). Taylor (2013: 274) reported it from the coastal forest mosaic from southeastern
Kenya to southeastern Tanzania and from the Zambezian woodland in eastern Zimbabwe. The Durban
Natural Science Museum also has a specimen from Mozambique (ACR 2016: 998). Based on these data,
the SDM map shows that the species might potentially occur in southern Uganda and in the border area
between the DRC and Tanzania, and that the currently reported locality might be too far to the west.
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VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
Fig. 7. Distribution maps. A–B. Nanonycteris veldkampii (Jentink, 1888). C–D. Megaloglossus
woermanni Pagenstecher, 1885. E–F. Myonycteris angolensis (Bocage, 1898). A, C, E. Distribution in
the CRB area. B, D, F. Pan-African distribution.
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European Journal of Taxonomy 382: 1–327 (2017)
Myonycteris torquata (Dobson, 1878)
Fig. 8C–D
Cynonycteris torquata Dobson, 1878: 71, 76.
* Myonycteris wroughtoni Andersen, 1908: 450.
Bergmans (1976) reviewed the genus Myonycteris and referred to Eisentraut (1963) and Brosset (1966a,
b), who suggested that wroughtoni should be considered a subspecies of M. torquata, a view that
was followed by Hayman & Hill (1971), Jones (1971) and Koopman (1975, 1994). Bergmans (1976:
203–204) found that wroughtoni differs from torquata mainly in its somewhat larger greatest skull and
forearms lengths and by a larger relative M2 length, and concluded that these differences are insuficient
to warrant a subspeciic division.
Hayman et al. (1966: 29, map 16) reported a total of nine specimens, all from the northeastern part of the
DRC: provinces of Bas-Uélé, Haut-Uélé and Nord-Kivu. We were able to ind data for 583 specimens
from almost all over the DRC. The map provided by Thomas & Henry (2013c: 275) is quite different
as they do not mention M. torquata from the provinces of Tshopo and Tshuapa. The reason for this is
probably the extremely recent nature of the collections: from 2010 onwards, although the species was
previously reported from Kisangani (Tshopo Province) by Ifuta et al. (1987: 113).
Tribe Plerotini Bergmans, 1997
Genus Plerotes K. Andersen, 1910
Plerotes anchietae (Seabra, 1900)
Fig. 8E–F
Epomophorus anchietæ Seabra, 1900: 116.
* Plerotes anchietae (Seabra, 1900).
Hayman et al. (1966: 29) mention three specimens from the DRC, two from Lusinga (RBINS 10685 and
10686) and one from Panda (RMCA 26210). The latter specimen, however, was examined by Bergmans
(1989: 147), who found it to be an immature Epomophorus (probably minor). Bergmans (1989: 147)
also referred to another specimen in the RMCA collection that was supposed to be from Likasi (former
Jadotville), but indicated that this bat could not be traced. The information provided on the museum’s
holdings accessed through Wim Wendelen did not mention any specimen from Likasi either.
Tribe Rousettini K. Andersen, 1912
Genus Rousettus Gray, 1821
Rousettus aegyptiacus (E. Geoffroy St.-Hilaire, 1810)
Fig. 9A–B
Pteropus Egyptiacus E. Geoffroy Saint-Hilaire, 1810: 96.
* Rousettus aegyptiacus leachi (Smith, 1823): 433.
Happold (2013d: 290) distinguishes four African subspecies in R. aegyptiacus, of which leachii occurs
from NE DRC to Ethiopia and southwards to South Africa, as well as on the islands of Pemba, Zanzibar
and Maia. A second subspecies, unicolor (Gray, 1870), occurs from Senegal to Cameroon, Bioko Island,
and southwards to Angola. Bergmans (1994: 91) indicates that in leachii, a weaker skull and dentition,
as compared to the typical subspecies, justiies the separation as a subspecies, but he also indicates
that the variation within leachii is very large and that it would be worth examining a larger series from
over its entire distribution area. Hayman & Hill (1971: 12) furthermore mention that the morphological
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Fig. 8. Distribution maps. A–B. Myonycteris relicta Bergmans, 1980. C–D. Myonycteris torquata
(Dobson, 1878). E–F. Plerotes anchietae (Seabra, 1900). A, C, E. Distribution in the CRB area. B, D,
F. Pan-African distribution.
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European Journal of Taxonomy 382: 1–327 (2017)
and geographic limits of aegyptiacus and leachii are not yet clear, and intergrading may occur in the
equatorial zone.
The distribution information provided by Hayman et al. (1966: 30, map 17) suggested that R. aegyptiacus
only occurred along the eastern border of the DRC (Ituri, Nord- and Sud-Kivu Provinces) and in the
extreme south (Haut-Katanga Province) and the extreme west (Kongo Central Province). However,
RBINS 12932 and 12933, reported by Hayman et al. (1966: 30) and Bergmans (1994: 90) from Kivu,
were actually from Buta (Bas-Uélé Province; Tom Geerinckx, pers. comm.). More recently collected
specimens further extend the occurrence of this species to the west of the DRC (Kisangani, Boende).
These records indicate that two subspecies might occur in CRB: leachii in the eastern part of the DRC and
in Rwanda and Burundi, and unicolor in the western part of the country. The subspeciic assignment of
the specimens from the intermediairy localities remains uncertain.
Tribe Scotonycterini Bergmans, 1997
Genus Casinycteris Thomas, 1910
Casinycteris argynnis Thomas, 1910
Fig. 9C–D
* Casinycteris argynnis Thomas, 1910: 111.
Hayman et al. (1966: 21, map 1) only mentioned three specimens from Irangi, Stanleyville and Medje in
the northeastern part of the DRC and one further unnumbered and unmapped specimen from Luluabourg
in the former Kasai Province. Meirte (1983) and Bergmans (1991: ig 4) already expanded its distribution
in the DRC further to the west, and Hassanin et al. (2015: 198) indicated that C. argynnis is endemic
to the Congo Basin, from southeastern Cameroon and eastern Gabon in the west, across southwestern
Central African Republic, the Republic of the Congo and all provinces of the DRC, except Bas-Congo,
to the Mitumba Mountains in eastern DRC. The latter, however, doesn’t seem to be substantiated entirely
by specimens we were able to trace.
Genus Scotonycteris Matschie, 1894
Scotonycteris bergmansi Hassanin et al., 2015
Fig. 9E–F
Scotonycteris bergmansi Hassanin, Khouider, Gembu, Goodman, Kadjo, Nesi, Pourrut, Nakouné &
Bonillo, 2015: 206.
* Scotonycteris zenkeri Matschie, 1894: 202.
Hassanin et al. (2015) reviewed the Scotonycterini tribe, which contained four species: Scotonycteris
zenkeri Matschie, 1894, Casinycteris argynnis Thomas, 1910, C. campomaanensis Hassanin, 2014, and
C. ophiodon (Pohle, 1943). Their analyses of cytochrome b and nuclear DNA revealed that S. zenkeri
should be split up into four clades representing three species: the extralimital (to CRB) S. zenkeri and
S. occidentalis Hayman, 1947, and S. bergmansi Hassanin et al., 2015 occurring in the rainforests
of Gabon, Equatorial Guinea, Republic of Congo, southern Central African Republic, and eastern
Democratic Republic of the Congo. Hassanin et al. (2015) also found suficient differences between
the population from the western part of the distribution area (Gabon to Central African Republic) and
the eastern part of the DRC to recognize two subspecies: the western S. b. bergmansi and the eastern
S. b. congoensis Hassanin et al., 2015.
Hayman et al. (1966: 21) reported this species (as S. zenkeri) from Beni and Kiloboze (with the erroneous
number RMCA 3145), in the extreme eastern part of the DRC. The new records extend the occurrence of
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VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
Fig. 9. Distribution maps. A–B. Rousettus aegyptiacus (E. Geoffroy St.-Hilaire, 1810). C–D. Casinycteris
argynnis Thomas, 1910. E–F. Scotonycteris bergmansi Hassanin et al., 2015. A, C, E. Distribution in the
CRB area. B, D, F. Pan-African distribution.
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European Journal of Taxonomy 382: 1–327 (2017)
the species much more to the west, up to the region around Boende in Tshuapa Province. Kipalu (2009)
reported a specimen of S. zenkeri from near Mbanza-Ngungu (Kongo Central Province), but no details
were given on voucher specimens.
Tribe Stenonycterini Nesi, Kadjo, Pourrut, Shongo, Cruaud & Hassanin, 2012
Genus Stenonycteris Gray, 1870
Stenonycteris lanosus (Thomas, 1906)
Fig. 10A–B
Rousettus lanosus Thomas, 1906: 137.
*Rousettus (Stenonycteris) lanosus Thomas, 1906.
The separation of Stenonycteris from Rousettus was already suggested by Leche (1921) and Kingdon
(1974) [in Bergmans (1994: 111)], but was only proven by genetic sequence analyses (Juste B. et al.
1999; Almeida et al. 2011; Nesi et al. 2012). Using cytochrome b sequences, Nesi et al. (2012) found
that the genus Rousettus was paraphyletic, and that Stenonycteris should be considered as a separate
genus, even constituting its own tribe: Stenonycterini.
Hayman et al. (1966: 20, map 18) reported this species from the Kivu Province in the eastern part
of the DRC. The additional specimens reported here are also mainly from the Nord- and Sud-Kivu
Provinces, with extensions into Rwanda and Burundi. The only outlier is RMCA 36250 from Tshibasha,
near Kananga (Kasaï-Central Province). The SDM map shows that this specimen is the westernmost
representative of the species, but it also shows that the distribution is rather patchy to the east too, with
a cluster of specimens in Kenya, and some dispersed records in Tanzania and Ethiopia.
Family Hipposideridae Lydekker, 1891
Genus Asellia Gray, 1838
Asellia tridens (E. Geoffroy St.-Hilaire, 1813)
Fig. 10C–D
Rhinolophus tridens E. Geoffroy Saint-Hilaire, 1813: 260.
Aulagnier (2013: 362) indicates that Asellia tridens is a desert species that occurs in the Sahara and Sahel
savanna areas of northern Africa, which is quite some distance from the Kivu area in the DRC, where
the 18 specimens reported here have been collected. However, the Paris museum also has a specimen
from Zanzibar, which suggests that the species might occur more to the south than generally accepted.
Genus Doryrhina Peters, 1871
Very recently, Foley et al. (2017: 12) analysed the cytochrome b sequences of the members of the genus
Hipposideros and found this for “H. cyclops” to be that different that they assigned this species to a
separate genus: Doryrhina. This is also conirmed by the karyotype, which contains 36 chromosomes,
whereas the members of the genus Hipposideros only have 32.
Doryrhina cyclops (Temminck, 1853)
Fig. 10E–F
Phyllorrhina cyclops Temminck, 1853: 75.
* Hipposideros cyclops (Temminck, 1853).
Hayman et al. (1966: 44–45, map 51) only mentioned specimens from the northeastern part of the DRC
(provinces of Nord-Kivu, Haut-Uélé, Bas-Uélé and Tshopo). Here, we report on additional specimens
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VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
Fig. 10. Distribution maps. A–B. Stenonycteris lanosus Thomas, 1906. C–D. Asellia tridens (E. Geoffroy
St.-Hilaire, 1813). E–F. Doryrhina cyclops (Temminck, 1853). A, C, E. Distribution in the CRB area.
B, D, F. Pan-African distribution.
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European Journal of Taxonomy 382: 1–327 (2017)
from over almost the entire northern part of the DRC, and down along the Congo River to Kinshasa. The
SDM map is very similar to the one presented by Fahr (2013d: 381), although we do believe that there
is a connection between the populations from the DRC and those from Gabon and Cameroon (see also
ACR 2016: 172).
Genus Hipposideros Gray, 1831
Hipposideros abae J.A. Allen, 1917
Fig. 11A–B
Hipposideros abæ J.A. Allen, 1917: 432.
* Hipposideros abae J.A. Allen, 1917.
Hipposideros abae is in the DRC only known from the extreme northeastern part of the country. Most
records are from the Garamba National Park and from Aba (Haut-Uélé Province). There is one additional
specimen from Digba (Bas-Uélé Province). The map provided by Happold (2013h: 372) shows a very
patchy distribution from Guinea-Bissau in West Africa to northwestern Uganda. Based on climatic data,
we suspect that the species might occur even further to the east through South Sudan to western Ethiopia.
Hipposideros beatus K. Andersen, 1906
Fig. 11C–D
Hipposiderus beatus K. Andersen, 1906: 275, 279.
* Hipposideros beatus Andersen, 1906.
* Hipposideros beatus maximus Verschuren, 1957: 365.
Hayman et al. (1966: 41, map 47) reported this species from the Garamba National Park and Faradje in
Haut-Uélé Province, from Buta in Bas-Uélé Province, and from Ikengo in Equateur Province, all in the
northern part of the DRC. Here, we report on additional specimens from the provinces of Ituri, Mongala,
and Tshopo (also in the northern part of the DRC), Sud-Kivu, and from Thysville (=Mbanza-Ngungu)
in the Kongo Central Province. Also for this species, Happold (2013i: 374) gives a patchy Pan-African
distribution from Sierra Leone in West Africa to the eastern DRC. In this area it is found in closed
forests, degraded areas within the lowland rainforest, coastal woodlands and coastal scrub, and mosaics
of thickets and grasslands, usually (possibly always) near rivers or swamps.
Recently, Heller & Volleth (2016: 3) reported on signiicant differences in echolocation calls for the
H. beatus specimens in the DRC and those from West Africa. They do not explicitly separate the two as
different species, but they suggest this should be looked at further. If this proves to be correct, beatus
might be restricted to West Africa (possibly to the DRC border area) and maximus might need to be
raised to species level.
Hipposideros caffer (Sundevall, 1846)
Fig. 11E–F
Rhinolophus caffer Sundevall, 1846: 118.
* Hipposideros caffer (Sundevall, 1846).
* Hipposideros caffer caffer (Sundevall, 1846).
* Hipposideros caffer tephrus Cabrera, 1906: 358.
* Hipposideros nanus J.A. Allen, 1917: 434.
The identiication of a specimen from Nagero (southern edge of the Garamba National Park, Haut-Uélé
Province; RBINS 14637) as H. caffer tephrus by Hayman et al. (1966: 44) is rather strange, as this
subspecies (currently considered as a separate species) occurs in Morocco and the dry sub-Saharan belt
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VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
Fig. 11. Distribution maps. A–B. Hipposideros abae J.A. Allen, 1917. C–D. Hipposideros beatus
(K. Anderson,1906). E–F. Hipposideros caffer (Sundevall, 1846). A, C, E. Distribution in the CRB
area. B, D, F. Pan-African distribution.
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European Journal of Taxonomy 382: 1–327 (2017)
from Mauritania and Guinea to Sudan and Yemen at about 15° N (Vallo et al. 2009; Bernard & Happold
2013b: 377). Bernard & Happold (2013b: 376) assigned the specimens from the northeastern part of
the DRC to H. c. nanus and those from the western part (Kongo Central Province) to H. c. angolensis
(Seabra, 1898), leaving the remainder to H. c. caffer.
On the map presented by Bernard & Happold (2013b: 377), the distribution of H. caffer only covers the
area around the mouth of the Congo River and the eastern part of the DRC and the western frontiers of
Rwanda and Burundi. Our data indicate that the species occurs much further to the west in the northern
half of the DRC. We therefore believe the distribution is much more continuous, connecting the West
African populations with those from Central and eastern Africa (see also ACR 2016: 164).
Hipposideros camerunensis Eisentraut, 1956
Fig. 12A–B
* Hipposideros camerunensis Eisentraut, 1956: 526.
Hipposideros camerunensis is a very rare species, with a very scattered distribution consisting of a few
specimens from Cameroon, eastern DRC, Uganda and western Kenya. Happold (2013j: 379) suggests
that this disjunct distribution might lead to the possibility that the various populations might be (sub-)
speciically distinct.
Hipposideros fuliginosus (Temminck, 1853)
Fig. 12C–D
Phyllorrhina fuliginosa Temminck, 1853: 77.
* Hipposideros fuliginosus (Temminck, 1853).
Hayman et al. (1966: 45, map 52) mention only three specimens from two localities in the DRC in the
collection of the BMNH: the Ituri River (Ituri Province) and Ngombe (Kasaï Province). The irst one is
probably BMNH 1930.11.11.144, but the second one could not be traced, and is, therefore, not plotted on
the distribution map (Fahr 2013e: 384 also indicated that this specimen needs re-examination). We did
ind some additional specimens from Epulu (Ituri Province) and Yalosemba (Mongala Province), which
resulted in the northern distribution of the species. Fahr (2013e: 384) plotted an additional locality near
Yalosemba, but south of the Congo River, which could not be traced.
Hipposideros ruber (Noack, 1893)
Fig. 12E–F
Phyllorhina rubra Noack, 1893: 586.
* Hipposideros caffer ruber (Noack, 1893).
Although longtime considered to be a subspecies of H. caffer, ruber is currently generally accepted to
represent a separate species. Vallo et al. (2009: 193) recognized a number of lineages in H. ruber, which
in Central Africa include two sister clades, which become sympatric in Cameroon. This suggests that
more than one species is involved, but the names for these taxa have not yet been determined.
The distribution map given by Happold (2013l: 394) passes almost entirely around the DRC, except for
the extreme northeastern and southeastern parts of the country. However, it does include both Rwanda
and Burundi. Our data suggest that the distribution is wider in the DRC, where the species might only
be missing in the central part of the country.
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VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
Fig. 12. Distribution maps. A–B. Hipposideros camerunensis Eisentraut, 1956. C–D. Hipposideros
fuliginosus (Temminck, 1853). E–F. Hipposideros ruber (Noack, 1893). A, C, E. Distribution in the
CRB area. B, D, F. Pan-African distribution.
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European Journal of Taxonomy 382: 1–327 (2017)
Genus Macronycteris Gray, 1866
In their very recently published study of the genus Hipposideros, Foley et al. (2017) found that the Cyt-b
sequences for Hipposideros commersoni E. Geoffroy Saint-Hilaire, 1813, H. cryptovalorona Goodman
et al., 2016, H. gigas (Wagner, 1845) and H. vittatus (Peters, 1852) are quite different from those of the
other representatives of this genus, and therefore, they resurrected the genus Macronycteris Gray, 1866.
This division is also supported by the deviant karyotype, which in Hipposideros counts 32 chromosomes
and 52 in species assigned to Macronycteris.
Macronycteris gigas (Wagner, 1845)
Fig. 13A–B
Rhinolophus gigas Wagner, 1845: 148.
* Hipposideros commersoni gigas (Wagner, 1845).
For a long time “Hipposideros commersoni” was considered to occur across the African continent, but
McWilliam (1982: 9) found gigas to be speciically different from commersoni, which ultimately led to
the restriction of the usage of the name “Hipposideros commersoni” for populations from Madagascar
only. Happold (2013k: 385) indicates that both gigas and vittatus (or its synonym marungensis) were
reported to occur sympatrically, which makes them separate species as well (see also Simmons 2005:
372, 377). Unfortunately, however, as mentioned by these two authors, this split probably makes it
necessary to re-examine all material collected prior to 2005 to assess their identiication.
Although Hayman et al. (1966: 44) report the two “subspecies”, they only provided a map for
H. commersoni gigas (map 50), which also contained the data for the other form. The distribution map
provided by Happold (2013k: 386) indicates that “H. gigas” primarily occurs in western Africa to the
Republic of the Congo, coastal Angola and in the coastal areas around the border between Kenya and
Tanzania. Furthermore, she plots a few isolated records in the eastern part of the DRC. The SDM map
is fairly similar, although we connect the western African populations with those from the DRC and
believe the eastern African populations represent Macronycteris vittatus rather than M. gigas.
Macronycteris vittatus (Peters, 1852)
Fig. 13C–D
Phyllorrhina vittata Peters, 1852: 32.
* Hipposideros commersoni marungensis (Noack, 1887): 272.
The distribution map for “H. vittatus” given by Happold (2013m: 396) indicates that this species
primarily occurs in southern Africa (northern Namibia, northern Botswana, Zimbabwe and southern
Mozambique) and along the coast of Kenya and southern Somalia. Additionally, she plotted a few
individual localities in northern Nigeria, northern Central African Republic, southwestern Angola,
Tanzania and southwestern DRC. This leaves most of the territory of the DRC without any records.
Based on the records we assigned to this species (see also ACR 2016) and the climatic data, we also
found a strong indication that this species primarily will occur in southern Africa, but in a wider band
extending to central Angola, Zambia and southern DRC. Furthermore, we expect it to occur in a narrow
band along the northern frontier of the DRC, extending into southern Cameroon and coastal West Africa.
However, as indicated above, the identiication of these specimens might need re-assessment.
Family Megadermatidae H. Allen, 1864
See Handley (1980) regarding the correct formation of the family name.
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VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
Fig. 13. Distribution maps. A–B. Macronycteris gigas (Wagner, 1845). C–D. Macronycteris vittatus
(Peters, 1852). E–F. Cardioderma cor (Peters, 1872). A, C, E. Distribution in the CRB area. B, D, F.
Pan-African distribution.
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European Journal of Taxonomy 382: 1–327 (2017)
Genus Cardioderma Peters, 1873
Cardioderma cor (Peters, 1872)
Fig. 13E–F
Megaderma cor Peters, 1872: 194.
The distribution area for Cardioderma cor reported by Happold (2013n: 404) covers part of northeastern
Africa, from eastern Sudan at the border with Eritrea, over Eritrea and central Ethiopia and central
Kenya to northern Tanzania, with an extention into western Somalia and some individual localities
in eastern Ethiopia and easternmost Somalia. The westernmost occurrence of C. cor is northeastern
Uganda and just across the border into South Sudan.
The DRC specimen reported here was collected by the Congolese authors at Rubi-Tele in Tshopo
Province, which is a westward extension of about 800 km from the hereto closest known locality.
Genus Lavia Gray, 1838
Lavia frons (Geoffroy, 1810)
Fig 14A–B
Megaderma frons E. Geoffroy Saint-Hilaire, 1810: 192.
* Lavia frons frons (E. Geoffroy St.-Hilaire, 1810).
In CRB, Lavia frons is widely distributed along the eastern part of the DRC, Rwanda and Burundi.
The records from the western part of the DRC are more patchy and were all previously reported by
Hayman et al. (1966: 37, map 35). Its Pan-African distribution stretches in a band between 15° N and
10° S, where it occurs in the woodland savanna zones from West Africa to eastern Africa, in coastal
mosaic, open degraded riverine forests in the Rainforest BZ of the central Congo Basin, East African
Coastal Forest Mosaic and miombo woodland. In arid areas, it is found in bush country surrounding
lakes, marshes and along rivers (Happold 2013o: 407).
Family Rhinolophidae Gray, 1825
Genus Rhinolophus Lacépède, 1799
Rhinolophus alcyone Temminck, 1852
Fig. 14C–D
Rhinolophus alcyone Temminck, 1852: 80.
* Rhinolophus alcyone alcyone Temminck, 1852.
Hayman et al. (1966: 38, map 37) only report two specimens belonging to this species, one from Buluku
and one from the upper Makpe River, both in the Garamba National Park, Haut-Uélé Province. Two
additional specimens are from Lukolela (Mai-Ndombe Province, on the border with the Republic of
Congo) and Scierie Forest (Maniema Province). Although specimens were reported from the Republic
of Congo by Dowsett et al. (1991: 256), these were from the coastal province of Kioulou at quite some
distance from Lukolela. Happold (2013e: 311) mentions that this species primarily occurs in the lowland
rainforests, but also in dense relict and riverine forests north of the rainforest.
Rhinolophus blasii Peters, 1867
Fig. 14E–F
Rhinolophus blasii Peters, 1867: 17.
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VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
Fig. 14. Distribution maps. A–B. Lavia frons (E. Geoffroy St.-Hilaire, 1810). C–D. Rhinolophus alcyone
Temminck, 1852. E–F. Rhinolophus blasii Peters, 1867. A, C, E. Distribution in the CRB area. B, D, F.
Pan-African distribution.
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European Journal of Taxonomy 382: 1–327 (2017)
* Rhinolophus blasii empusa (Andersen, 1904: 378).
In the CRB region, this species is represented by a single specimen from Moba, Tanganyika Province.
Happold (2013f: 312) mentions that R. blasii empusa occurs from the southern DRC to the former
Transvaal (RSA). The map she presents also indicates that the Pan-African distribution for R. blasii
contains three centres of gravity: one in the southern half of Africa, starting in the southern DRC and
Tanzania to the northeastern part of the RSA; a second one in Ethiopia and a third north of the Sahara
in Morocco, Algeria and Tunisia. Based on SDM data, we suspect that there might be a connection
between the southern and the eastern areas, but the connection towards the Maghrabian populations
doesn’t seem to be straightforward. This might warrant further investigation to conirm the speciic
relation between these two separate areas.
Happold (2013f: 313) also plotted a second specimen in the neighbourhood of Likasi (Haut-Katanga
Province), but that specimen could not be traced.
Rhinolophus clivosus Cretzschmar, 1826
Fig. 15A–B
* Rhinolophus clivosus Cretzschmar, 1826: 47.
* Rhinolophus clivosus zuluensis (Andersen, 1904): 383.
Hayman et al. (1966: 38) mention two different taxa: R. clivosus and R. clivosus zuluensis, where the
latter name is applied to specimens from the (south)western part of the DRC (Sud-Kivu, Haut-Lomami
and Haut-Katanga). However, Bernard & Happold (2013a: 316) mention coastal South Africa as region
for the zuluensis subspecies. They do not provide any information on the identity of the subspecies
occurring in the DRC. Based on the boundaries set by Bernard & Happold (2013a: 316), the northern
specimens (from Nord- and Sud-Kivu and from Rwanda and Burundi) might possibly be assigned to
keniensis Hollister, 1916, and those from Haut-Lomami and Haut-Katanga to augur K. Andersen, 1904.
The subspeciic assignment of the westernmost specimen (RMCA 7718 from Kisantu, Kongo Central
Province) cannot be performed as no other specimens from that area are known.
The Rwandan locality here attributed to Hayman et al. (1966) refers to three specimens from BugoyeKisenyi, Western Prefecture (RMCA 12265 to 12267), which were originally assigned to Rhinolophus
fumigatus.
Recent research on nuclear introns by Dool et al. (2016: 202) indicates that R. clivosus is paraphyletic, which
would suggest that some of the currently recognized subspecies might represent valid, separate species.
Stoffberg et al. (2012) already found that the mtDNA control region of the southern African R. clivosus
specimens differed considerably from that of northern populations (as much as they differ from
R. ferrumequinum). They suggest to call these R. geoffroyi Smith, 1829. However, they also point out that
it “… is not currently recognized as a separate species because the name was unidentiiable and the type
specimen apparently lost …” This was already mentioned by Ellerman et al. (1953: 56), Hayman (1954:
284), and Csorba et al. (2003). Stoffberg et al. (2012) do not provide any information that substantiates
the validity of geoffroyi, so we are reluctant to use this name. They also indicate that “R. geoffroyi”
includes four subspecies: geoffroyi in western RSA, zuluensis in eastern RSA, augur in central RSA and
Botswana and zambesiensis in Mozambique, Zimbabwe, Zambia and the southeastern-most part of the
DRC (Sakania Territory, Haut-Katanga Province). The latter area is just southeast of the records from
Haut-Katanga we report here, and seems to suggest that they do not belong to “R. geoffroyi”. Stoffberg
et al. (2012: 6) considered specimens from Kenya to belong to R. clivosus, which might suggest that
the more northern CRB specimens (Sud- and Nord-Kivu, Burundi and Rwanda) could be assigned to
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VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
Fig. 15. Distribution maps. A–B. Rhinolophus clivosus Cretzschmar, 1826. C–D. Rhinolophus eloquens
K. Andersen, 1905. E–F. Rhinolophus fumigatus Rüppell, 1842. A, C, E. Distribution in the CRB area.
B, D, F. Pan-African distribution.
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European Journal of Taxonomy 382: 1–327 (2017)
R. clivosus. On the other hand, Benda & Vallo (2012: 83) indicate that all populations occurring in the
savanna belt from southeastern RSA to Kenya belong to one taxon, for which augur K. Andersen, 1904
is the prior available name. However, pending further research into the systematics of the R. clivosus
complex, we tentatively retain the name R. clivosus for all the CRB specimens.
Cotterill (2002: 166) described a new closely related horseshoe bat (Rhinolophus sakejiensis) from
Kavunda (11°17′ S, 24°21′ E), between the Sakeji and Zambezi Rivers in Zambian Northwest Province.
This locality is less than 10 km from the border with the DRC. It would, therefore, also be very likely
that this species occurs in the DRC, but it has not yet been recovered from there.
Rhinolophus eloquens K. Andersen, 1905
Fig. 15C–D
Rhinolophus Hildebrandti eloquens K. Andersen, 1905: 74.
* Rhinolophus hildebrandti eloquens Andersen 1905.
Although described as a subspecies of hildebrandtii, eloquens was widely accepted to represent a
separate species, already beginning with Andersen (1905: 660; 1907: 31, 34), and further Allen (1939),
Koopman (1975: 389), Simmons (2005: 354) and Cotterill (2013a: 323). However, some authors, such
as Hayman & Hill (1971: 22), placed eloquens in R. fumigatus, whereas Kock (1969: 121) considered
it a subspecies of R. aethiops (which is currently also considered a synonym of fumigatus). Cotterill
(2013a: 324) indicates that eloquens can be distinguished from either fumigatus and hildebrandtii by its
size, but the dimensions show a considerable overlap (see key).
The map presented by Cotterill (2013a: 324) shows a very restricted distribution in eastern Africa, with
occurrences in Kenya, Uganda, South Sudan, Rwanda and extreme northeastern DRC. This agrees very
well with the distribution we postulate, although we believe it might also occur in southern Ethiopia.
Rhinolophus fumigatus Rüppell, 1842
Fig. 15E–F
*Rhinolophus fumigatus Rüppell, 1842: 132.
In their synonymy list for R. fumigatus, Hayman et al. (1966: 38) include Rhinolophus abae J.A. Allen,
1917, and they refer to Koopman, who considered abae to be very close to fumigatus at the species
level and indicated that abae might only be a subspecies of fumigatus. The records for these two taxa
were grouped together in the text, but they still provided two separate maps: 36 for R. abae and 40 for
R. fumigatus. These maps indicate that the two taxa occur in the northeastern RDC. Further fumigatus
records were reported from Rwanda and the southern provinces of Tanganyika, Lualaba and HautKatanga. Hayman et al. (1966) also reported a series from 74 km on the Banana-Boma Road (RMCA
20649, 20651 to 20653), but these were re-identiied as Rhinolophus lobatus (= R. landeri; Wim
Wendelen, pers. comm.). However, two other R. fumigatus specimens have surfaced from the Kongo
Central Province (Thysville: RBINS 22428 and 22429). Cotterill & Happold (2013a: 330) indicate
that R. fumigatus inhabits savanna areas (miombo and mopane woodlands in south-central Africa).
Their distribution map shows a patchy covering of sub-Saharan Africa consisting of a number of red
areas in West Africa, northeast Africa (including NE DRC, Rwanda, Uganda, Kenya and Ethiopia), and
southwestern Africa (Angola and Namibia) as well as a number of individual records in southeastern
Africa (Zambia, Zimbabwe, Mozambique). From the data in ACR (2016) we know that the distribution in
southern Africa is more extensive than Cotterill & Happold’s map suggests, and this probably might also
be the case in West Africa. As such, our distribution map is more extensive, indicating that R. fumigatus
38
VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
occurs over the major part of sub-Saharan Africa, with the exception of the coastal areas in West Africa,
the Central African forest region and extreme southern Africa.
Rhinolophus hildebrandtii Peters, 1878
Fig. 16A–B
Rhinolophus Hildebrandtii Peters, 1878: 195.
* Rhinolophus hildebrandti Peters, 1878.
Hayman et al. (1966: 39) distinguished two forms in R. hildebrandtii; one they identiied to subspeciic
level (see below) and one to speciic level. However, they only included one distribution map (map
41) for these two forms. The distribution within CRB was restricted to the eastern part of the DRC and
Rwanda, with specimens in the Congolese provinces of Nord-Kivu, Ituri and Haut-Uélé in the north on
the one hand and in Tanganyika, Haut-Lomami and Haut-Katanga Provinces in the south on the other
hand. Some of the specimens from Rwanda (Magachi, INPBC V1903; Lugarama, INPBC 3041) were
re-identiied as R. fumigatus, which might account for the remark made by Monfort (1992: 146) on the
species being plotted on map 41 in Hayman et al. (1966), but being absent in their listing. Additional
specimens of R. hildebrandtii are reported from Burundi and the southern Congolese provinces, but
also from Gangala-na-Bodio (Haut-Uélé Province, RMCA 26664 and 26665) and Ibembo (Bas-Uélé
Province, RMCA 89.004-M-0002). Based on their geographic location, the specimens from the Uélé
Provinces might possibly need to be re-assigned to R. eloquens, but even for that species, the Ibembo
specimen was found too far to the west.
Rhinolophus hilli Aellen, 1973
Fig. 16C–D
Rhinolophus hilli Aellen, 1973: 101.
Although the type specimen of Rhinolophus hilli was already collected in 1964, it was only described
after the publication of Hayman et al. (1966). The species is only known from two specimens: the type
specimen from Uwinka, at the top of a hill at 2300 m, and another specimen from Ruta Bansugera
(=Routabansougera) at the bottom of the same hill at 1750 m and 8 km to the northwest (Baeten et al.
1984: 194; Fahr 2013b: 335).
Rhinolophus kahuzi Fahr & Kerbis Peterhans, 2013
Fig. 16E–F
Rhinolophus kahuzi Fahr & Kerbis Peterhans, 2013: 192, 194.
This recently described species is only known from the type specimen (Kerbis Peterhans et al. 2013:
194).
Rhinolophus landeri Martin, 1838
Fig. 17A–B
Rhinolophus landeri Martin, 1838: 101.
* Rhinolophus landeri lobatus Peters, 1852: 41.
The form lobatus is currently still accepted to be a valid subspecies of R. landeri, occurring from Sudan
and Ethiopia, southwards to the former Transvaal (RSA) (Happold 2013g: 340). The nominate subspecies
(R. l. landeri) occurs from Gambia to Cameroon and further south to the mouth of the Congo River. This
means that the specimens from Banana (RMCA 20746 and 20747) and from 74 km on the Banana-Boma
Road (RMCA 20643 to 20653) might be assigned to R. landeri landeri and all other specimens to R. l.
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European Journal of Taxonomy 382: 1–327 (2017)
Fig. 16. Distribution maps. A–B. Rhinolophus hildebrandtii Peters, 1878. C–D. Rhinolophus hilli
Aellen, 1973. E–F. Rhinolophus kahuzi Fahr & Kerbis Peterhans, 2013. A, C, E. Distribution in the
CRB area. B, D, F. Pan-African distribution.
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VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
lobatus. The distribution map presented by Happold (2013g: 340) suggests there might be a connection
between the eastern and western populations in the DRC, but that is not yet supported by evidence.
Based on climatic data, the SDM map suggests that there probably might not be a direct connection
between the eastern and western populations, but that the latter could be connected to the specimens
from Angola, which implies that the specimens from the Kongo Central Province may possibly be
assigned to R. l. angolensis Seabra, 1898. To resolve the subspeciic status of these specimens, further
material from western Gabon, Congo, Angola and DRC is required.
Rhinolophus ruwenzorii Hill, 1942
Fig. 17C–D
* Rhinolophus ruwenzorii J. Eric Hill, 1942: 1.
Rhinolophus ruwenzorii has a very restricted distribution of approximately 360 km2 in the DRC provinces
of Nord-Kivu and Ituri (Fahr 2013c: 342). Some authors (e.g., Smith & Hood 1980) consider ruwenzorii
to be a subspecies of the west African R. maclaudi, which would result in an extremely disjunct
distribution for this species (Baeten et al. 1984). However, Fahr et al. (2002) showed that both forms
differ substantially and should be considered as separate species.
Rhinolophus simulator K. Andersen, 1912
Fig. 17E–F
Rhinolophus simulator K. Andersen, 1912: 384.
The 52 specimens of R. simulator reported here were already collected in the irst half of the 1960s at a
number of localities in the provinces of Haut Katanga, Lualaba and Haut-Lomami (southeastern DRC).
These localities form a westwards extension of the species in central Africa, but Cotterill & Happold
(2013b: 352) mention its presence in northern Zambia. Furthermore they plotted individual localities
in Tanzania, Uganda and South Sudan, as well as in a few small areas in central Ethiopia and Malawi
and in a larger area stretching from southern Zambia, over almost entire Zimbabwe, southwestern
Mozambique and northern RSA. They also report its presence in a few localities in the border area
between Guinea and Liberia and in central Nigeria and western Cameroon. ACR (2016: 1075) reported
additional localities from Botswana and Namibia. Based on bioclimatic data, the species might also
occur in central Angola, which would suggest that the southern African distribution area might connect
to Cameroon along the coast.
Rhinolophus swinnyi Gough, 1908
Fig. 18A–B
* Rhinolophus swinnyi Gough, 1908: 71.
Hayman et al. (1966: 40) mention 25 specimens in the collection of the Tervuren Museum (RMCA) from
Banana, Baudouinville, Kakontwe and Lubudi Cave, but contrary to all other taxa, no museum numbers
were provided. A recent catalog list of the museum (Wim Wendelen, pers. comm.) doesn’t mention any
specimens belonging to this species either.
Based on the currently available data, R. swinnyi is restricted to the extreme southeastern part of the
DRC (Haut-Lomami and Haut-Katanga Provinces). The Pan-African distribution of the species includes
eastern (Tanzania) and southern Africa, where it occurs from the southern DRC, over Zambia, Zimbabwe,
Malawi, southern Mozambique and extreme eastern RSA.
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European Journal of Taxonomy 382: 1–327 (2017)
Fig. 17. Distribution maps. A–B. Rhinolophus landeri Martin, 1838. C–D. Rhinolophus ruwenzorii
J. Eric Hill, 1942. E–F. Rhinolophus simulator K. Andersen, 1912. A, C, E. Distribution in the CRB
area. B, D, F. Pan-African distribution.
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VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
Fig. 18. Distribution maps. A–B. Rhinolophus swinnyi Gough, 1908. C–D. Rhinolophus willardi Kerbis
Peterhans & Fahr, 2013. E–F. Cloeotis percivali Thomas, 1901. A, C, E. Distribution in the CRB area.
B, D, F. Pan-African distribution.
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European Journal of Taxonomy 382: 1–327 (2017)
Rhinolophus willardi Kerbis Peterhans & Fahr, 2013
Fig. 18C–D
Rhinolophus willardi Kerbis Peterhans & Fahr, 2013: 190.
Rhinolophus willardi is known from two localities in the Sud-Kivu Province, where the holotype and three
paratypes were collected (Kerbis Peterhans et al. 2013: 191). These are the only known specimens of this species.
Family Rhinonycteridae Gray, 1866
From their study of 19 nuclear exon and intron gene fragments, Foley et al. (2014) demonstrated that the
genera Cloeotis, Rhinonicteris, Paratriaenops and Triaenops differed signiicantly enough from the genera
Asellia, Aselliscus, Coelops and Hipposideros to assign them to a separate family, the Rhinonycteridae.
Genus Cloeotis Thomas, 1901
Cloeotis percivali Thomas, 1901
Fig. 18E–F
Cloeotis Percivali Thomas, 1901: 28.
* Cloeotis percivali australis Roberts, 1917: 264.
In the CRB area, Cloeotis percivali is restricted to the southeastern Congolese provinces of Haut-Lomami
and Haut-Katanga. Strangely enough, Jacobs (2013: 365) only refers to one locality: Shinkolobwe. Its PanAfrican distribution reaches from southern DRC over Zambia and Zimbabwe to northwestern RSA and
Swaziland. All of these records are assigned to C. p. australis. The nominate subspecies is restricted to
southeastern Kenya and northeastern Tanzania, where it is known from a few isolated specimens.
Suborder Vespertilioniformi Hutcheon & Kirsch, 2006
Family Emballonuridae Gervais, 1855
Genus Coleura Peters, 1868
Coleura afra (Peters, 1852)
Fig. 19A–B
Emballonura afra Peters, 1852: 51.
* Coleura afra (Peters, 1852).
No further locality records for this species were found besides those mentioned by Hayman et al. (1966:
31), although we were able to track additional specimens from these localities. One locality (Kodja Hill,
Haut-Uélé Province) is not indicated on our map as we could not trace the underlying material. Schwarz
(1920: 1050) mentions this/these specimen(s) to be collected by the Alexander-Gosling expedition.
Genus Saccolaimus Temminck, 1838
Saccolaimus peli (Temminck, 1853)
Fig. 19C–D
Taphozous peli Temminck, 1853: 82.
* Taphozous (Saccolaimus) peli Temminck, 1853.
Saccolaimus has variably been assigned (sub)generic status by various authors, but Barghoorn (1977: 5)
ranked it as a genus based on its skull proportions and ossiication, and the morphology of the auditory
bullae (see also ig. 88 in Happold 2013p).
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VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
Fig. 19. Distribution maps. A–B. Coleura afra (Peters, 1852). C–D. Saccolaimus peli (Temminck,
1853). E–F. Taphozous mauritianus E. Geoffroy St.-Hilaire, 1818. A, C, E. Distribution in the CRB
area. B, D, F. Pan-African distribution.
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European Journal of Taxonomy 382: 1–327 (2017)
Our distribution map hardly differs from the one presented by Hayman et al. (1966: map 24). We have
a few extra records in the Nord-Kivu and Ituri Provinces in the northeastern part of the DRC, and in the
Mongala and Tshuapa Provinces in the northwestern part of the country. The general map shows that
S. peli is primarily a forest species.
Genus Taphozous E. Geoffroy Saint-Hilaire, 1818
Taphozous mauritianus E. Geoffroy St.-Hilaire, 1818
Fig. 19E–F
* Taphozous mauritianus E. Geoffroy Saint-Hilaire, 1818: 127.
This species was reported by Hayman et al. (1966: 31, map21) from all over CRB. The extra specimens
we report here ill in a few gaps, but do not add any further information. The general distribution map
indicates that T. mauritianus potentially occurs over the entire sub-Saharan part of Africa, with the
exception of the Kalahari and a major part of the RSA.
Taphozous nudiventris Cretzschmar, 1830
Fig. 20A–B
Taphozous nudiventris Cretzschmar, 1830: 70.
* Taphozous (Liponycteris) nudiventris (Cretzschmar, 1830).
One single specimen from Cell II in the Garamba National Park (Haut-Uélé Province) was reported by
Hayman et al. (1966: 32). No further specimens were found by us. The SDM map of the species indicates
that its occurrence in the extreme northeastern corner of the DRC is not completely unexpected.
Taphozous perforatus E. Geoffroy St.-Hilaire, 1818
Fig. 20C–D
Taphozous perforatus E. Geoffroy Saint-Hilaire, 1818: 126.
* Taphozous sudani Thomas, 1915: 561.
Rosevear (1965: 150–151) compared material of both T. perforatus and T. sudani, and found that the
only character distinguishing the two taxa is the darker pelage in sudani, generally combined with white
wings. He also found a specimen with intermediate characters, which led him to retain sudani only as a
mere subspecies of perforatus. This view was followed by most subsequent authors (e.g., Kock 1969:
74; Hayman & Hill 1971: 16; Koopman 1975: 370).
With the exception of a few extra specimens from the same localities, all the material reported in this
paper was already mentioned by Hayman et al. (1966: 32). The SDM map shows that T. perforatus
primarily occurs in sub-Saharan Africa, above the equator, with a few records reported from northern
Botswana and southern Zimbabwe.
Family Nycteridae Van der Hoeven, 1855
Genus Nycteris G. Cuvier & E. Geoffroy Saint-Hilaire, 1795
Nycteris arge Thomas, 1903
Fig. 20E–F
* Nycteris arge Thomas, 1903: 633.
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VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
Fig. 20. Distribution maps. A–B. Taphozous nudiventris Cretzschmar, 1830. C–D. Taphozous perforatus
E. Geoffroy St.-Hilaire, 1818. E–F. Nycteris arge Thomas, 1903. A, C, E. Distribution in the CRB area.
B, D, F. Pan-African distribution.
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European Journal of Taxonomy 382: 1–327 (2017)
The distribution of this species is primarily restricted to the northern part of the DRC, with a few
specimens from the Upemba National Park (Haut-Lomami Province) and Luebo (Kasaï Province),
already reported by Hayman et al. (1966: 33, map 26), and some additional specimens from the Kananga
(Kasaï-Central Province) and Kilembe (Kwilu Province) areas. In general, its distribution is mainly
restricted to the rainforest zones and the rainforest-savanna mosaics (Fahr 2013f: 442).
Nycteris grandis Peters, 1865
Fig. 21A–B
* Nycteris grandis Peters, 1865: 351, 358.
Both the species distribution model and the distribution within CRB of Nycteris grandis are more or less
similar to those of the previous species. For this species its core distribution within the DRC is in the
northern part of the country. Although the SDM map also indicates that the species occurs in East Africa
down to Zimbabwe and southern Mozambique, it has not been found in the southeastern provinces of
the DRC.
Nycteris hispida (Schreber, 1774)
Fig. 21C–D
Vespertilio hispidus Schreber, 1774: 169, 188.
* Nycteris hispida (Schreber, 1774).
Nycteris hispida has a very wide distribution, both in CRB and Pan-African. In the DRC it is found over
almost the entire country, and it is also known from both Rwanda and Burundi. The SDM map indicates
that it occurs almost over the entire sub-Saharan African continent, with the exception of the extreme
eastern part. It is also missing from Namibia, Botswana and Zimbabwe (except for the most northern parts
of these latter two countries), and from most of the RSA, where it only occurs along the eastern coast line.
Nycteris intermedia Aellen, 1959
Fig. 21E–F
* Nycteris intermedia Aellen, 1959: 218.
In a note on page 35, Hayman et al. (1966) mention for N. intermedia: “A doubtfully valid species”.
However, the multivariate biometrical analyses by Van Cakenberghe & De Vree (1985) showed that this
species can clearly be distinguished from the larger N. arge and the smaller N. nana.
Contrary to the very patchy distribution map presented by Fahr (2013g: 450), we believe this species has
a more extensive distribution over the African rainforest than all of the members of the Nycteris arge
group, since representatives of this species might have been misidentiied as N. arge. However, we do
caution that in the DRC, the species has only been found in the northeastern part of the country (Ituri,
Nord- and South-Kivu Provinces).
Nycteris macrotis Dobson, 1876
Fig. 22A–F
* Nycteris macrotis Dobson, 1876: 80.
* Nycteris aethiopica Dobson, 1878: 162, 165.
For a very long time, up to the early 1960s and 1970s, N. aethiopia has been considered a separate
species (from N. macrotis). Hayman et al. (1966: 33, 35, 36) followed that view, but also referred
to Koopman, who indicated that aethiopica and macrotis are conspeciic. Hayman & Hill (1971: 19)
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VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
Fig. 21. Distribution maps. A–B. Nycteris grandis Peters, 1865. C–D. Nycteris hispida (Schreber, 1774).
E–F. Nycteris intermedia Aellen, 1959. A, C, E. Distribution in the CRB area. B, D, F. Pan-African
distribution.
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European Journal of Taxonomy 382: 1–327 (2017)
still mention the two as separate species, but note that their distinctness is by no means clear, as was
suggested by Koopman (1965) and Kock (1969). The multivariate analyses by Van Cakenberghe &
De Vree (1985) conirmed that the two forms are conspeciic.
Nycteris macrotis has a scattered distribution over CRB, where it occurs in almost all regions, except
for the northwestern part of the DRC. The SDM map suggests a wide distribution across Africa south
of the Sahara.
Nycteris major (K. Andersen, 1912)
Fig. 22C–D
Petalia major K. Andersen, 1912: 547.
* Nycteris major (Andersen, 1912).
Hayman et al. (1966: 36) only mention the type specimen of N. avakubia as representative of N. major
in the DRC. However, on map 32, they have two dots, and neither of them refers to the type locality of
avakubia, which is Avakubi (Tshopo Province). The two dots seem to refer to the two AMNH specimens
reported by Allen (1917: 427) as belonging to N. major (49399 from the Garamba National Park and
49422 from Faradje), but of which Koopman (1965: 6) indicated that they actually belong to N. macrotis.
Koopman (1965) furthermore refers to two additional specimens reported by Schouteden (1947):
one from Macaco (Kasaï Province; reported for the irst time by Kershaw 1923) and one from Boma
(Kongo Central Province). These specimens were re-identiied by Hayman et al. (1966: 35) and Van
Cakenberghe & De Vree (1985: 80) as N. macrotis.
Nycteris nana (K. Andersen, 1912)
Fig. 22E–F
Petalia nana K. Andersen, 1912: 547.
* Nycteris nana (Andersen, 1912).
Hayman et al. (1966: map 33) restricted the occurrence of N. nana in the DRC to the current provinces
of Ituri and Haut-Uélé, with one record from Ibembo (Bas-Uélé Province). All of these are in the
northeastern part of the DRC. Further specimens are recorded from as far west as Duma, on the border
with the Central African Republic, and as far south as Kananga (Central-Kasaï Province). Due to the
regular collection of specimens belonging to this species, Fahr (2013h: 455) doesn’t believe the disjunct
distribution he presented to be the result of insuficient sampling. However, he did not include the record
from Duma (SMF 6390) nor the one from Wafanya (RMCA 89.020-M-0042), so the actual distribution
of the species might be more wide than he suggested. Additionally, Fahr (2013h) also indicates that
N. nana is recorded in drier types of forests than the other representatives of the N. arge group (arge,
intermedia and major).
Nycteris parisii (de Beaux, 1924)
Fig. 23A–B
Petalia parisii de Beaux, 1924 (for 1923): 254.
The systematic position of Nycteris parisii isn’t completely clear yet. Van Cakenberghe & De Vree
(1985: 81) included it in N. woodi K. Andersen, 1914, but Cotterill (2013b: 456) considered it a separate
species.
There is one specimen from Mount Né, near Blukwa (Ituri Province), which is in between the eastern
African records from Ethiopia and Somalia on the one hand and the record from Cameroon on the other
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VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
Fig. 22. Distribution maps. A–B. Nycteris macrotis Dobson, 1876. C–D. Nycteris major (K. Andersen,
1912). E–F. Nycteris nana (K. Andersen, 1912). A, C, E. Distribution in the CRB area. B, D, F. PanAfrican distribution.
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European Journal of Taxonomy 382: 1–327 (2017)
hand. Cotterill (2013b: 457) indicates that it occurs in the Sudan savanna, the Somalia-Masai bushland
and the coastal forest mosaic biotic zones.
Nycteris thebaica Geoffroy, 1818
Fig. 23C–D
Nycteris Thebaicus E. Geoffroy Saint-Hilaire, 1818: 119.
* Nycteris capensis Smith, 1829: 434.
* Nycteris thebaica E. Geoffroy St.-Hilaire, 1818.
As all recent authors, Bernard & Happold (2013c: 458) consider capensis to be a subspecies of N. thebaica,
which is distributed over Tanzania, SE DRC and Zambia to Cape of Good Hope. As such, the records
reported by Hayman et al. (1966: 34, map 27) from the northern provinces of Nord-Kivu and Ituri do not
belong to this form. Furthermore, most of the specimens from Kateke River (RBINS 10813 to 10834)
reported by Hayman et al. (1966: 34) as N. capensis were re-identiied by Van Cakenberghe & De Vree
(1985: 80) as N. macrotis. Most of the records reported here are from the eastern part of CRB (provinces
Ituri, Nord- and South-Kivu, Haut-Lomami and Haut-Katanga in the DRC and in Rwanda). The only
exception is a series of ive specimens, collected by Brian Robbins in Yalosemba in Mongala Province.
The distribution of the species is primarily sub-saharan Africa, but also along the Atlantic coast up to
Morocco and along the River Nile and part of the coast of the Red Sea up to the Mediterranean.
Family Molossidae Gervais, 1856
At the time of Hayman et al. (1966), most of the African molossids were assigned to the genus Tadarida,
which was split up in a number of subgenera. Freeman (1981: 36), however, considered these subgenera
to be valid genera.
Genus Chaerephon Dobson, 1874
Chaerephon aloysiisabaudiae (Festa, 1907)
Fig. 23E–F
Nyctinomus aloysii-sabaudiæ Festa, 1907: 1.
* Tadarida (Chaerephon) aloysii-sabaudiae (Festa, 1907).
* Tadarida (Tadarida) brunnea (Seabra, 1900): 83.
Hayman et al. (1966: 61) mentioned one DRC record, an immature specimen from Avakubi
(AMNH 48934), which according to Koopman (1965: 23) represents T. (C.) aloysiisabaudiae. They
also mentioned (p. 67) one specimen from Eala (RMCA 6565, Equateur Province) belonging to
T. (T.) brunnea. According to Simmons (2005: 449), Nyctinomus brunneus is a synonym of Tadarida
aegyptiaca. However, Fenton & Peterson (1972: 19) re-examined this specimen and found that it
represented T. aloysiisabaudiae. A third specimen was collected in Yalosemba (Mongala Province).
A further specimen (RBINS 13826 from Iso.4, Garamba National Park) was identiied as T. trevori by
Hayman et al. (1966: 65), but was re-identiied as T. aloysiisabaudiae by Peterson (1972: 2). All of
these localities are in the northern part of the DRC. Fahr (2013k: 494) indicates that C. aloysiisabaudiae
is mainly found in the rainforest and adjacent northern and eastern rainforest-savanna mosaics, one
record very marginally in the Sudan savanna. Specimens were reported from Côte d’Ivoire, Ghana,
Cameroon, Gabon, Central African Republic, DRC, South Sudan and Uganda. The SDM map suggests
it might occur as far west as Liberia and as far east as central Ethiopia, western Kenya and northwestern
Tanzania. Furthermore, it might extend southwards to southern DRC and northeastern Angola.
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VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
Fig. 23. Distribution maps. A–B. Nycteris parisii (de Beaux, 1924). C–D. Nycteris thebaica E. Geoffroy
St.-Hilaire, 1818. E–F. Chaerephon aloysiisabaudiae (Festa, 1907). A, C, E. Distribution in the CRB
area. B, D, F. Pan-African distribution.
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European Journal of Taxonomy 382: 1–327 (2017)
Chaerephon ansorgei (Thomas, 1913)
Fig. 24A–B
Nyctinomus ansorgei Thomas, 1919: 318.
* Tadarida (Tadarida) ansorgei (Thomas, 1913).
Hayman et al. (1966: 67, map 108) reported ansorgei from a number of localities in the eastern part of
the DRC: the Garamba National Park and Faradje (Haut-Uélé Province) and from Jadotville (=Likasi)
and Mwadingusha (Haut Katanga Province). The specimen from Vitshumbi (RMCA 17076) was reidentiied by R.L. Peterson as belonging to Chaerephon bivittatus. Additional material was collected in
Rwanda (Gabiro and Rukara) and in the DRC near Lake Kivu (Sud-Kivu Province) and in the Kikwit
area (Kwilu Province). The latter is a major extention to the west of the DRC, but was overlooked by
Cotterrill (2013b: 496). His map shows a scattered sub-Saharan distribution, composed of two larger areas:
central Ethiopia and southwestern Zimbabwe, a few smaller areas in northeastern DRC, northeastern
Zimbabwe, southwestern Kenya and the southern border between Nigeria and Cameroon, and some
individual localities in eastern Côte d’Ivoire, northern Cameroon, northern Central African Republic,
southern South Sudan, southwestern Tanzania, southeastern DRC, Zimbabwe and northern Angola. The
SDM map suggests a potentially wider distribution in southern Africa, reaching further southwards in
Angola towards northern Namibia.
Chaerephon bemmeleni (Jentink, 1879)
Fig. 24C–D
Nyctinomus Bemmeleni Jentink, 1879: 125.
* Tadarida (Tadarida) cistura (Thomas, 1903): 502.
Based on morphological comparisons, Peterson (1971b: 1347) found that cistura is only an eastern
African subspecies of bemmeleni. Hayman et al. (1966: 67, map 110) reported the species from two
localities in the Nord-Kivu Province, and no additional specimens were found.
Fahr (2013l: 498) indicates that C. bemmeleni is known from ive localities in West Africa (Sierra Leone,
Liberia, Côte d’Ivoire and Cameroon) and from approximately 17 localities in eastern Africa, ranging
from South Sudan, over Uganda and northeastern DRC into the border area between Kenya and Tanzania.
Furthermore, he also plotted an additional locality in Maniema Province, but without providing details.
Chaerephon bivittatus (Heuglin, 1861)
Fig. 24E–F
Nyctinomus bivittatus Heuglin, 1861: 4, 13.
One immature specimen belonging to this species is known from the DRC. This specimen was originally
identiied as Tadarida ansorgei, but as indicated above, was re-identiied by R. L. Peterson as belonging
to the current species. Cotterill (2013d: 500) reported this species from a number of scattered areas in
eastern Africa, reaching from northwestern Eritrea over central Ethiopia, western Kenya, the coastal
area between Kenya and Tanzania, northern Zambia, southern Zambia, central Zimbabwe, the border
between Zimbabwe and Mozambique, and also from a small area on the border between Uganda and
South Sudan. This latter area is the closest record of C. bivittatus to the DRC.
Chaerephon chapini J.A. Allen, 1917
Fig. 25A–B
Chærephon (Lophomops) chapini J.A. Allen, 1917: 460, 461.
* Tadarida (Chaerephon) chapini J.A. Allen, 1917.
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VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
Fig. 24. Distribution maps. A–B. Chaerephon ansorgei (Thomas, 1913). C–D. Chaerephon bemmeleni
(Jentink, 1879). E–F. Chaerephon bivittatus (Heuglin, 1861). A, C, E. Distribution in the CRB area. B,
D, F. Pan-African distribution.
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Hayman et al. (1966: 61, map 92) reported chapini from three localities: Banana Creek (Kongo Central
Province), Léopoldville (Kinshasa City Province) and Faradje (Haut-Uélé Province). The two former are
in the western part of the DRC, whereas the latter is in the northeast. There is one additional specimen
from Masha in northwestern Rwanda. Happold & Cotterill (2013: 504) plotted two additional localities in
southeastern DRC, but in their text they only refer to northeastern and southwestern DRC. Furthermore,
they plotted individual localities in Côte d’Ivoire, Ghana, South Sudan, Ethiopia, Uganda, Zambia,
Zimbabwe, and Namibia, as well as small distribution areas in western Kenya, northern Botswana and
the western boundary between Namibia and Angola. The SDM map indicates that the species can occur
in the countries between Ghana and South Sudan, although the Central African Republic doesn’t seem
to be that favorable. Furthermore, it might occur in a major part of Tanzania, Malawi and the western
part of Mozambique.
Chaerephon gallagheri (Harrison, 1975)
Fig. 25C–D
Tadarida (Chaerephon) gallagheri Harrison, 1975: 313.
Also this species was described after the publication of Hayman et al. (1966) and is known from one
specimen only (Harrison 1975: 313; Cotterill 2013f: 305).
Chaerephon leucogaster (A. Grandidier, 1869)
Fig. 25E–F
* Nyctinomus leucogaster A. Grandidier, 1869: 337.
In Mammals of Africa, Happold (2013v: 528) includes leucogaster in Tadarida pumila. However, we
follow Peterson et al. (1995: 162) and Simmons (2005: 434) in retaining it as a separate species. Some of
the forms we consider to be synonyms of leucogaster were treated as synonyms of pumilus by Hayman
et al. (1966: 62): Chaerephon (Lophomops) cristatus J.A. Allen, 1917 and Chaerephon frater J.A. Allen,
1917. Simmons (2005) indicates that the distribution area of this species extends from Ethiopia to
Ghana, including Nigeria, the DRC, Mali, and Madagascar. ACR (2016: 1130) mentions furthermore
specimens from Cameroon, Central African Republic, Chad, Congo, Côte d’Ivoire, Mayotte, Sierra
Leone, Tanzania and Zambia.
Chaerephon major (Trouessart, 1897)
Fig. 26A–B
[Nyctinomus (Nyctinomus) pumilus] var. Major Trouessart, 1897: 146.
* Tadarida (Chaerephon) major (Trouessart, 1897).
Hayman et al. (1966: 61, map 93) reported this species from the Garamba National Park and Faradje and
Aba (Haut-Uélé Province). Additional specimens were found in Rwindi (Nord-Kivu Province), Tandala
(Sud-Ubangi Province) and Mbandaka (Equateur Province). All of these localities are in the northern
half of the DRC, with the latter two on the western side. These additional records are not mentioned by
Happold (2013s: 517), who mapped the species from a number of distinct areas: one starting from the
Garamba National Park along the River Nile up to Abu Hamad in Sudan, a second on the eastern side
of Lake Victoria, a third along the coast of Kenya and Tanzania, and inally a larger area in West Africa,
starting in eastern Guinea and reaching the western part of Nigeria. Based on the material in ACR (2016:
1131) and the climatological data, we believe the species can occur further to the west (to Senegal) and
to the east (in western Ethiopia). Additionally ACR (2016: 1131) refers to material from Cameroon and
the Central African Republic, bridging the gap between 10° E and 33° E referred to by Happold (2013s:
517).
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Fig. 25. Distribution maps. A–B. Chaerephon chapini J.A. Allen, 1917. C–D. Chaerephon gallagheri
(Harrison, 1975). E–F. Chaerephon leucogaster (A. Grandidier, 1869). A, C, E. Distribution in the CRB
area. B, D, F. Pan-African distribution.
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European Journal of Taxonomy 382: 1–327 (2017)
Chaerephon nigeriae Thomas, 1913
Fig. 26C–D
Chærephon nigeriæ Thomas, 1913: 319.
* Tadarida (Chaerephon) nigeriae Thomas, 1913.
* Tadarida (Chaerephon) spillmanni (Monard, 1933): 51.
Contrary to most other changes proposed by Koopman, Hayman et al. (1966: 64) did not follow Koopman
(1965: 21), who suggested that there is “no good reason to regard spillmanni as anything except a
subspecies of T. (C.) nigeriae.” They indicated (p. 62) that the different wing colour separates the two
species. According to their maps (94 and 97), the dark-winged nigeriae occurs in the northeastern part
of the DRC (Haut-Uélé Province) and the light-winged spillmanni in the southeastern part (Tanganyika,
Haut-Katanga and Haut-Lomami Provinces). One additional record comes from Tandala (Equateur
Province), in the northwestern part of the DRC. With the exception of the last record, Cotterill &
Happold (2013d: 524) plotted all of the DRC specimens. The southeastern records are part of an area
also covering the north/western parts of Zambia. A further large area is found in Zimbabwe-Botswana,
and they also drew smaller areas in Ghana, southern Nigeria, northern Cameroon, northern Central
African Republic, central Ethiopia and south-central Angola, and individual localities in Guinea, Niger,
South Sudan, Tanzania and Namibia. The SDM map suggests a much wider distribution in southern
Africa (supported by material from northeastern RSA; ACR 2016: 1133).
Chaerephon pumilus (Cretzschmar, 1826)
Fig. 26E–F
Dysopes pumilus Cretzschmar, 1826: 69.
* Tadarida (Chaerephon) pumila (Cretzschmar, 1826).
Hayman et al. (1966: 62, map 95) reported this species from almost over the entire CRB area, with the
exception of the northwestern and central parts of the DRC. The only more central locality is Stanleyville
(= Kisangani, Tshopo Province), where a single specimen was collected in the 19th century (RMCA 283).
The additional records, we have found, seem to conirm this distribution. The distribution map given
by Happold (2013v: 529) shows two main areas where C. pumilus occurs: one in West Africa, where
it occurs from Senegal to Nigeria and northern Cameroon, although lacking in the coastal forest, and
a second in eastern Africa, from the Red Sea coast in Eritrea over eastern Sudan, South Sudan and
western Ethiopia, eastern DRC, Zambia, northern Zimbabwe, Malawi, Mozambique to northeastern
RSA. It also occurs in southern Kenya, reaching the coast, going north into Somalia and south into
Tanzania and Mozambique. Furthermore, she indicates a peculiar extension across the southern DRC to
reach the Atlantic coast in Cabinda and Angola, and going south almost to the border with Namibia.
ACR (2016: 387, 1135) mentions additional records from Cameroon, Gabon and Congo which narrow
the gap between the western and eastern distribution ranges. However, there still seems to be a gap
in the north between 20° and 25° E in Chad, Sudan, Central African Republic and the DRC (with the
exception of the single specimen from Kisangani), which might continue further in central and eastern
Angola and Namibia towards the RSA. This would suggest that the western DRC specimens, as well as
those from Angola, might be closer related to the specimens from western Africa than to those from the
eastern part of the distribution area. A connection between the western and eastern populations might
indeed be along the line suggested by Happold (2013v), but it might also be that the specimens from
the Kasaï Provinces are the easternmost representatives of the western populations or the westernmost
representatives of the eastern populations.
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VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
Fig. 26. Distribution maps. A–B. Chaerephon major (Trouessart, 1897). C–D. Chaerephon nigeriae
Thomas, 1913. E–F. Chaerephon pumilus (Cretzschmar, 1826). A, C, E. Distribution in the CRB area.
B, D, F. Pan-African distribution.
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European Journal of Taxonomy 382: 1–327 (2017)
Chaerephon russatus J.A. Allen, 1917
Fig. 27A–B
Chærephon russatus J.A. Allen, 1917: 458.
* Tadarida (Chaerephon) russata J.A. Allen, 1917.
The only specimens belonging to this species reported by Hayman et al. (1966: 64, map 96) were from
the type locality in Haut-Uélé Province, northeastern DRC. There is one additional record from Tandala
(Sud-Ubangi Province), northwestern DRC. The distribution map presented by Happold (2013w: 531)
shows that the few known localities, where this species was found, are almost on a straight line reaching
from southern Côte d’Ivoire, southern Ghana, Cameroon, northern DRC to southwestern Kenya.
Genus Mops Lesson, 1842
Subgenus Mops Lesson, 1842
Mops (Mops) condylurus (A. Smith, 1833)
Fig. 27C–D
Nyctinomus Condylurus A. Smith, 1833: 54.
* Tadarida (Mops) condylura (A. Smith, 1833).
* Tadarida (Mops) condylura condylura (A. Smith, 1833).
Hayman et al. (1966: 64) indicated that many of the specimens they reported under this name could
actually belong to one of the next species (M. niveiventer). The major part of the specimens they mention
are from the eastern part of the DRC (provinces Haut-Uélé, Ituri, Nord-Kivu, Sud-Kivu, Maniema, HautLomami, Haut-Katanga) and Rwanda. Their two remaining localities were from more western areas:
Lubondai-Tshimbulu (Kasaï-Central Province) and Léopoldville (Kinshasa City Province). We found
additional specimens from localities in Rwanda and Burundi, the Sud-Kivu, Tanganyika and KasaïCentral Provinces, and from further to the west (Kikwit, Kwilu Province). These localities were all in
the general area already reported by Hayman et al. (1966). However, we also found two records from
the northwestern part of the DRC (Duma and Tandala, both in Sud-Ubangi Province).
The map given by Happold (2013r: 505) shows a major distribution area in eastern Africa, ranging from
southeastern Sudan, over South Sudan, eastern DRC, Uganda, Rwanda, Burundi, southern DRC, most
of Angola, Zambia, Malawi, southern Mozambique to northeastern RSA. In Kenya, there is a small
band along the border with Tanzania from Uganda to the coast, where it turns northwards to reach
into Somalia. The absence of the species in almost entire Tanzania and northern Mozambique is rather
strange, although a single locality is plotted on the coastal border between these two countries. In western
Africa, there are three areas: one encompassing The Gambia, southern Senegal and Guinea-Bissau, a
second in Sierra Leone, and a third covering eastern Côte d’Ivoire to central Nigeria. Happold (2013r)
also marked a small area in Ethiopia. Our data (ACR 2016: 1153) suggest that M. condylurus might
occur more extensively in Ethiopia, as well as in Tanzania and Mozambique, and also further to the
north in West Africa. The distribution in southern DRC and Angola, however, might not be as extensive
as proposed by Happold (2013r).
Mops (Mops) congicus J.A. Allen, 1917
Fig. 27E–F
Mops congicus J.A. Allen, 1917: 467.
* Tadarida (Mops) congica J.A. Allen, 1917.
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VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
Fig. 27. Distribution maps. A–B. Chaerephon russatus J.A. Allen, 1917. C–D. Mops (Mops) condylurus
(A. Smith, 1833). E–F. Mops (Mops) congicus J.A. Allen, 1917. A, C, E. Distribution in the CRB area.
B, D, F. Pan-African distribution.
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European Journal of Taxonomy 382: 1–327 (2017)
Besides the type specimen, Hayman et al. (1966: 65) mention one extra locality: Medje. Additional
occurrences of this species are reported from Tandala (Sud-Ubangi Province), Yalosemba (Mongala
Province) and Tshopo. Fahr (2013m: 508) indicated that congica was found at a few isolated localities
from southern Cameroon (south of the Sanaga River), northern DRC (north of the Congo River) and
Uganda.
See also Mops (Mops) trevori J.A. Allen, 1917 below.
Mops (Mops) demonstrator (Thomas, 1903)
Fig. 28A–B
Nyctinomus demonstrator Thomas, 1903: 504.
* Tadarida (Mops) demonstrator (Thomas, 1903).
For this species, Hayman et al. (1966: 65) mention a number of localities in or near the Garamba National
Park (Haut-Uélé Province). A number of specimens they mentioned as Tadarida (Mops) niveiventer
from Keyberg (near Lubumbashi, Haut-Katanga Province) were re-identiied by R.L. Peterson (Wim
Wendelen, pers. comm.). This is rather strange as Fahr (2013n: 510) indicates that demonstrator only
occurs in the northeastern part of the DRC, northwards along the River Nile to northern South Sudan
and along the Blue Nile in Sudan. He also plotted a number of individual localities in Cameroon, Côte
d’Ivoire, Ghana and Burkina Faso. All of these are north of the equator. However, there is one additional
specimen from Mushe, Rwanda (RMCA 37983), which more or less bridges the gap between HautUélé and Haut-Katanga. Based on climatic data, the SDM map makes the extension towards Rwanda
conceivable, but a further expansion to Haut-Katanga is still dubious.
Mops (Mops) midas (Sundevall, 1843)
Fig. 28C–D
Dysopes midas Sundevall, 1843 (for 1842): 207.
* Tadarida (Mops) midas (Sundevall, 1843).
Hayman et al. (1966: 66, map 103) reported specimens from in and near the Garamba National Park
(Haut-Uélé Province), from Mwiga and Kamohohora (Nord-Kivu Province) and from Uvira (Sud-Kivu
Province). There is one additional specimen from the Garamba Park, which apparently was overlooked
by Hayman et al., as were two further specimens from Bwamanda (Sud-Ubangi Province). These two
latter specimens are also not plotted by Cotterill & Happold (2013c: 519) on their distribution map. This
map indicates that M. midas occurs in a large number of isolated areas and individual localities, primarily
in the eastern and southern part of Africa, and in two small areas in northern Cameroon and southern
Burkina Faso, and one locality in northern Senegal. The SDM map also shows a patchy distribution, but
nevertheless more extensive, especially in southern Africa, but also in the northern part of the distribution
area, extending more northwards than suggested by Cotterill & Happold (2013c: 519).
Mops (Mops) niangarae J.A. Allen, 1917
Fig. 28E–F
Mops niangaræ J.A. Allen, 1917: 468.
* Tadarida (Mops) niangarae J.A. Allen, 1917.
As indicated by Happold (2013u: 522) the speciic status of Mops niangarae remains uncertain. Hayman
et al. (1966: 66) considered it a separate species, but noted that Koopman (1965: 25) already mentioned
that it closely resembled M. congicus, except for the absence of a band connecting the ears. Hayman &
Hill (1971: 60) also consider it a valid species, but refer to Peterson (in litt.), who believed the missing
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VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
Fig. 28. Distribution maps. A–B. Mops (Mops) demonstrator (Thomas, 1903). C–D. Mops (Mops)
midas (Sundevall, 1843). E–F. Mops (Mops) niangarae J.A. Allen, 1917. A, C, E. Distribution in the
CRB area. B, D, F. Pan-African distribution.
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European Journal of Taxonomy 382: 1–327 (2017)
connection between the ears to be an artifact and thought that niangarae is identical to trevori, but not
conspeciic with congicus [contrary to Happold (2013u: 522) and Simmons (2005: 443), who indicated
that Hayman & Hill (1971) consider niangarae to be a subspecies of congicus]. Freeman (1981: 111)
retained niangarae as a distinct species based on its more robust jaw and separated ears. This view is
followed by Simmons (2005: 443), who suggested that a more formal revision of the trevori/congicus
(and niangarae) complex is needed, but Thorn et al. (2009: 68) retain it as a synonym of M. trevori.
Due to the fact that the species is only known from the type specimen, the general distribution map only
shows a single locality mark.
Mops (Mops) niveiventer Cabrera & Ruxton, 1926
Fig. 29A–B
Mops angolensis niveiventer Cabrera & Ruxton, 1926: 594.
* Tadarida (Mops) chitauensis J. Eric Hill, 1937: 2–3.
* Tadarida (Mops) niveiventer (Cabrera & Ruxton, 1926).
Aellen (1966: 71, map 98) reported six specimens from Elisabethville (= Lubumbashi) and Jadotville
(= Likasi) in southeastern DRC. Hayman & Hill (1971: 60) considered chitauensis as a synonym of
niveiventer, which was followed by Koopman (1975: 421), Freeman (1981: 160) and Simmons (2005:
443).
Hayman et al. (1966: 65) did not provide a separate map for this species, but probably included
the specimen on the map for Tadarida (Mops) condylura (map 99). They reported the species from
Astrida (= Butare, Rwanda), Samba (Maniema Province), a number of localities around Elisabethville
(= Lubumbashi) and Jadoville (= Likasi) in Haut-Katanga Province, and from Luluabourg (= Kananga,
Kasaï-Central Province). Additional records were found from Rwanda, Burundi, the area around
Kananga and the area around Kikwit (Kwilu Province).
Cotterill (2013g: 526) mentioned that niveiventer is found in wetter savannas (miombo woodland).
On his map, he marked an area covering southeastern DRC, western Zambia and extreme eastern
Angola and a second area on the border between Zambia and northwestern Mozambique. Furthermore,
he plotted a number of individual localities in Rwanda, the DRC, Zambia and Angola. Our map suggests
a more uniform distribution from central Angola, the entire territory of Zambia and reaching the western
part of Malawi, and a second area connecting the remaining DRC localities with those in Rwanda and
Burundi.
Mops (Mops) trevori J.A. Allen, 1917
Fig. 29C–D
Mops trevori J.A. Allen, 1917: 469.
* Tadarida (Mops) congica J.A. Allen, 1917.
Hayman et al. (1966: 65) considered trevori to be a synonym (or – following Koopman 1965: 26 – at
most as subspecies) of Tadarida (Mops) congica. However, based on size and subtle differences in the
ear and facial features, Peterson (1972: 10) separated the smaller trevori from the larger congicus. This
view was followed by Freeman (1981: 159, 160) and Simmons (2005: 444).
Besides the type specimen, Hayman et al. (1966: 65) mention one extra locality: Iso.4 (RBINS 13826).
As indicated above, however, this specimen was re-identiied as T. aloysiisabaudiae by Peterson
(1972: 2), leaving the type specimen of M. trevori as the only representative of this species in the DRC.
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VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
Fig. 29. Distribution maps. A–B. Mops (Mops) niveiventer Cabrera & Ruxton, 1926. C–D. Mops (Mops)
trevori J.A. Allen, 1917. E–F. Mops (Xiphonycteris) brachypterus (Peters, 1852). A, C, E. Distribution
in the CRB area. B, D, F. Pan-African distribution.
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European Journal of Taxonomy 382: 1–327 (2017)
Happold (2013z: 538) marked the northeastern corner of the DRC and central Uganda as the core distribution
area for trevori, with some further individual localities in Guinea, eastern Côte d’Ivoire, western Ghana,
and western Nigeria. A record from the Central African Republic needs further investigation as it might
represent either congica or trevori (Fahr 2013m; Happold 2013z). Concerning the habitat preferences of
the two species, Peterson (1972: 17) stated that trevori is a forest edge or open forest dweller.
Subgenus Xiphonycteris Dollman, 1911
Mops (Xiphonycteris) brachypterus (Peters, 1852)
Fig. 29E–F
Dysopes brachypterus Peters, 1852: 59.
* Tadarida (Mops) leonis (Thomas, 1908): 373.
Following El-Rayah (1980, 1981), Simmons (2005: 442) and Happold (2013q: 501), we consider leonis
to be a synonym of brachypterus.
Hayman et al. (1966: 66, map 102) report T. (M.) leonis from six localities, of which we were unable
to trace the underlying BMNH specimens for the three southernmost records (Sud-Kivu Province).
The three other localities are in the northeastern part of the DRC (Nord-Kivu, Haut-Uélé and Tshopo
Provinces). The species was further reported from the northwestern part of the country (Yalosemba,
Mongala Province and Tandala, Sud-Ubangi Province), but also from the southwestern part (Thysville
[= Mbanza-Ngungu], Kongo Central Province and Léopoldville [= Kinshasa]). These last four localities
were not plotted on the distribution map presented by Happold (2013q: 502). On her map, she painted
four areas where this species occurs: northeastern DRC (with two individual localities in Uganda), the
east African coast (from Malindi in Kenya to Mozambique Island), coastal West Africa (from Sierra
Leone to western Togo, and from western Nigeria to Equatorial Guinea) expanding to the western part
of the Central African Republic. The SDM map shows an almost similar image, although the western
DCR records connect Happold’s northeastern DRC area with the West African one.
Mops (Xiphonycteris) nanulus J.A. Allen, 1917
Fig. 30A–B
Mops (Allomops) nanulus J.A. Allen, 1917: 477.
* Tadarida (Mops) nanula (J.A. Allen, 1917).
This species was reported by Hayman et al. (1966: 66, map 104) from nine localities in the northeastern,
southeastern and southwestern parts of the DRC. The specimens from Butembo (Nord-Kivu Province)
were re-identiied by Frits De Vree as belonging to M. (X.) spurrelli. Additional specimens were
reported from the Kikwit area (Kwilu Province), Lulimbi (Nord-Kivu Province), the Isa River (Ituri
Province), and Aba (Haut-Uélé Province). All of these are in the same general area. However, M. (X.)
nanulus was also collected at Yalosemba (Mongala Province) and Tandala (Sud-Ubangi Province) in the
northwestern part of the DRC. These two latter localities were not included on the map presented by
Happold (2013t: 521). Her map shows a patchy distribution, with two major areas in the DRC: one in
the southwestern part of the country and a second reaching from western South Sudan over northeastern
DRC through Uganda to western Kenya. Furthermore, there are smaller areas in the border area of
southern Nigeria and Cameroon, southern Ghana and Sierra Leone, as well as individual localities along
the border of Liberia and Côte d’Ivoire and between Benin and Nigeria, in Cameroon, and western
Ethiopia. Our Pan-African distribution map suggests that the species can occur from Sierra Leone to
Togo and from southern Nigeria over Cameroon, northern Congo, southern Central African Republic,
northern DRC, southern South Sudan and Uganda to western Kenya, and as well as from Cameroon,
eastern Gabon, central Congo into southwestern and southcentral DRC.
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VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
Fig. 30. Distribution maps. A–B. Mops (Xiphonycteris) nanulus J.A. Allen, 1917. C–D. Mops
(Xiphonycteris) spurrelli (Dollman, 1911). E–F. Mops (Xiphonycteris) thersites (Thomas, 1903). A, C,
E. Distribution in the CRB area. B, D, F. Pan-African distribution.
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European Journal of Taxonomy 382: 1–327 (2017)
Mops (Xiphonycteris) spurrelli (Dollman, 1911)
Fig. 30C–D
Xiphonycteris spurrelli Dollman, 1911: 211.
* ? Xiphonycteris spurrelli Dollman, 1911.
Although Hayman et al. (1966: 8) did not mention this species explicitly in their ‘Catalog of Species’,
they do include it with a question mark in their ‘Nominal List of Congo Bats’, and in their key, referring
to Koopman (1965: 31), who tentatively identiied a skin from Luluabourg (= Kananga, Kasaï Central
Province) as belonging to this species. The specimens from Butembo (Nord-Kivu Province) were
reported as Tadarida (Mops) nanula, but were re-identiied as belonging to spurrelli (see above). Further
specimens belonging to this species were collected in the northwestern part of the DRC: Yalosemba
(Mongala Province) and Tandala (Sud-Ubangi Province). Happold (2013x: 533) ignored the record
from Luluabourg and also missed out on the northwestern DRC records, mentioning it only from the
northeastern part of the country. Furthermore, she also plotted individual localities from the Central
African Republic, Equatorial Guinea, Togo, western Côte d’Ivoire and southern Guinea, and three
small areas in Cameroon, southern Ghana and the border area between Sierra Leone and Liberia. The
SDM map is very similar, but we suspect the areas to be more continuous north of the Central African
rainforest and the record from Luluabourg suggests it also occurs south of the rainforest. Whether it
occurs in the forest itself is not yet clear.
Mops (Xiphonycteris) thersites (Thomas, 1903)
Fig. 30E–F
Nyctinomus thersites Thomas, 1903: 634.
* Tadarida (Mops) thersites (Thomas, 1903).
Hayman et al. (1966: 66, map 106) mentioned this species from ive localities: two in the northeast of the
DRC (Avakubi, Tshopo Province and Medje, Haut-Uélé Province) and three in the southwestern part of
the country (Luluabourg [= Kananga], Kasaï Central Province, Luebo, Kasaï Province and Ndwa, MaiNdombe Province). Additional specimens have been collected at a number of localities in Rwanda, in
Kwilu Province (Mosenge and Kikwit), in Kongo Central Province (Banana), but also in Yalosemba and
on the road between Bombanga and Bumba (Mongala Province) and Tandala (Sud-Ubangi Province), in
the northwest of the DRC. Although the SDM map shows more red than the one presented by Happold
(2013y: 536), it looks very similar, with a northern and southern-central branch in the DRC, passing
around the Congolian rainforest. We, however, suspect that the two branches might reunite in the eastern
part of the DRC.
Genus Myopterus E. Geoffroy Saint-Hilaire, 1818
Myopterus daubentonii Desmarest, 1820
Fig. 31A–B
Myopterus daubentonii Desmarest, 1820: 132.
* Myopterus albatus Thomas, 1915: 469.
Koopman (1989: 9) indicated that he believed that the larger forms of the genus Myopterus (daubentonii,
senegalensis and albatus) are all the same species. Adam et al. (1993: 317) suggested that M. albatus
might only be a mere subspecies of M. daubentonii. This view was also followed by Simmons (2005:
446) and Fahr (2013i: 476). Besides the specimens from Bunia (Ituri Province), Niangara (Haut-Uélé
Province) and the Uélé River mentioned by Hayman et al. (1966: 68, map 113), no further representatives
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Fig. 31. Distribution maps. A–B. Myopterus daubentonii Desmarest, 1820. C–D. Myopterus whitleyi
(Scharff, 1900). E–F. Otomops martiensseni (Matschie, 1897). A, C, E. Distribution in the CRB area.
B, D, F. Pan-African distribution.
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of this species are known from the CRB area. The distribution map provided by Fahr (2013i: 477)
includes additional specimens from northern Central African Republic, Côte d’Ivoire and Senegal.
Myopterus whitleyi (Scharff, 1900)
Fig. 31C–D
Mormopterus Whitleyi Scharff, 1900: 569.
* Myopterus whitleyi (Scharff, 1900).
Hayman et al. (1966: 68, map114) reported M. whitleyi from three localities: Léopoldville (= Kinshasa,
Kinshasa City Province), Luluabourg (= Kananga, Kasaï Central Province) and Beno (Mai-Ndombe
Province), all in the southwestern part of the DRC. Additional specimens were collected in the Kikwit
area (Kwilu Province) and in Banana (Kongo Central Province), also in the southwestern part of the
DRC. One further specimen, however, was collected in the northeastern part of the DRC: 10 km N of
Kasenye (Ituri Province). Aellen (1952: 99) reported the species from Coquilhatville (= Mbandaka,
Equateur Province), but that record could not be traced. The distribution map presented by Fahr (2013j:
478) shows two red areas: one encompassing the southwestern DRC records (except for Banana, which
is plotted separately) and a second in coastal Nigeria/Cameroon (from the mouth of the Niger to the
Mount Cameroon area). Furthermore, he plotted single localities in Ghana, the Central African Republic,
Gabon, and Uganda. The SDM map looks very similar, although it doesn’t reach Ghana, as we were
unable to trace the speciic locality for that country. The map also suggests that M. whitleyi does not
occur in the Congolian rainforest, but encircles it both on the north and the south, and the two branches
meet again in the Ugandan-Tanzanian border area.
Genus Otomops Thomas, 1913
Otomops martiensseni (Matschie, 1897)
Fig. 31E–F
Nyctinomus martiensseni Matschie, 1897: 84.
* Otomops martiensseni (Matschie, 1897).
Otomops martiensseni was reported by Hayman et al. (1966: 68, map 112) from three localities in the eastern
part of the DRC (Poko in Bas-Uélé Province, the Garamba National Park in Haut-Uélé Province and the
Upemba National Park in Haut-Lomami Province) and from one locality in Rwanda (Kigali). Additional
specimens were collected in Burundi (Rwegura and the Kibira National Park) and in southeastern DRC
(Marungu, Tanganyika Province). Yalden & Happold (2013: 481) indicate that the distribution of this
species in Africa is wide, but disjunct, with records from Côte d’Ivoire and Ghana in West Africa, the
Central African Republic, DRC, Uganda, Rwanda and Angola in Central Africa, the RSA, Zimbabwe and
Malawi in southern Africa, and a range along the border between Tanzania and Kenya, reaching to the
border with Uganda. They also mark a few small areas in Ethiopia and Djibouti. However, the specimens
from Ethiopia, Djibouti, Eritrea and Kenya have been found to represent a recently described species,
Otomops harrisoni Ralph, Richards, Taylor, Napier & Lamb, 2015. Ralph et al. (2015: 27) also indicate
that the range of their new species might include additional localities in Uganda, and their MapEnt map
on page 17 suggests that this species also has a high probability of occurring in the CRB area.
Genus Tadarida Rainesque, 1814
Tadarida fulminans (Thomas, 1903)
Fig. 32A–B
Nyctinomus fulminans Thomas, 1903: 501.
* Tadarida (Tadarida) fulminans (Thomas, 1903).
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Fig. 32. Distribution maps. A–B. Tadarida fulminans (Thomas, 1903). C–D. Tadarida ventralis
(Heuglin, 1861). E–F. Miniopterus fraterculus Thomas & Schwann, 1906. A, C, E. Distribution in the
CRB area. B, D, F. Pan-African distribution.
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Hayman et al. (1966: 68, map 111) reported T. fulminans from two localities: Nyakibanda (Rwanda) and
Rumangabo (Nord-Kivu Province). Additional specimens were collected from two further localities in
Rwanda: Nyanza and Rukara. On the African continent, T. fulminans is restricted to eastern Africa, from
Kenya to extreme northern RSA. On his distribution map, Cotterill (2013e: 512) marked three areas: on
the border between Kenya and Uganda, in northeastern Zimbabwe, reaching into Mozambique and in
northwestern Zimbabwe, reaching into Zambia. All other occurrences are individual localities. Based
on climatic data, the SDM map indicates a potential wider occurrence in southeastern Africa and two
areas in the northern part: one in northeastern DRC, reaching into Uganda and Rwanda, and another in
western Kenya, with a corridor-like connection between the latter and the southern range.
Tadarida ventralis (Heuglin, 1861)
Fig. 32C–D
Nyctinomus (Dysopes) ventralis Heuglin, 1861: 4, 11.
* Tadarida (Tadarida) africana (Dobson, 1876): 348.
Hayman et al. (1966: 67, map 107) reported that only one specimen was found in the CRB area: RMCA
29241 from Alima (Nord-Kivu Province). Until that time, T. africana was considered to be a valid
species, occurring in southern Africa. Kock (1975: 6) discussed the features of an original specimen of
Nyctinomus ventralis and concluded that africana is a synonym of ventralis. This view is followed by
most recent authors.
No further specimens from the CRB area were found.
Cotterill (2001a: 201) indicates that T. ventralis has a wide and highly scattered distribution across
eastern tropical Africa, which is conirmed by the distribution map presented by Cotterill (2013h: 539).
The range covers a major part of western Kenya and the joining area in northern Tanzania. Furthermore,
he marked individual localities in Eritrea, Ethiopa, South Sudan, DRC, Zambia, Malawi, northwestern
Mozambique, Zimbabwe and possibly northeastern RSA. The SDM map looks very similar, although
we excluded the RSA records, as we could not trace the speciic specimens.
Family Miniopteridae Dobson, 1875
During a long period, the genus Miniopterus was considered to represent a separate subfamily within the
Vespertilionidae. However, based on dental characters, the shape of the cranium, the double articulation
of the humerus-scapulum, the length of the second phalanx of the third inger, the karyotype, the absence
of a baculum and reproductive differences, Mein & Tupinier (1977: 210) suggested that the distinction
should be made at the family level. This was later conirmed with molecular data (Hoofer & Van Den
Bussche 2003: 1; Miller-Butterworth et al. 2007: 1553).
Genus Miniopterus Bonaparte, 1837
As mentioned by Happold (2013ap: 711), representatives of the genus Miniopterus show a remarkable
degree of morphological uniformity, which makes it dificult to identify the various forms. In continental
Africa, she recognizes six species, of which we believe four occur in the CRB region. However, we
cannot exclude that some of the specimens were erroneously identiied.
Miniopterus fraterculus Thomas & Schwann, 1906
Fig. 32E–F
Miniopterus fraterculus Thomas & Schwannn, 1906: 162.
* Miniopterus schreibersi villiersi Aellen, 1956: 890.
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Koopman (1965: 18) refers to Harrison (1953: 71), who revised the Miniopterus forms of southern Africa,
where he distinguished two species: “the large natalensis and the small fraterculus”. Unfortunately,
however, this distinction doesn’t seem to hold (see key).
Hayman et al. (1966: 59–60) assigned most of their material to Miniopterus schreibersi villiersi and only
15 specimens from two localities to M. s. vicinior. The latter are plotted on their map 90, but no separate
map was presented for M. s. villiersi. Apparently, these specimens were included on map 88 (M. inlatus).
In “Mammals of Africa”, Bernard & Happold (2013d: 713; 2013e: 719) restrict M. fraterculus to the
northwestern part of the DRC and M. natalensis to the southeastern part. We are following this division
for the CRB material, with specimens from the DRC provinces of Haut-Uélé, Ituri, Nord-Kivu, and
Sud-Kivu, as well as from Rwanda and Burundi being assigned to M. fraterculus and those from the
provinces of Tanganyika, Haut-Lomami and Haut-Katanga to M. natalensis. However, we also assigned
records from more southwestern localities to the latter species: Luluabourg (= Kananga, Kasaï-Central
Province) and near Thysville (= Mbanza-Ngongo, Kongo Central Province).
See also Miniopterus natalensis (A. Smith, 1833) below.
Miniopterus inlatus Thomas, 1903
Fig. 33A–B
Miniopterus inlatus Thomas, 1903: 634.
* Miniopterus inlatus rufus Sanborn, 1936: 112.
With the exception of FMNH 152883 and 152884 from 32 km SE of Mbuji-Mayi (Kasaï Oriental
Province), all other specimens are from the eastern part of the CRB area (DRC provinces of Haut-Uélé,
Ituri, Tanganyika, Haut-Lomami and Haut-Katanga; Rwanda and Burundi).
Happold (2013aq: 715) indicated that M. inlatus occurs widespread south of the Sahara, but mostly
in isolated localities. Contrary to her map, we excluded the specimens from Ethiopia, as we follow
Peterson et al. (1995: 120), Lavrenchenko et al. (2004b: 144), Simmons (2005: 519) and Juste et al.
(2007: 28) in considering africanus as a separate species. We also believe that the species occurs far
further to the south, into the RSA.
Miniopterus minor Peters, 1867
Fig. 33C–D
* Miniopterus minor Peters, 1867 (for 1866): 870.
Hayman et al. (1966: 60) reported M. minor from one locality: Thysville (= Mbanza Ngungu, Kongo
Central Province). Two specimens from caves near Thysville (RMCA 10061, 22501) were originally
identiied as M. schreibersi villiersi, but were re-identiied as M. minor, and this was also the case for a
specimen from Kisala (INPBC W1462). All these localities are in the extreme western part of the DRC.
Happold (2013ar: 717) indicates that M. minor is distributed over three areas in continental Africa: on
São Tomé, in western DCR and southwestern Congo and in the coastal boundary area between Kenya
and Tanzania. The species is furthermore also reported from the Central African Republic, the Comoro
Islands and from Madagascar (ACR 2016: 1191).
Miniopterus natalensis (A. Smith, 1833)
Fig. 33E–F
Vespertilio Natalensis A. Smith, 1833: 59.
* Miniopterus natalensis arenarius Heller, 1912: 2.
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Fig. 33. Distribution maps. A–B. Miniopterus inlatus Thomas, 1903. C–D. Miniopterus minor Peters,
1867. E–F. Miniopterus natalensis (A. Smith, 1833). A, C, E. Distribution in the CRB area. B, D, F.
Pan-African distribution.
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VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
* Miniopterus schreibersi vicinior J.A. Allen, 1917: 450.
* Miniopterus schreibersi villiersi Aellen, 1956: 890.
Originally described as a subspecies of breyeri, vicinior was moved to M. natalensis by Sanborn (1936:
110). However, most later authors assigned the form to M. schreibersi, but we follow Simmons (2005:
521) and Monadjem et al. (2013: 138), who included it back in M. natalensis. On the other hand, Thorn
et al. (2009: 49) included vicinior specimens from Uganda in M. fraterculus.
Aellen (1956) described villiersi as a subspecies of M. inlatus. Most recent authors, though, include it
in M. schreibersii, but Fahr et al. (2006: 173), Decher et al. (2010: 170) and Schaer et al. (2013: 17416,
suppl. 1) consider it a valid species, restricted to western Africa. Decher et al. (2016: 274, 276) are a
bit dubious in using M. inlatus villiersi in their systematic review and M. villiersi in their discussion
sections. Furthermore, Anciaux de Faveaux (1978a: 479) mentioned that he originally assigned all of his
material to either Miniopterus inlatus villiersi or Miniopterus natalensis, but that – according to Felten
and Aellen – actually almost all of his specimens belonged to Miniopterus schreibersi natalensis. Based
on this, we consider the DRC villiersi specimens to represent M. natalensis.
See also Miniopterus fraterculus Thomas & Schwann, 1906 above.
Family Vespertilionidae Gray, 1821
Simmons (2005) subdivided the African Vespertilionidae into four subfamilies: Kerivoulinae, Myotinae,
Vespertilioninae and Miniopterinae. The latter has been raised to family level (see above), whereas
Koubínová et al. (2013: 9) distinguished one additional subfamily: Scotophilinae.
Subfamily Kerivoulinae Miller, 1907
Genus Kerivoula Gray, 1842
Kerivoula argentata Tomes, 1861
Fig. 34A–B
* Kerivoula argentata Tomes, 1861: 31–32.
Eight specimens from three localities in the southern part of the DRC were mentioned by Hayman et al.
(1966: 58, map 84). More recently, one additional specimen was collected much more to the north in
Boende (Tshuapa Province). The map given by Cotterill (2013i: 726) indicates that the core distribution
area for K. argentata covers western Mozambique, eastern Zimbabwe, southern Malawi, southeast to
northwest Zambia and southeastern DRC. Furthermore, he plotted some individual localities in the
RSA, Namibia, Tanzania, Kenya, Angola (?) and Luluabourg (= Kananga) in the DRC. Our distribution
map (based on data from ACR 2016: 1199) suggests a distribution which is restricted to the southeastern
part of the continent, reaching from the northeastern part of the RSA to southern Kenya, whereby the
records from Luluabourg and Boende seem to be located too far to the northeast and should probably
be re-examined.
Kerivoula cuprosa Thomas, 1912
Fig. 34C–D
Kerivoula cuprosa Thomas, 1912: 41.
In the DRC, Kerivoula cuprosa is known from three localities: Koteli (Bas-Uélé Province), Eala
(Equateur Province) and Wafanya (Tshuapa). The specimens from Koteli and Eala were reported as
K. smithii by Hayman et al. (1966: 59), but were re-identiied by Jakob Fahr as K. cuprosa (Fahr 2013r:
728). Kityo et al. (2009: 135) indicate that four cuprosa specimens were reported by Schouteden (1947):
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European Journal of Taxonomy 382: 1–327 (2017)
Eala, Koteli, Bambesa and Lisala, but were not discussed by Hayman et al. (1966). However, three
of these were explicitly included in K. smithii by Hayman et al. (1966: 59), and the remaining one
(RMCA 14263) in reality came from Gumba in the Lisala Territory. RMCA 14263 actually includes
two specimens, one of which was identiied as a Nycteris sp. by Danny Meirte and one as a Kerivoula
sp. by Jakob Fahr (Wim Wendelen, pers. comm.). Besides the three currently reported DRC localities,
Fahr (2013r: 728) also mentions this species from Cameroon, Côte d’Ivoire and Guinea. The SDM
map shows that cuprosa is restricted to the central African rain forest block, but this should probably
be extended to the West African block as well (the records from Côte d’Ivoire and Guinea were not
included when generating this map, as the underlying specimens could not be traced).
Kerivoula lanosa (A. Smith, 1847)
Fig. 34E–F
Vespertilio lanosus A. Smith, 1847: pl. 50 and text.
* Kerivoula harrisoni muscilla (Thomas, 1906): 294.
* Kerivoula harrisoni lucia (Hinton, 1920): 240.
Rosevear (1965: 307–308) doubted the validity of the characters separating harrisoni from lanosa,
i.e., the number of cusps of the upper incisors (two in harrisoni and one in the other species) and the
length of the rostrum (comparing c-m3 with m3-m3, the irst being greater than the second in harrisoni
and smaller in the other species). This was further investigated by Hill (1977: 628), who considered
harrisoni to be a subspecies of lanosa.
Hayman et al. (1966: 58, map 86) mention three specimens from the eastern part of the DRC: one
from Pawa (Haut-Uélé Province) [considered to belong to K. harrisoni bellula Aellen, 1959 by Aellen
1959: 225], one from Lukulu (Tanganyika Province) and one from Funda Biabo (Lualaba Province).
One additional specimen was collected along the road between Kamituga and Kampene in Sud-Kivu
Province. The SDM map conirms the distribution suggested by Cotterill (2013j: 730), who mentions it
is known from widespread, scattered records in the rain forest, the northern rain forest-savanna mosaic,
Somalia-Masai bushland, coastal forest mosaic, Zambezian woodland, Afromontane-Afroalpine and
south-west Cape biotic zones, but we suggest it might occur further to the west in northern Namibia and
south-central to coastal Angola.
Kerivoula phalaena Thomas, 1912
Fig. 35A–B
Kerivoula phalæna Thomas, 1912: 281.
* Kerivoula phalaena Thomas, 1912.
Although not separately listed in their catalog part, Hayman et al. (1966: 57) refer to this species, as
Koopman (1965: 31) mentioned a specimen from Lukolela (Mai-Ndombe Province) in the AMNH. Two
additional specimens have been collected since then: one from Wafanya (Tshuapa Province) and one
from Irangi (Sud-Kivu Province). Fahr (2013t: 732) indicated that K. phalaena is mainly recorded
from the rainforest zones in western and central Africa, with outlying records from the AfromontaneAfroalpine zones in northeastern DRC and western Uganda and from the Guinea Savanna BZ in Côte
d’Ivoire. This is conirmed by our map, which is based on climatic data and suggests that K. phalaena
might occur even further to the east.
Kerivoula smithii Thomas, 1880
Fig. 35C–D
Kerivoula Smithii Thomas, 1880: 166.
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Fig. 34. Distribution maps. A–B. Kerivoula argentata Tomes, 1861. C–D. Kerivoula cuprosa Thomas,
1912. E–F. Kerivoula lanosa (A. Smith, 1847). A, C, E. Distribution in the CRB area. B, D, F. PanAfrican distribution.
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* Kerivoula smithi Thomas, 1880.
* Kerivoula ? aerosa Tomes, 1858: 333.
Hayman et al. (1966: 58) mention a specimen from Bukavu (RMCA 23172; Sud-Kivu Province) as
possibly belonging to “K. aerosa”. They also indicated that there is some doubt about the African origin of
the types of this species. As aerosa is currently assigned to the mainly southeast Asian genus Phoniscus,
it has been suggested that they actually came from the Malay Archipelago (Happold 2013as: 735). In the
meantime, the Bukavu specimen has been re-identiied by Jakob Fahr as belonging to K. smithii. He also
re-identiied three other animals: one from Koteli (RMCA 16190; Bas-Uélé Province), one from Eala
(RMCA 16189; Equateur Province) and one from Gumba Island (RMCA 14263; Mongala Province).
The two former represent K. cuprosa, whereas the latter could only be identiied as Kerivoula sp.
We also report four additional localities: Kalehe Territory (Sud-Kivu Province), Ukaika (Ituri Province),
Aketi (Bas-Uélé Province) and Wafanya (Tshuapa Province), with the exception of the latter all in the
northeastern part of the DRC. Fahr (2013u: 733) indicates that K. smithii is primarily found in the rain
forest zones of southeastern Nigeria to the DRC and from the Afromontane-Afroalpine and the SomaliaMasai bushland zones in Kenya. The SDM map conirms this view.
Subfamily Myotinae Tate, 1942
Genus Myotis Kaup, 1829
Myotis bocagii (Peters, 1870)
Fig. 35E–F
Vespertilio Bocagii Peters, 1870: 125.
* Myotis bocagei (Peters, 1870).
* Myotis bocagei bocagei (Peters, 1870).
* Myotis bocagei hildegardeae Thomas, 1904: 209.
* Myotis bocagei cupreolus Thomas, 1904: 407.
Besides 40 specimens identiied to the speciic level, Hayman et al. (1966: 51) recognized three subspecies,
of which hildegardeae is currently included in the nominate subspecies. This agrees with a statement
made by Hayman [in Fain 1953: 99]: “ … since the more specimens [of M. b. hildegardeae] I see, the
less I think it possible to maintain hildegardeae as a valid race”. Happold (2013an: 692) indicates that
M. b. bocagii occurs in the woodland savannah from Ethiopia to Angola and the former Transvaal, and
M. b. cupreolus in the rainforest from Liberia to the DRC. In the CRB area, Hayman et al. (1966, map
68) reported M. bocagii from the eastern part (provinces Bas-Uélé, Haut-Uélé, Ituri, Nord-Kivu, SudKivu, Tanganyika, Haut-Lomami and Haut-Katanga), Rwanda, and from the western part (provinces
Kongo Central, Kinshasa and Kwilu) of the DRC. They did not report any specimens from the more
central areas or from the northwestern part of the country. We were able to trace specimens from the
latter area and from almost all of the northern half of the country, as well as from Rwanda and Burundi,
but none from the south-central provinces (Kasaï-Central, Kasaï Oriental, Lualaba). The distribution
map given by Happold (2013an: 693) doesn’t include any of the additional northern specimens and,
therefore, separates the eastern distribution patches from the western ones. Our map suggests that both
are connected and that only the border zone between the DRC and Angola might not be an optimal
habitat for M. bocagii.
Myotis tricolor (Temminck, 1832)
Fig. 36A–B
Vespertilio tricolor Temminck, 1832: 106.
* Myotis tricolor (Temminck, 1832).
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VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
Fig. 35. Distribution maps. A–B. Kerivoula phalaena Thomas, 1912. C–D. Kerivoula smithii Thomas,
1880. E–F. Myotis bocagii (Peters, 1870). A, C, E. Distribution in the CRB area. B, D, F. Pan-African
distribution.
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In general, M. tricolor primarily occurs in eastern and southern Africa, from Ethiopia to the RSA, only
touching the eastern border of CRB (Bernard 2013: 707). The record from a cave near Thysville (RMCA
16450; Hayman et al. 1966: 52) is, therefore, already far to the west, but not impossible as Koopman
(1989: 7) reported a specimen from Liberia, and ACR (2016: 1207) reported another from Nigeria.
Myotis welwitschii (Gray, 1866)
Fig. 36C–D
Scotophilus welwitschii Gray, 1866: 211.
* Myotis welwitschi venustus (Matschie, 1899).
The validity of the subspecies venustus was already doubted prior to Hayman et al. (1966), e.g., by
Kershaw (1922: 181) and Frechkop (1954: 19), and this view was later conirmed by Kock (1967: 324).
In the DRC, Hayman et al. (1966: 52, map 71) reported this species from Nord-Kivu Province in the
east and from the southeastern provinces of Haut-Lomami, Haut-Katanga and Lualaba. More recently,
it has been found in Rwanda and Burundi. Happold (2013ao: 709) gives a patchy distribution in eastern
and southern Africa, from Ethiopia to the RSA, with an extension into the southern part of the DRC and
Angola. She also plotted two separate localities in Guinea and Cameroon.
Subfamily Scotophilinae Gray, 1866
Genus Scotophilus Leach, 1821
Scotophilus dinganii (A. Smith, 1833)
Fig. 36E–F
Vespertilio Dinganii A. Smith, 1833: 59.
* Scotophilus nigrita (Schreber, 1775): 58.
* Scotophilus nigrita herero Thomas, 1906: 174.
As indicated below, S. nigrita was erroneously used to identify the medium-sized African Scotophilus
species. Robbins (1978: 212) showed that the correct name for this species should be S. dinganii.
Hayman et al. (1966: 57, map 83) reported “S. nigrita” from over almost the entire DRC and from
northern Rwanda, with the exception of the northwestern part of the DRC and the most central part.
Here, we include additional specimens from the northwestern, southwestern and southeastern parts of
the country as well as from Rwanda and Burundi. This still leaves the central part of the DRC unoccupied
by this species. The only central DRC record is a specimen from Boende (Tshuapa Province), but this is
a juvenile specimen, which is probably too young to be identiied with certainty.
Furthermore, we removed a number of localities from the northeastern part of the DRC as the underlying
specimens belonged to the next species.
Happold (2013am: 674) based her distribution map on data from Robbins et al. (1985: 63), which resulted
in a rather detailed map with a lot of inger-like extrusions and areas which almost touch one another.
We believe that the species has a wider distribution, only lacking in eastern Somalia, southern Namibia
and most of the RSA and probably also from the west and central African rainforest as mentioned
by Happold. However, we need to point out that some of the east African records might need to be
reassigned to a separate species, e.g., S. colias as was tentatively suggested by Vallo et al. (2011: 350).
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Fig. 36. Distribution maps. A–B. Myotis tricolor (Temminck, 1832). C–D. Myotis welwitschii (Gray,
1866). E–F. Scotophilus dinganii (A. Smith, 1833). A, C, E. Distribution in the CRB area. B, D, F. PanAfrican distribution.
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Scotophilus leucogaster (Cretzschmar, 1826)
Fig. 37A–B
Nycticejus leucogaster Cretzschmar, 1826: 71.
* Scotophilus leucogaster (Cretzschmar, 1826).
Hayman et al. (1966: 56–57, map 82) mention one locality in the DRC (Faradje, Haut-Uélé Province),
and refer to Koopman (1965: 17) who indicated that Pachyotus altilis (the name under which Allen 1917:
447 reported these specimens) is a synonym of S. l. leucogaster. Allen (1917) and Hayman et al. (1966)
report on two specimens, whereas the AMNH catalog mentions three. The third specimen (AMNH
49283, a foetal specimen) has the same original number (1865) as AMNH 49015 (an adult female).
Only one additional specimen has been reported since: MHNG 1896.016 from Mweya, Burundi.
The map presented by Van Cakenberghe & Happold (2013i: 677) shows S. leucogaster to be present in a
number of areas south of the Sahara: Senegal/Gambia; Guinea/Sierra Leone; Burkina Faso/northern Ghana
to southwestern Chad; central Sudan; central Ethiopia; northeastern DRC; southwestern Zambia/northern
Namibia/northern Botswana/western Zimbabwe. They also plotted individual records from Mauritania,
Côte d’Ivoire, Central African Republic, Uganda, Kenya, western Angola and Namibia, and Mozambique.
The SDM map suggests a sub-Saharan distribution, with a northward extention along the River Nile. The
species probably will not occur in eastern Ethiopia, Eritrea or Somalia, in the central African rain forest
or in southern Africa, except for the northeastern tip of the RSA and southern Mozambique.
Scotophilus nigrita (Schreber, 1774)
Fig. 37C–D
Vespertilio Nigrita Schreber, 1774: 171.
* Scotophilus gigas Dobson, 1875: 122.
Until Robbins (1978: 212), the largest representative of the genus Scotophilus was called S. gigas,
whereas this should be S. nigrita, a name that had been used for the medium-sized African form for
decades. Unfortunately, this led to confusion that left its mark on a lot of subsequent publications, where
it wasn’t always clear whether nigrita was used to designate the large or the medium-sized form.
Hayman et al. (1966: 56, map 81) reported “S. gigas” from one locality: Albertville (= Kalemie) in
Tanganyika Province. The only other specimen from the CRB area was collected in Bujumbura, Burundi.
With the exception of two small areas in Ghana, Happold (2013am: 679) only plotted individual
localities on her distribution map, which shows a major gap between the locality in western Nigeria
(the easternmost for West Africa) and central Sudan. Further to the south, there are records from the
DRC, coastal Kenya and Tanzania, and Mozambique. ACR (2016: 1120) provides some additional
records from Niger, northern Cameroon, Sudan and South Sudan, which more or less bridge the gap
between West Africa and Sudan, and which also may conirm the assignment of the Sudanese specimens
to the nominate subspecies as mentioned by Happold (2013am: 678). The SDM map also suggests that
such a connection might well be possible through the Central African Republic and central Cameroon
or through central Chad and northern Cameroon. Although the species hasn’t been conirmed from the
central African rainforest, its occurrence in the West African forest block makes it plausible.
Scotophilus nux Thomas, 1904
Fig. 37E–F
* Scotophilus nigrita nux Thomas, 1904: 208.
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Fig. 37. Distribution maps. A–B. Scotophilus leucogaster (Cretzschmar, 1826). C–D. Scotophilus
nigrita (Schreber, 1774). E–F. Scotophilus nux Thomas, 1904. A, C, E. Distribution in the CRB area. B,
D, F. Pan-African distribution.
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Thomas (1915: 468) already mentioned “The conspicuous difference between the colour of S. nux and
that of all forms of S. nigrita makes me now think that it should be recognized as a distinct species, and
not merely a subspecies of the older known form.” This view, however, was ignored by most subsequent
authors, until it was resurrected by Robbins (1983: 19) and Robbins et al. (1985: 53, 71).
Hayman et al. (1966: 57) only mentioned specimens from the northeastern part of the DRC (BasUélé, Haut-Uélé and Ituri Provinces). One additional specimen was collected in Kongolo (Tanganyika
Province) in southeastern DRC. Van Cakenberghe & Happold (2013j: 681) indicate that the species
is recorded from the rainforest and the northern and eastern rainforest-savanna mosaics from Sierra
Leone in the west to Cameroon, and from northwestern DRC to western Kenya. We agree with this
distribution, although it occurs further to the south in the DRC than previously accepted. The climatic
data used for the SDM map conirm that the gap between Cameroon and the DRC is probably due to
insuficient sampling, as suggested by Van Cakenberghe & Happold. They also conirm the gap between
southwestern Ghana and southern Nigeria, which might be linked to the “Dahomey Gap”, but further
investigation in that area is needed to conirm this.
Subfamily Vespertilioninae Gray, 1821
Genus Glauconycteris Dobson, 1875
Glauconycteris alboguttata J.A. Allen, 1917
Fig. 38A–B
* Glauconycteris alboguttatus J.A. Allen, 1917: 449.
The only known representatives of this species in the CRB area are the three specimens reported by
Hayman et al. (48, 49, map 60) and one additional specimen from Yalosemba (Mongala Province; CM
86658) reported by Eger & Schlitter (2001: 7) and Thorn et al. (2009: 61). Besides these four specimens,
Happold (2013aa: 564) indicates that the species also occurs in the coastal area of Cameroon. Based on
climatic data, we suspect that the species can occur over the central African forest belt from southern
Nigeria to Rwanda and Burundi.
Glauconycteris argentata (Dobson, 1875)
Fig. 38C–D
Chalinolobus argentatus Dobson, 1875: 385.
* Glauconycteris argentata (Dobson, 1875).
Hayman et al. (1966: 49) mention 72 records for this species, of which at least one (RMCA 16213;
Mukishi, Haut-Lomami Province) was re-identiied as G. variegata (Wim Wendelen, pers. comm.).
They also they mentioned the same museum numbers for the series from Boma and Butembo. These
numbers apply for the Butembo specimens and the correct numbers for the Boma series are 6571 to 6576
and 7250). Their map 61 contains two localities we cannot identify from the specimens they listed. One
of these is most probably Ibembo (Bas-Uélé Province), where they reported G. beatrix (see map 62), and
another one is in Kasaï Province.
Within the CRB area, G. argentata occurs in three separate regions: one along the eastern border of the
DRC (Ituri, Nord- and Sud-Kivu Provinces) and Rwanda and Burundi. A second region is at the mouth
of the Congo River (Kongo Central Province), and a third runs from Mbandaka (Equateur Province) over
Boende and Ikela (Tshuapa Province), Lodja (Sankuru Province) to Kananga (Kasaï-Central Province).
This agrees very well with the map given by Happold (2013ab: 565), although she also cites the “Ibembo”
record listed above. Extralimitally, the species occurs in the coastal areas of Cameroon and Equatorial
Guinea, southwestern Kenya, northeastern Tanzania, northern Malawi and Angola. The SDM map
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Fig. 38. Distribution maps. A–B. Glauconycteris alboguttata J.A. Allen, 1917. C–D. Glauconycteris
argentata (Dobson, 1875). E–F. Glauconycteris beatrix Thomas, 1901. A, C, E. Distribution in the CRB
area. B, D, F. Pan-African distribution.
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suggests that G. argentata might be more widespread over central and eastern Africa, from southern
Nigeria in the west to Kenya and Tanzania in the east, between 5° N and 10° S.
Glauconycteris beatrix Thomas, 1901
Fig. 38E–F
* Glauconycteris beatrix Thomas, 1901: 256.
Glauconycteris beatrix was reported by Hayman et al. (1966: 49, map 62) from two localities in
northern DRC: Ibembo in Bas-Uélé Province and Paulis (= Isiro) in Haut-Uélé Province. Additional
specimens are reported from Irangi (Sud-Kivu), Aketi (Bas-Uélé), Wafanya (Tshuapa) and Kimbinga,
near Kikwit (Kwilu). The specimens from Kimbinga were erroneously reported as Miniopterus minor
by Van Cakenberghe et al. (2000: 308). Happold (2013ac: 566) only mapped the specimens from Irangi,
and except for the coastal areas of Cameroon and Equatorial Guinea, she only plotted individual records
for Côte d’Ivoire, Ghana, southern Nigeria, Gabon and northern Angola. Our distribution map suggests
that G. beatrix could occur in the forest belts of both West and Central Africa, ranging from Sierra Leone
in the west to western Kenya in the east.
Glauconycteris curryae Eger & Schlitter, 2001
Fig. 39A–B
Glauconycteris curryi Eger & Schlitter, 2001: 2.
This species is known from only seven specimens from ive rainforest localities in Cameroon and one in
the DRC (Yalosemba, Mongala Province) (Eger 2013: 568).
Glauconycteris humeralis J.A. Allen, 1917
Fig. 39C–D
* Glauconycteris humeralis J.A. Allen, 1917: 448.
Glauconycteris humeralis was considered to be a valid species by Hayman et al. (1966: 49), but some
later authors (e.g., Koopman 1971; Eger & Schlitter 2001; Monadjem et al. 2010b) considered it to be a
subspecies of G. beatrix. Heller et al. (1994: 6) state that humeralis could merely represent the spotted
variety of beatrix, if it wasn’t for the different form of the baculum in the two species, as was already
illustrated by Hill & Harrison (1987: Fig. 19).
Hayman et al. (1966: 49, map 63) mention six specimens from three localities, of which the
BMNH specimen from Irangi could not be traced. However, the species has been conirmed from
that locality by material in the collection of the SMF. All of these localities were in the northeastern
part of the DRC. Other specimens were collected near Epulu (Ituri Province, northeastern DRC) and
Yalosemba (Mongala, northwestern DRC) and in Kananga (Kasaï-Central Province, southern DRC).
The latter specimen was apparently overlooked by Happold (2013ad: 571), as she only plotted the
northern specimens. When the Kananga specimen is included, the distribution of the species covers the
Congolese rain forest from eastern Cameroon over northern Congo and reaching into the rainforestsavanna mosaic in Uganda and western Kenya (see also Happold 2013ad: 571).
Glauconycteris poensis (Gray, 1842)
Fig. 39E–F
Kerivoula poensis Gray, 1842: 258.
* Glauconycteris poensis (Gray, 1842).
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Fig. 39. Distribution maps. A–B. Glauconycteris curryae Eger & Schlitter, 2001. C–D. Glauconycteris
humeralis J.A. Allen, 1917. E–F. Glauconycteris poensis (Gray, 1842). A, C, E. Distribution in the CRB
area. B, D, F. Pan-African distribution.
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In the DRC, G. poensis was collected at four localities in the northern half of the country, two in the
northwestern part that were already reported by Hayman et al. (1966: 49, map 64), and two from the
northeastern part. The one from Irangi (Nord-Kivu Province) is a relatively recent one, but the one
from Faradje (Haut-Uélé) was already collected by Lang and Chapin. Allen (1917) doesn’t report any
G. poensis specimens in their collection, and the only Glauconycteris from Faradje that he mentions
is G. papilio (= G. variegata). The original entry in the AMNH catalog mentions 15 G. papilio
specimens from Niangara, Faradje and Aba, which were changed into Chalinolobus variegata (49056–
9), Glauconycteris variegata variegata (49060–4), Chalinolobus (Glauconycteris) variegata papilio
(49065) and Glauconycteris variegata variegata (49066–70). The online catalog, however, mentions
the entire series, with the exception of 49066, as Glauconycteris variegata variegata. That specimen is
identiied as G. poensis. Happold (2013ae: 575) overlooked this record when drawing the map for this
species. Her map indicates that G. poensis is primarily a West African species, with some outliers in the
DRC. There is also a doubtful record from Marangu (Tanzania) of which J. Kingdon (in Grimshaw et al.
1995) suggests that it was misidentiied, and most likely belongs to G. variegata. The online catalog still
mentions the underlying specimens (219772–4) as G. poensis. The SDM map indicates that this locality
is situated far to the east, but it is not impossible that poensis could occur over there.
Glauconycteris superba Hayman, 1939
Fig. 40A–B
Glauconycteris superba Hayman, 1939: 219.
* Glauconycteris superba superba Hayman, 1939.
At the time of Hayman et al. (1966) only two specimens of this remarkably coloured bat were known:
the holotype of the species from the DRC and a specimen from Ghana, which was considered to belong
to a separate subspecies, G. superba sheila Hayman, 1947. In 1972, a third specimen was collected in
Côte d’Ivoire. In the next 40 years, not a single specimen of this species was found, but in 2012 two
animals were collected: one in the Bangangai Game Reserve, South Sudan (Reeder et al. 2013), and
one from Mbiye Island, near Kisangani, Tshopo Province, DRC (Gembu Tungaluna et al. 2013: 18). In
2015, another specimen was collected on Mbiye Island. This is the irst time the species was collected
at the same locality, suggesting that G. superba is not as rare as was assumed during the irst 40 years
after its initial discovery.
As if to conirm the previous statement, after the manuscript for this paper was submitted, Ing et al.
(2016) and Hassanin et al. (2017) reported on ten additional specimens from Mbiye island and the
Yoko Forest Reserve (00°17′ N, 25°17′ E). Hassanin et al. (2017) also described a new species of
Glauconycteris, which is not covered in the current paper.
Based on a number of morphological characters, Reeder et al. (2013) created a separate genus for
superba: Niumbaha. These characters include the considerably larger skull; more striking body patterns;
not excessively foreshortened (i.e., longer) rostrum; more robust muzzle; unreduced relative canine size;
unelongated wing tips (second phalanxes); more robust, subquadrangular ears, with a larger free lobe
at the inner margin; a more strongly curved tragus; position of the nostrils. Although these are quite
important differences, we remain reluctant to follow Reeder et al. (2013) in separating superba from the
genus Glauconycteris, due to the fact that the morphological characters were probably only examined
on one specimen and some of them might be the result of the large size of superba. The rejection of
Niumbaha as a separate genus is conirmed by the DNA analyses by Hassanin et al. (2017).
Given the fact that G. superba was rarely collected in the past, it is very dificult to assess its general
distribution. The SDM map suggests that it may occur in the forest zones of West and Central Africa.
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Fig. 40. Distribution maps. A–B. Glauconycteris superba Hayman, 1939. C–D. Glauconycteris
variegata (Tomes, 1861). E–F. Hypsugo anchietae (Seabra, 1900). A, C, E. Distribution in the CRB
area. B, D, F. Pan-African distribution.
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Glauconycteris variegata (Tomes, 1861)
Fig. 40C–D
Scotophilus variegatus Tomes,1861: 31, 36.
* Glauconycteris variegata papilio (Thomas, 1905): 77.
Although most recent authors consider papilio to be a synonym of G. variegata (e.g., Simmons 2005) or
even G. v. variegata (e.g., Lanza et al. 2015), Monadjem et al. (2010b: 417) – referring to a statement
made by Hayman (1963: 107) on the possible speciic status of papilio – suggest that a detailed study of
this species group is necessary to resolve this matter.
For the DRC, Hayman et al. (1966: 50, map 66) indicate that G. variegata primarily occurs in the
northeastern part of the country (Haut-Uélé and Nord-Kivu Provinces) and in the southeast (HautLomami and Haut-Katanga Provinces). From the western part of the country, they report it from Boma
(Kongo Central Province) and Kwamouth (Mai-Ndombe Province). The second record they mapped
from Kongo Central Province could not be traced, but they also did not map Mbala in Tanganyika
Province. We could only add two additional localities: one on the northwestern bank of Lake Tanganyika
(Sud-Kivu Province) and at Boteka (Equateur Province). The map given by Happold (2013af: 577)
shows a very patchy distribution area stretching from Senegal to Somalia and southwards to northeastern
RSA. Based on specimen data from ACR (2016: 1230) and climatic data, our distribution map ills in
most of the gaps between the patches.
Genus Hypsugo Kolenati, 1856
Hypsugo anchietae (Seabra, 1900)
Fig. 40E–F
Vesperugo Anchieta Seabra, 1900: 26.
Two specimens from Lubenge (Tanganyika Province) were re-identiied by the irst author as belonging
to this species. Kearney (2013b: 611) indicates that the species occurs from southwestern to eastern
Angola, and through southeastern DRC, western Zambia, Zimbabwe to eastern RSA and Swaziland,
where it is found in riverine, coastal and scrub forests as well as bushveld biotopes.
Hypsugo crassulus (Thomas, 1904)
Fig. 41A–B
* Pipistrellus crassulus Thomas, 1904: 206.
As indicated by Fahr (2013q: 617) crassulus has been moved into various (sub)genera or “collective
groups” over the past few years: Pipistrellus, P. (Vansonia), “Attalepharca”, Hypsugo and Nycticeinops.
The currently most accepted view is that it belongs to the genus Hypsugo.
Hayman et al. (1966: 52, map 72) reported it from two localities: Bokuma (Equateur Province) and
Mulolo-Kisanga (Maniema Province), and more recently it was also found at Irangi (Sud-Kivu Province).
The distribution map provided by Fahr (2013q: 618) only shows a very restricted red area covering the
eastern parts of the Kivu Provinces reaching towards Maniema Province. Furthermore, there are single
records from Guinea, in West Africa, over Côte d’Ivoire, southern Cameroon, northwestern DRC (the
Bokuma record), northern Angola, southern South Sudan, southern Uganda and southwestern Kenya.
Based on climatic data, the SDM map covers this area, but also suggests it might occur in Equatorial
Guinea, Gabon, southern Congo, and southwestern and southeastern DRC.
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Fig. 41. Distribution maps. A–B. Hypsugo crassulus (Thomas, 1904). C–D. Hypsugo cf. eisentrauti
(Hill, 1968). E–F. Hypsugo musciculus (Thomas, 1913). A, C, E. Distribution in the CRB area. B, D, F.
Pan-African distribution.
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Hypsugo cf. eisentrauti (Hill, 1968)
Fig. 41C–D
Pipistrellus eisentrauti Hill, 1968: 45.
“Hypsugo eisentrauti” was collected at two localities in the DRC: Lukolela (Mai-Ndombe Province)
and Kabobo Forest (Sud-Kivu Province) and in one locality in Rwanda: Nyungwe Forest Reserve.
However, as already indicated by Van Cakenberghe & Happold (2013b: 622), the specimens from
Lukolela do not agree with the type specimens of eisentrauti from Cameroon. And this is also the case
for the animal from Rwanda. Huhndorf & Kaleme (2008: 26) and Kerbis Peterhans et al. (2013: 189)
assigned the Kabobo Forest specimen to “H. cf. eisentrauti”. Tentatively, we assign this name to all of
the DRC and Rwandan specimens.
Hypsugo musciculus (Thomas, 1913)
Fig. 41E–F
* Pipistrellus musciculus Thomas, 1913: 316.
Similar to H. crassulus, H. musciculus has been assigned to various supra-speciic units: Pipistrellus,
P. (Hypsugo), “Attalepharca” and Hypsugo (see Van Cakenberghe & Happold 2013e: 637). Here, we
follow Simmons (2005: 491) in assigning it to the genus Hypsugo.
Hayman et al. (1966: map 74) have two localities plotted on their map: Bokuma in Equateur Province
and Avakubi in Tshopo Province. For the latter, however, they refer (p. 53) to Koopman (1965: 13) who
indicated that this specimen agreed better with the original description of P. nanulus than with that of
P. musciculus. In the Paris museum, there is one other DRC specimen (from Bata, Kwilu Province) that
had been identiied as musciculus.
Hypsugo musciculus is primarily a West African species occurring from Sierra Leone in the west to
Cameroon and Gabon in the east, with a few records extending into the DRC.
Genus Laephotis Thomas, 1901
Laephotis angolensis Monard, 1935
Fig. 42A–B
Laephotis angolensis Monard, 1935: 45.
* Laephotis wintoni angolensis (Monard, 1935).
At the time Hayman et al. (1966) was written, the genus Laephotis was considered to be monotypic
(Hill 1974: 1), with the nominate subspecies restricted to Kenya and L. w. angolensis occurring in
Angola, Zambia, Botswana and the southern part of the DRC. However, based on cranial characteristics,
Peterson (1971a: 888) found the two to be speciically different.
Laephotis angolensis was reported by Hayman et al. (1966: 50, map 67) with 13 specimens from two
localities in Haut-Katanga, and no further localities have been added since. Of these 11 were found to
represent L. botswanae (Kearney & Seamark 2005: 73). Extralimitally, the species is further known
from Angola, Zimbabwe and Zambia.
Laephotis botswanae Setzer, 1971
Fig. 42C–D
Laephotis botswanae Setzer, 1971: 260.
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Fig. 42. Distribution maps. A–B. Laephotis angolensis Monard, 1935. C–D. Laephotis botswanae
Setzer, 1971. E–F. Mimetillus moloneyi (Thomas, 1891). A, C, E. Distribution in the CRB area. B, D,
F. Pan-African distribution.
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This species was reported from two localities near Lubumbashi (Haut-Katanga Province) in the extreme
southeastern part of the DRC. The distribution map provided by Kearney (2013a: 580) indicates that the
species occurs in a number of scattered areas in southern Africa: southeastern DRC and northwestern
Zambia, central and southern Zimbabwe reaching into northern RSA, northwestern Botswana and
northeastern Namibia, southern Malawi, and in some individual localities in Angola, southeastern
RSA and northern Malawi. Kearney (2013a: 580), however, also pointed out that the relationship
between angolensis and botswanae is obscure, because of the scarcity of the former, but she also refers
to Setzer (1971: 262), who indicated that the two species might possibly occur sympatrically.
Genus Mimetillus Thomas, 1904
Mimetillus moloneyi (Thomas, 1891)
Fig. 42E–F
Vesperugo (Vesperus) Moloneyi Thomas, 1891: 528.
* Mimetillus moloneyi (Thomas, 1891).
Hayman et al. (1966: 50, map 68) mention this species from a number of localities in the northwestern
part of the DRC (Bas- and Haut-Uélé, Ituri, Tshopo Provinces), but also from the southwestern Landana
(“Lower Congo”). This last locality, however, is in Cabinda (Angola). The southern occurrence of
M. moloneyi in the DRC is conirmed by records from Luluabourg (= Kananga) in Central Kasaï Province
and Mosenge in Kwilu Province. These records were not included by Fahr (2013o: 587), who, therefore,
plotted only a narrow band in the northern part of the DRC and individual localities in the border area
between DRC, Uganda and Rwanda in the east, the border between the DRC and Zambia in the south
and in Cabinda and the mouth of the Congo River in the west. He also stated that it is conspicuously
absent from most of the central Congo Basin. Our additional records, however, suggest that it might be
present in the fringes of the rain forest and other forest types and savannas reaching to the Kenyan and
Tanzanian coast in the east and Sierra Leone and Guinea in the west.
Mimetillus thomasi Hinton, 1920
Fig. 43A–B
Mimetillus thomasi Hinton, 1920: 240.
Although Fahr (2013o: 586) considers thomasi to be a subspecies of M. moloneyi, we follow Cotterill
(2001b: 215), who proposed that the two are distinct evolutionary species, a view that was followed
by Monadjem et al. (2010a: 390, 2010b: 444). The distribution map presented by Monadjem et al.
(2010b: 444) indicates that M. thomasi occurs in the northern part of southern Africa, from Angola over
southern DRC, Zambia and Zimbabwe to western and northern Mozambique. The DRC specimen from
Elisabethville (= Lubumbashi, Haut-Katanga Province) reported here its within this range.
Genus Neoromicia Roberts, 1926
Neoromicia brunnea (Thomas, 1880)
Fig. 43C–D
Vesperugo (Vesperus) brunneus Thomas, 1880: 165.
One specimen belonging to this species was collected at Lukolela (Mai-Ndombe Province, northwestern
DRC). Fahr (2013p: 613) indicates that this species has a West African distribution, covering an area
along the coast of Cameroon and Equatorial Guinea, and individual localities in Sierra Leone, Liberia,
Côte d’Ivoire and Ghana. ACR (2016: 1237), furthermore, mentions specimens from Congo, Gabon and
Nigeria. The Lukolela specimen is the easternmost record of this species.
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Fig. 43. Distribution maps. A–B. Mimetillus thomasi Hinton, 1920. C–D. Neoromicia brunnea (Thomas,
1880). E–F. Neoromicia capensis (A. Smith, 1829). A, C, E. Distribution in the CRB area. B, D, F. PanAfrican distribution.
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Neoromicia capensis (A. Smith, 1829)
Fig. 43E–F
Vespertilio Capensis A. Smith, 1829: 435.
* Eptesicus capensis (A. Smith, 1829).
* Eptesicus garambae” J.A. Allen, 1917: 445.
* Eptesicus minutus” (Temminck, 1835): 209.
* Eptesicus grandidieri” (Dobson, 1876): 500.
Hayman et al. (1966: 46) already pointed to the confused status of the nomenclature of the small, darkwinged African members of the genus Eptesicus, where they indicated that E. capensis might include
the following synonyms: garambae, minutus, grandidieri and possibly pusillus. Although the situation
at the species level has been resolved, the supraspeciic (generic) assignment of the African taxa once
considered to belong to either Eptesicus or Pipistrellus still remains uncertain. Currently, these taxa
are assigned to Eptesicus, Pipistrellus, Neoromicia and Hypsugo, of which the former does not have
representatives in the CRB area. One of the major reasons for this confusion is that there are hardly
any morphological characters which easily distinguish these genera (see Van Cakenberghe & Happold
2013a: 600). Based on bacular morphology, Hill & Harrison (1987: 261) suggested that capensis should
be transferred from the genus Eptesicus to the genus Pipistrellus and more speciically to the subgenus
Neoromicia. Based on dental characters, Menu (1987: 33) placed capensis in the genus Nycterikaupius.
Karyological data led Volleth et al. (2001: 25) to elevate the subgenus Neoromicia to generic rank.
Distributionwise, Hayman et al. (1966: 46) provided data for four taxa: capensis, “garambae”,
“grandidieri” and “minutus”, which were plotted on maps 54, 55, 56 and 57. The specimens identiied
as belonging to the latter two taxa were only reported from the area of the Garamba National Park
(Haut Uélé Province). Specimens belonging to capensis were reported from two additional localities in
the northern part of the DRC and also from a number of localities in the southern former provinces of
Katanga and Kasai, as well as from Léopoldville (= Kinshasa). The SDM map indicates that the core
area of distribution for N. capensis is southern Africa, south of the DRC, but also that specimens have
been reported from as far north as Guinea, Ghana, Togo and Cameroon in western Africa and South
Sudan and Ethiopia in the east. The map provided by Kearney (2013c: 615) shows a gap in NE (sic)
Angola / SW DRC, and one from the central DRC over Tanzania to Mozambique, which might relect
insuficient sampling. However, she also left a gap between the DRC specimens and those from Cameroon,
suggesting that the West African populations might be isolated from the remainder of the distribution
area and represent another taxon. This might either be a different species or a form of capensis, which
has not yet been named, as Kearney doesn’t mention any subspecies from that part of the continent.
Neoromicia guineensis (Bocage, 1889)
Fig. 44A–B
Vesperus guineensis Bocage, 1889: 6.
The two specimens assigned to this species were collected by Herbert Lang and James Paul Chapin,
and were reported by Allen (1917: 445) as belonging to Eptesicus minutus minutus (Temminck,
1835). Hayman et al. (1966: 47) refer to these specimens as “Eptesicus minutus”, indicating that the
name minutus was used in a tentative sense. However Rosevear (1965: 255) pointed out that minutus
Temminck, 1835 is a synonym of Eptesicus capensis. Koopman (1965: 15) tentatively identiied these
two specimens as Eptesicus pusillus, but Koopman (1975: 407) furthermore pointed out that pusillus
was based on two sources: Leconte (1858) and Noack (1889). The former was found to be unidentiiable
and the latter to be a synonym of Pipistrellus nanus. Koopman’s igure 39 indicates that the extralimital
(to Sudan) specimen from northeastern DRC is considered to belong to Eptesicus guineensis.
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Fig. 44. Distribution maps. A–B. Neoromicia guineensis (Bocage, 1889). C–D. Neoromicia nana
(Peters, 1852). E–F. Neoromicia rendalli (Thomas, 1889). A, C, E. Distribution in the CRB area. B, D,
F. Pan-African distribution.
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Van Cakenberghe & Happold (2013d: 625) indicate that guineensis primarily occurs disjunctly north of
the equator in a band from Senegal, Guinea, northern Côte d’Ivoire, Ghana, Burkina Faso, northern Togo
and Benin, and in a second band on the border between Nigeria and Cameroon, reaching to southern
Chad. Furthermore, it occurs in a few smaller areas in central Sudan, the border between South Sudan
and Uganda and in northwestern DRC. There are also some isolated records from Liberia, Nigeria, and
western and eastern Ethiopia. The identiications of the specimens from Congo and southern DRC were
found to be erroneous.
Neoromicia nana (Peters, 1852)
Fig. 44C–D
Vespertilio nanus Peters, 1852: 63.
* Pipistrellus culex Thomas, 1911: 458.
* Pipistrellus nanus nanus Peters, 1852.
Although they followed Koopman (1965) in many of his other conclusions, Hayman et al. (1966: 52)
did not follow Koopman’s view (1965: 12) on culex, which he tentatively considered a subspecies of
P. stamplii (Jentink, 1888), but of which he also stated that it possibly only represents a small-sized
population of P. nanus. Rosevear (1965: 268), Koopman (1966: 160) and Kock (1969: 170) considered
culex to be a synonym of P. nanus. Over the past few decades Vespertilio nanus has changed genus a
number of times (Happold 2013ah: 639): Pipistrellus, Neoromicia, Hypsugo. The current assignment to
Neoromicia was based on karyotype data (Volleth et al. 2001: 44; Kearney et al. 2002: 55).
Hayman et al. (1966: 52, 53–55, maps 73, 76) reported N. nana from all over the CRB area, and some
of the gaps have been illed in with additional specimens. The only exception seems to be the central
part of the DRC (the area around the Salonga National Park), but that might possibly be attributed to
the impenetrability of the region rather than the real absence of the species. The SDM map looks very
similar to the one presented by Happold (2013ah: 640), although we show a number of hatched areas
where the climatic circumstances might not be optimal for the species, e.g., the central forest area in
the DRC (but see above), the border area between the DRC and Angola, eastern Angola and western
Tanzania.
Neoromicia rendalli (Thomas, 1889)
Fig. 44E–F
Vesperugo (Vesperus) Rendalli Thomas, 1889: 362.
* Eptesicus rendalli (Thomas, 1889).
Hayman et al. (1966: 47, map 58) mention primarily specimens from the Garamba National Park area
(Haut Uélé Province), and some single records from the provinces of Ituri, Tanganyika, Kasaï Central
and Tshuapa. They also mention a specimen from Butembo (Nord-Kivu Province; RMCA 26585), but
that specimen should be an Epomophorus anurus [= E. labiatus] (Wim Wendelen, pers. comm.). The
patchy distribution map given by Van Cakenberghe & Happold (2013g: 646) shows three red zones
in the CBR area: one in the northeastern part of the DRC reaching northern Rwanda, a second in the
northwestern part on the boundary between the DRC, Congo and the Central African Republic and a
third in the province of Haut-Lomami towards Kasaï. Based on climatic data, our general map suggests
a more continuous sub-Saharan distribution from about 15° N to 15° S (following the borders between
Angola / Namibia and Zambia / Botswana + Zimbabwe) and further south to southern Mozambique and
Swaziland, with possibly the exception of the central Congolese rain forest.
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Neoromicia somalica (Thomas, 1901)
Fig. 45A–B
Vespertilio minutus somalicus Thomas, 1901: 32.
Representatives of this species were collected at four localities, two in western DRC (Lukolela, MaiNdombe Province and Mbwambala, Kwilu Province), one in northeastern DRC (Mboga, Ituri Province),
and one in Rwanda (Karama). The SMF has furthermore a specimen from a non-disclosed locality in
the DRC. Van Cakenberghe & Happold (2013h: 654) show that the distribution area of somalica forms a
narrow band from southern Senegal over Guinea, Sierra Leone, northern Côte d’Ivoire, southern Burkina
Faso, Togo, Benin, Nigeria, northern Cameroon, Central African Republic, South Sudan, splitting up
in a branch towards Ethiopia and northern Somalia and a second branch to Uganda, eastern Rwanda,
northern Tanzania, southern Kenya and reaching into southern Somalia. There are also scattered records
from Congo and the DRC and some uncertain records from southern Tanzania, Mozambique, Zambia,
Zimbabwe, Botswana, Angola and RSA. As indicated by Van Cakenberghe & Happold, some of these
latter records might represent the N. zuluensis.
Neoromicia tenuipinnis (Peters, 1872)
Fig. 45C–D
Vesperus tenuipinnis Peters, 1872: 236.
* Eptesicus tenuipinnis (Peters, 1872).
* Eptesicus tenuipinnis ater (J.A. Allen, 1917): 443.
Hayman et al. (1966: 47, 48, map 59) showed that tenuipinnis was frequently captured in the
southwestern part of the DRC (from Banana in Kongo Central Province to the area of Mbandaka in
Equateur Province) and in the northeastern part (provinces of Bas-Uélé, Haut-Uélé, Ituri and NordKivu). In the southeast they reach Kabongo (northern Haut-Lomami Province). The species seems to
be missing in the southeastern provinces of Haut-Katanga and Tanganyika. More recently, specimens
were collected in the northwestern part of the country (Tandala, Sud-Ubangi Province and Yalosemba,
Mongala Province), and in northern Rwanda and western Burundi. Fahr (2013r: 656) reported that
tenuipinnis is recorded from the lowland rainforest, swamp forest and coastal forests as well as from
montane forest, mangroves, forest-savanna mosaic, Isoberlinia woodland, Acacia-Commiphora
bushland and thicket and miombo woodland. Within the rainforest, it seems to prefer drier types of
forest along the periphery.
Neoromicia zuluensis (Roberts, 1924)
Fig. 45E–F
Eptesicus zuluensis Roberts, 1924: 60.
As mentioned above, confusion with N. somalica might have led to some mis-identiications. However,
one specimen in the collection of the Zürich Museum from Mimuli (Rwanda) was identiied as belonging
to this species. The distribution map presented by Happold et al. (2013: 658) indicates that the species
seems to cover two major distribution areas, one covering the northeastern part of Uganda and the western
part of Kenya, reaching into central Ethiopia, and a second reaching from the southeastern DRC over
major parts of Zambia and Zimbabwe to northeastern and eastern Botswana and the northeastern
RSA. They also indicate its presence at separate localities in eastern Kenya, Angola, Namibia, Malawi
and central RSA. ACR (2016: 1264) furthermore mentions specimens from Mozambique, South Sudan
and Swaziland.
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Fig. 45. Distribution maps. A–B. Neoromicia somalica (Thomas, 1901). C–D. Neoromicia tenuipinnis
(Peters, 1872). E–F. Neoromicia zuluensis (Roberts, 1924). A, C, E. Distribution in the CRB area. B, D,
F. Pan-African distribution.
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Genus Nycticeinops Hill & Harrison, 1987
Nycticeinops schlieffenii (Peters, 1860)
Fig. 46A–B
Nycticejus Schlieffenii Peters, 1860: 224.
* Nycticeius (Scoteinus) schlieffeni Peters, 1860.
* Nycticeius (Scoteinus) schlieffeni albiventer Thomas & Wroughton, 1908: 540.
Based on the shape of the baculum, Hill & Harrison (1987: 254) created a new genus (Nycticeinops),
which contains schlieffenii as the only species. Happold (2013ag: 595) indicated that four subspecies
were recognized by Hayman & Hill (1971: 36), but these are of dubious validity.
Hayman et al. (1966: 55–56, map 78) report schlieffenii from the northeastern part of the DRC, where
they plotted three localities. Of these, Niangara needs to be removed as they stated on page 56 that
Koopman (1965: 17) reassigned this specimen to Scotoecus hirundo hindei. Furthermore, Hayman et al.
(1966) mention specimens from the southeastern part of the DRC (Tanganyika, Haut-Katanga and HautLomami Provinces). The locality on Lake Tanganyika they mention is probably (Qua) Mpala, the type
locality of Scotophilus minimus Noack, 1887, but this specimen could not be traced and is, therefore,
not included here. More to the west, they also mentioned a specimen from Luluabourg. In the collection
of the Tervuren museum, there is also an additional specimen from “Bas Congo” (RMCA 65b), which
probably was overlooked by Hayman et al. (1966), as they report a Myotis bocagii from the same
locality (RMCA 65). Recently, an additional specimen was collected by the team of the University of
Kisangani at Aketi (Bas-Uélé Province).
The map presented by Happold (2013ag: 595) shows two major distribution areas: the Sudanian
woodlands in West Africa (from the Mauritanian-Senegalese border in the west to Nigeria and northern
Cameroon in the east). The second area covers a major part of eastern Africa (from the coastal area in
Sudan and Eritrea, splitting in two narrow stretches in Ethiopia, combining again to cover central Kenya,
passing through central Tanzania to the southeastern DRC, Zambia, extending to the border between
Angola and Namibia, Malawi, northern Zimbabwe, central and eastern Mozambique to the extreme
northeastern RSA), also including a separate, small area in the northeastern DRC. The specimens
reported here conirm the species’ presence in the two DRC regions, but also suggest that the species
extends more to the west.
Genus Pipistrellus Kaup, 1829
Pipistrellus grandidieri (Dobson, 1876)
Fig 46C–D
Vesperugo (Vesperus) grandidieri Dobson, 1876: 500.
* Eptesicus grandidieri (Dobson, 1876).
Although Hayman et al. (1966: 46) report six “Eptesicus grandidieri” specimens from the DRC, none
of these actually belong to that species, but rather to Neoromicia capensis. However, two additional
specimens from Resha (Burundi; SMF 57586 and 57587) were found to belong to grandidieri, so the
species does occur in the CRB area.
Thorn et al. (2007) re-evaluated the known material of grandidieri and concluded that this form should
be assigned to a separate subgenus (Afropipistrellus) within the genus Pipistrellus. Van Cakenberghe &
Happold (2013c: 623) present an extremely patchy distribution range, which covers a number of isolated
localities ranging from the Nigerian/Cameroonian border to southern Somalia, and southwards to central
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Fig. 46. Distribution maps. A–B. Nycticeinops schlieffenii (Peters, 1860). C–D. Pipistrellus grandidieri
(Dobson, 1876). E–F. Pipistrellus hesperidus (Temminck, 1840). A, C, E. Distribution in the CRB area.
B, D, F. Pan-African distribution.
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VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
Angola and southern Mozambique. Here, we extend the distribution to Guinea and Liberia in West
Africa, where “Pipistrellus cf. grandidieri” was reported by Decher et al. (2016: 273).
Pipistrellus hesperidus (Temminck, 1840)
Fig. 46E–F
Vespertilio hesperida Temminck, 1840: 211.
This species used to be included in Pipistrellus kuhlii, but as found by Göpfert et al. (1995: 68) and
Volleth et al. (2001: 28), the sub-Saharan specimens have a different chromosome number than the
specimens from Europe and northern Africa. For these sub-Saharan populations, Kock (2001: 277) reinstated the name hesperidus.
Specimens belonging to this species were collected at three localities in the CRB area: two in Rwanda
(Mutura and Kitabi) and one in Burundi (Nyamugari Hill). Kearney (2013d: 630) indicates that
P. hesperidus is distributed in a narrow band in eastern Africa, reaching from southwestern Eritrea,
over western Ethiopia, eastern South Sudan to northern Uganda, where it splits into two branches, one
along the Kenyan-Tanzanian border reaching the coast, and a second along the great lakes to eastern
Zimbabwe and the eastern RSA, reaching just north of Port Elizabeth. Additionally, she marked the
presence of the species in the Mount Cameroon area and in Djibouti, with some further individual
localities in northeastern Somalia, western Zambia, Angola and southern RSA. Furthermore, she refers
to Jakob Fahr, who indicated that some records from West Africa might be misidentiied. The SDM map
conirms the presence of the species in eastern Africa. The records from West Africa might need to be
re-examined.
Pipistrellus nanulus Thomas, 1904
Fig. 47A–B
* Pipistrellus nanulus Thomas, 1904: 198.
Hayman et al. (1966: map 75) plotted one specimen from Avakubi, which is actually the same specimen
they plotted as “Pipistrellus musciculus” (see above). One additional record is from from Mosenge in
Kwilu Province.
The map given by Van Cakenberghe & Happold (2013f: 639) shows that P. nanulus primarily occurs
in the savannas of Sudan and Guinea, the northern rainforest-savanna mosaic in West Africa (from
Guinea to Togo and from southern Nigeria to Gabon, and from north-central Benin to central Nigeria)
and in the eastern rainforest-savanna mosaic in northeastern DRC, Uganda and western Kenya. Based
on additional material mentioned in ACR (2016: 1276), our distribution map covers a more continuous
area from Senegal to western Kenya, with a small gap at the Dahomey Gap in western Africa. The map
also shows that it could occur over almost the entire northern half of the DRC, with the exception of the
central part.
Pipistrellus rueppellii (Fischer, 1826)
Fig. 47C–D
Vespertilio Rüppelii J.B. Fischer, 1826: 109.
* Pipistrellus (Scotozous) rüppelli (Fischer, 1826).
* Pipistrellus (Scotozous) rüppelli fuscipes (Thomas, 1913): 315.
Happold (2013ai: 648) mentions that fuscipes is the subspecies of P. rueppellii occurring from Ethiopia to
northern Angola, which would cover the CRB area. Hayman et al. (1966: 55, map 77) report the species
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from the northeastern part of the DRC (provinces Bas-Uélé, Tshopo, Ituri and Nord-Kivu), as well as
from the southeast (Tanganyika and Haut-Katanga Provinces) and southwest parts (Kongo Central,
Kasaï and Mai-Ndombe Provinces). The two additional records (RMCA 12247 from Mongbwalu and
RMCA 993 from Lukula) were probably overlooked by Hayman et al. (1966), as these specimens were
present in the Tervuren collection at the time.
Happold’s (2013ai: 648) distribution map essentially shows the species following the River Nile
southwards, along the great lakes to northern Zimbabwe and central Mozambique, speading in the
southern part where it should occur over most of Zambia. Additionally, the species occurs in northern
DRC, along the lower Congo River and up the Kasaï River; along the Moroccan-Algerian border
crossing into the latter country; and along the Senegalese-Mauritanian border. Furthermore, there are
some individual records from Egypt, Ethiopia, eastern Tanzania, Botswana, RSA, Angola, Gabon,
Central African Republic, Chad, Nigeria, Libya, Tunisia and Morocco. The species seems to be lacking
from West Africa, although climatic data used for the SDM map suggest that it might occur in that part
of the continent.
Genus Scotoecus Thomas, 1901
Scotoecus albofuscus (Thomas, 1890)
Fig. 47E–F
Scotophilus albofuscus Thomas, 1890: 84.
* Nycticeius (Scotoecus) albofuscus woodi (Thomas, 1917): 280.
Although Hayman et al. (1966: 56) considered Scotoecus to be a subgenus of Nycticeius, its generic
status was already restored by Rosevear (1965: 246) and Koopman (1965: 17), who were followed by
Hill (1974b: 171), Koopman (1993: 226, 1994: 127) and Simmons (2005: 461, 464).
Only one specimen of this species has been reported from the CRB area up until now: RMCA 22412
from Baudouinville (= Moba), Tanganyika Province. It was irst mentioned by Hayman (1957: 44)
and subsequently conirmed by Fain (1959: 161), Hill (1974b: 177), Anciaux de Faveaux (1983: 29),
Monadjem et al. (2010b: 506) and Happold (2013aj: 668).
The distribution map presented by Happold (2013aj: 668) clearly illustrates the rarity of this species in
scientiic collections, as only individual records are plotted, with the exception of small areas in Sierra
Leone and The Gambia. The species occurs from the latter country in West Africa to Northern Cameroon
and then continuing from northwestern Uganda, over Tanzania, southeastern DRC and Mozambique to
the RSA. This leaves a wide gap between Cameroon and Uganda. ACR (2016: 1283) mentions three
specimens from the Central African Republic, which narrow this gap a bit, but it nevertheless remains
wide. We concur with Ansell & Dowsett (1988: 43), who state that “this species is rarely collected,
though it has a wide range from Senegal to southern Mozambique. It is no doubt more widespread in
Malawi than the single recorded locality would indicate”, and believe this is also the case for many other
countries. The SDM map suggests that S. albofuscus potentially can occur along the northern border of
the DRC, where it can bridge the gap between the two areas where it has been found up until now.
Scotoecus hindei Thomas, 1901
Fig. 48A–B
Scotoecus Hindei Thomas, 1901: 264.
* Nycticeius (Scotoecus) hirundo hindei (Thomas, 1901).
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VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
Fig. 47. Distribution maps. A–B. Pipistrellus nanulus Thomas, 1904. C–D. Pipistrellus rueppellii
(Fischer, 1826). E–F. Scotoecus albofuscus (Thomas, 1890). A, C, E. Distribution in the CRB area. B,
D, F. Pan-African distribution.
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The taxonomy of the dark-winged Scotoecus forms is not very clear. Pending a revision of this group,
Happold (2013ak: 669) considers all the dark-winged Scotoecus to belong to one species: S. hirundo.
However, we follow Ansell & Dowsett (1988: 44), Taylor & Van der Merwe (1998: 64), Cotterill (2001c:
219) and Simmons (2005: 464) in recognizing S. hindei as a separate species.
Scotoecus hindei has only been reported twice from the DRC: once from the southeast of the country
(Kapolowe, Haut-Katanga Province) and once from the northeast (Niangara, Haut-Uélé Province). Both
were already mentioned by Hayman et al. (1966: 56).
Since Happold (2013ak: 670) doesn’t recognize hindei as a separate species, our distribution map cannot
be compared with hers. Based on the data in ACR (2016: 1283). The SDM map suggests a primarily
eastern African distribution, from Eritrea in the north to southern Mozambique and northern RSA in
the south. It extends westwards across the northern part of the DRC, the Central African Republic,
Cameroon and Nigeria.
Fig. 48. Distribution maps. Scotoecus hindei Thomas, 1901. A. Distribution in the CRB area. B. PanAfrican distribution.
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Nominal list of bats of the Congo, Rwanda and Burundi (CRB) region
Present study
PTEROPODIFORMI
Hayman et al. (1966)
MEGACHIROPTERA
Pteropidae
Eidolon helvum (Kerr, 1792)
Pteropodidae
Eidolinae
Eidolon helvum (Kerr, 1792)
Rousettinae
Epomophorini
Epomophorus anselli Bergmans & Van Strien, 2004
Epomophorus crypturus Peters, 1852
Epomophorus gambianus (Ogilby, 1835)
Epomophorus anurus Heuglin, 1864
Epomophorus labiatus minor (Dobson, 1880)
Epomophorus crypturus Peters, 1852
Epomophorus labiatus (Temminck, 1837)
Epomophorus minimus Claessen & De Vree, 1991
Epomophorus labiatus minor (Dobson, 1880)
Epomophorus minor Dobson, 1880
Epomophorus wahlbergi haldemani (Halowell, 1846)
Epomophorus wahlbergi (Sundevall, 1846)
Epomops dobsoni (Bocage, 1889)
Epomops dobsonii (Bocage, 1889)
Epomops franqueti franqueti (Tomes, 1860)
Epomops franqueti (Tomes, 1860)
Hypsignathus monstrosus H. Allen, 1861
Hypsignathus monstrosus H. Allen, 1862
Micropteropus intermedius Hayman, 1963
Micropteropus intermedius Hayman, 1963
Micropteropus pusillus (Peters, 1867)
Micropteropus pusillus (Peters, 1868)
Nanonycteris veldkampii (Jentink, 1888)
Myonycterini
Megaloglossus woermanni Pagenstecher, 1885
Megaloglossus woermanni prigoginei Hayman, 1966
Rousettus (Lissonycteris) angolensis (Bocage, 1898)
Megaloglossus woermanni Pagenstecher, 1885
Myonycteris angolensis (Bocage, 1898)
Myonycteris relicta Bergmans, 1980
Myonycteris wroughtoni Andersen, 1908
Myonycteris torquata (Dobson, 1878)
Plerotes anchietae (Seabra, 1900)
Plerotes anchietae (Seabra, 1900)
Plerotini
Rousettini
Rousettus aegyptiacus leachi (Smith, 1823)
Rousettus aegyptiacus (E. Geoffroy St.-Hilaire, 1810)
Scotonycterini
Casinycteris argynnis Thomas, 1910
Casinycteris argynnis Thomas, 1910
Scotonycteris zenkeri Matschie, 1894
Scotonycteris bergmansi Hassanin et al., 2015
Stenonycterini
Rousettus (Stenonycteris) lanosus Thomas, 1906
MICROCHIROPTERA
Hipposideridae
Stenonycteris lanosus Thomas, 1906
Hipposideridae
Asellia tridens (E. Geoffroy St.-Hilaire, 1813)
Hipposideros cyclops (Temminck, 1853)
Doryrhina cyclops (Temminck, 1853)
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Present study
Hayman et al. (1966)
Hipposideros abae J.A. Allen, 1917
Hipposideros abae J.A. Allen, 1917
Hipposideros beatus Andersen, 1906
Hipposideros beatus K. Andersen, 1906
Hipposideros beatus maximus Verschuren, 1957
Hipposideros caffer (Sundevall, 1846)
Hipposideros caffer caffer (Sundevall, 1846)
Hipposideros caffer (Sundevall, 1846)
Hipposideros caffer tephrus Cabrera, 1906
Hipposideros nanus J.A. Allen, 1917
Hipposideros camerunensis Eisentraut, 1956
Hipposideros camerunensis Eisentraut, 1956
Hipposideros fuliginosus (Temminck, 1853)
Hipposideros fuliginosus (Temminck, 1853)
Hipposideros caffer ruber (Noack, 1893)
Hipposideros ruber (Noack, 1893)
Hipposideros commersoni gigas (Wagner, 1845)
Macronycteris gigas (Wagner, 1845)
Hipposideros commersoni marungensis (Noack, 1887)
Macronycteris vittatus (Peters, 1852)
Megadermidae
Megadermatidae
Cardioderma cor (Peters, 1872)
Lavia frons frons (Geoffroy, 1810)
Lavia frons (E. Geoffroy St.-Hilaire, 1810)
Rhinolophidae
Rhinolophus alcyone alcyone Temminck, 1852
Rhinolophus alcyone Temminck, 1852
Rhinolophus blasii empusa (Andersen, 1904)
Rhinolophus blasii Peters, 1867
Rhinolophus clivosus Cretzschmar, 1826
Rhinolophus clivosus Cretzschmar, 1828
Rhinolophus clivosus zuluensis (Andersen, 1904)
Rhinolophus hildebrandti eloquens Andersen, 1905
Rhinolophus eloquens K. Andersen, 1905
Rhinolophus fumigatus Rüppell, 1842
Rhinolophus fumigatus Rüppell, 1842
Rhinolophus hildebrandti Peters, 1878
Rhinolophus hildebrandtii Peters, 1878
Rhinolophus hilli Aellen, 1973
Rhinolophus kahuzi Fahr & Kerbis Peterhans, 2013
Rhinolophus landeri lobatus (Peters, 1852)
Rhinolophus landeri Martin, 1838
Rhinolophus ruwenzorii Hill, 1942
Rhinolophus ruwenzorii J. Eric Hill, 1942
Rhinolophus simulator K. Andersen, 1904
Rhinolophus swinnyi Gough, 1908
Rhinolophus swinnyi Gough, 1908
Rhinolophus willardi Kerbis Peterhans & Fahr, 2013
Hipposideridae
Cloeotis percivali australis Roberts, 1917
Coleura afra (Peters, 1852)
Rhinonycteridae
Cloeotis percivali Thomas, 1901
VESPERTILIONIFORMI
Emballonuridae
Coleura afra (Peters, 1852)
Taphozous (Saccolaimus) peli (Temminck, 1853)
Saccolaimus peli (Temminck, 1853)
Taphozous mauritianus Geoffroy, 1818
Taphozous mauritianus E. Geoffroy St.-Hilaire, 1818
Taphozous (Liponycteris) nudiventris (Cretzschmar, 1826)
Taphozous nudiventris Cretzschmar, 1830
Taphozous sudani Thomas, 1915
Taphozous perforatus E. Geoffroy St.-Hilaire, 1818
Nycteridae
Nycteris arge Thomas, 1903
Nycteris arge Thomas, 1903
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Nycteris grandis Peters, 1865
Nycteris grandis Peters, 1865
Nycteris hispida (Schreber, 1775)
Nycteris hispida (Schreber, 1774)
Nycteris intermedia Aellen, 1959
Nycteris intermedia Aellen, 1959
Nycteris aethiopica Dobson, 1878
Nycteris macrotis Dobson, 1876
Nycteris macrotis Dobson, 1876
Nycteris major (Andersen, 1912)
Nycteris major (K. Andersen, 1912)
Nycteris nana (Andersen, 1912)
Nycteris nana (K. Andersen, 1912)
Nycteris parisii (de Beaux, 1924)
Nycteris capensis Smith, 1829
Nycteris thebaica E. Geoffroy St.-Hilaire, 1818
Nycteris thebaica Geoffroy, 1818
Molossidae
Tadarida (Chaerephon) aloysii-sabaudiae (Festa, 1907)
Tadarida (Tadarida) brunnea (Seabra, 1900)
Chaerephon aloysiisabaudiae (Festa, 1907)
Tadarida (Tadarida) ansorgei (Thomas, 1913)
Chaerephon ansorgei (Thomas, 1913)
Tadarida (Tadarida) cistura (Thomas, 1903)
Chaerephon bemmeleni (Jentink, 1879)
Chaerephon bivittatus (Heuglin, 1861)
Tadarida (Chaerephon) chapini J. A. Allen, 1917
Chaerephon chapini J.A. Allen, 1917
Chaerephon gallagheri (Harrison, 1975)
Chaerephon leucogaster (A. Grandidier, 1869)
Tadarida (Chaerephon) major (Trouessart, 1897)
Chaerephon major (Trouessart, 1897)
Tadarida (Chaerephon) nigeriae Thomas, 1913
Chaerephon nigeriae Thomas, 1913
Tadarida (Chaerephon) spillmanni (Monard, 1932)
Tadarida (Chaerephon) pumila (Cretzschmar, 1826)
Chaerephon pumilus (Cretzschmar, 1826)
Tadarida (Chaerephon) russata J.A. Allen, 1917
Chaerephon russatus J.A. Allen, 1917
Tadarida (Mops) condylura (A. Smith, 1833)
Tadarida (Mops) condylura condylura (A. Smith, 1833)
Mops (Mops) condylurus (A. Smith, 1833)
Tadarida (Mops) congica J.A. Allen, 1917
Mops (Mops) congicus J.A. Allen, 1917
Tadarida (Mops) demonstrator (Thomas, 1903)
Mops (Mops) demonstrator (Thomas, 1903)
Tadarida (Mops) midas (Sundevall, 1843)
Mops (Mops) midas (Sundevall, 1843)
Tadarida (Mops) niangarae J.A. Allen, 1917
Mops (Mops) niangarae J.A. Allen, 1917
Tadarida (Mops) chitauensis Hill, 1937
Tadarida (Mops) niveiventer (Cabrera & Ruxton, 1926)
Mops (Mops) niveiventer Cabrera & Ruxton, 1926
Tadarida (Mops) congica J.A. Allen, 1917
Mops (Mops) trevori J.A. Allen, 1917
Tadarida (Mops) leonis (Thomas, 1908)
Mops (Xiphonycteris) brachypterus (Peters, 1852)
Tadarida (Mops) nanula J.A. Allen, 1917
Mops (Xiphonycteris) nanulus J.A. Allen, 1917
? Xiphonycteris spurrelli Dollman, 1911
Mops (Xiphonycteris) spurrelli (Dollman, 1911)
Tadarida (Mops) thersites (Thomas, 1903)
Mops (Xiphonycteris) thersites (Thomas, 1903)
Myopterus albatus Thomas, 1915
Myopterus daubentonii Desmarest, 1820
Myopterus whitleyi (Scharff, 1900)
Myopterus whitleyi (Scharff, 1900)
Otomops martiensseni (Matschie, 1897)
Otomops martiensseni (Matschie, 1897)
Tadarida (Tadarida) fulminans (Thomas, 1903)
Tadarida fulminans (Thomas, 1903)
Tadarida (Tadarida) africana (Dobson, 1876)
Tadarida ventralis (Heuglin, 1861)
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Present study
Hayman et al. (1966)
Vespertilionidae
Miniopteridae
Miniopterus fraterculus Thomas & Schwann, 1906
Miniopterus inlatus rufus Sanborn, 1936
Miniopterus inlatus Thomas, 1903
Miniopterus minor Peters, 1867
Miniopterus minor Peters, 1867
Miniopterus natalensis arenarius Heller, 1912
Miniopterus schreibersi vicinior (J.A. Allen, 1917)
Miniopterus natalensis (A. Smith, 1833)
Miniopterus schreibersi villiersi Aellen, 1956
Vespertilionidae
Kerivoulinae
Kerivoula argentata Tomes, 1861
Kerivoula argentata Tomes, 1861
Kerivoula cuprosa Thomas, 1912
Kerivoula harrisoni muscilla (Thomas, 1906)
Kerivoula harrisoni lucia (Hinton, 1920)
Kerivoula lanosa (A. Smith, 1847)
Kerivoula phalaena Thomas, 1912
Kerivoula phalaena Thomas, 1912
Kerivoula smithi Thomas, 1880
Kerivoula smithii Thomas, 1880
Kerivoula ? aerosa Tomes, 1858
Myotinae
Myotis bocagei (Peters, 1870)
Myotis bocagei bocagei (Peters, 1870)
Myotis bocagei cupreolus Thomas, 1904
Myotis bocagii (Peters, 1870)
Myotis bocagei hildegardeae Thomas, 1904
Myotis tricolor (Temminck, 1832)
Myotis tricolor (Temminck, 1832)
Myotis welwitschi venustus (Matschie, 1899)
Myotis welwitschii (Gray, 1866)
Scotophilinae
Scotophilus nigrita (Schreber, 1775)
Scotophilus nigrita herero Thomas, 1906
Scotophilus dinganii (A. Smith, 1833)
Scotophilus leucogaster (Cretzschmar, 1826)
Scotophilus leucogaster (Cretzschmar, 1826)
Scotophilus gigas Dobson, 1875
Scotophilus nigrita (Schreber, 1774)
Scotophilus nigrita nux Thomas, 1904
Scotophilus nux Thomas, 1904
Vespertilioninae
Glauconycteris alboguttatus J.A. Allen, 1917
Glauconycteris alboguttata J.A. Allen, 1917
Glauconycteris argentata (Dobson, 1875)
Glauconycteris argentata (Dobson, 1875)
Glauconycteris beatrix Thomas, 1901
Glauconycteris beatrix Thomas, 1901
Glauconycteris curryae Eger & Schlitter, 2001
Glauconycteris humeralis J.A. Allen, 1917
Glauconycteris humeralis J.A. Allen, 1917
Glauconycteris poensis (Gray, 1842)
Glauconycteris poensis (Gray, 1842)
Glauconycteris superba superba Hayman, 1939
Glauconycteris superba Hayman, 1939
Glauconycteris variegata papilio (Thomas, 1905)
Glauconycteris variegata (Tomes, 1861)
Hypsugo anchietae (Seabra, 1900)
Pipistrellus crassulus Thomas, 1904
Hypsugo crassulus (Thomas, 1904)
Hypsugo cf. eisentrauti (Hill, 1968)
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Pipistrellus musciculus Thomas, 1913
Hypsugo musciculus (Thomas, 1913)
Laephotis wintoni angolensis (Monard, 1935)
Laephotis angolensis Monard, 1935
Laephotis botswanae Setzer, 1971
Mimetillus moloneyi (Thomas, 1891)
Mimetillus moloneyi (Thomas, 1891)
Mimetillus thomasi Hinton, 1920
Neoromicia brunnea (Thomas, 1880)
Eptesicus capensis (Smith, 1829)
“Eptesicus garambae” J.A. Allen, 1917
Neoromicia capensis (A. Smith, 1829)
“Eptesicus grandidieri” (Dobson, 1876)
“Eptesicus minutus” (Temminck, 1835)
Neoromicia guineensis (Bocage, 1889)
Pipistrellus culex Thomas, 1911
Neoromicia nana (Peters, 1852)
Pipistrellus nanus nanus Peters, 1852
Eptesicus rendalli (Thomas, 1889)
Neoromicia rendalli (Thomas, 1889)
Neoromicia somalica (Thomas, 1901)
Eptesicus tenuipinnis (Peters, 1872)
Neoromicia tenuipinnis (Peters, 1872)
Eptesicus tenuipinnis ater (J.A. Allen, 1917)
Neoromicia zuluensis (Roberts, 1924)
Nycticeius (Scoteinus) schlieffeni Peters, 1860
Nycticeius (Scoteinus) schlieffeni albiventer
Thomas & Wroughton, 1908
Nycticeinops schlieffenii (Peters, 1859)
Eptesicus grandidieri (Dobson, 1876)
Pipistrellus grandidieri (Dobson, 1876)
Pipistrellus hesperidus (Temminck, 1840)
Pipistrellus nanulus Thomas, 1904
Pipistrellus nanulus Thomas, 1904
Pipistrellus (Scotozous) rüppelli (Fischer, 1826)
Pipistrellus (Scotozous) rüppelli fuscipes
(Thomas, 1913)
Pipistrellus rueppellii (Fischer, 1829)
Nycticeius (Scotoecus) albofuscus woodi (Thomas, 1917)
Scotoecus albofuscus (Thomas, 1890)
Nycticeius (Scotoecus) hirundo hindei (Thomas, 1901)
Scotoecus hindei Thomas, 1901
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European Journal of Taxonomy 382: 1–327 (2017)
Key to the bat species of the CRB region
This key is primarily modiied from Hayman et al. (1966), with additional information from Hayman &
Hill (1971), Bergmans (1997), Cotterill (2001b), Happold & Happold (2013), Kerbis Peterhans et al.
(2013) and Foley et al. (2014).
1. Cheekteeth without W-pattern of cusps; second inger with claw; tail absent or rudimentary;
interfemoral membrane reduced; ears very short, well separated, no tragus; margin of ear forming a
complete ring (family Pteropidae) ..................................................................................................... 9
– Cheekteeth with W-pattern of cusps; second inger entirely included in patagium and without claw;
tail generally well developed; interfemoral membrane developed; ears very short to very long,
widely separed or joined; tragus present in most cases (except for Rhinolophidae, Rhinonycteridae
and Hipposideridae) ........................................................................................................................... 2
2. Tail more or less completely included in uropatagium (in certain forms the last vertebra can be
free) ................................................................................................................................................... 3
– Tail not completely included in uropatagium .................................................................................... 8
3. No nasal appendages; tragus conspicuous; ears very short to very long, widely separated or joined,
but not fused in the mediosagittal plane .......................................................................................... 64
– Nasal appendages present .................................................................................................................. 4
4. No tragus or tragus inconspicuous; ears very short to medium, not fused mediosagitally ............... 5
– Tragus present (sometimes small); ears fused or very close to each other mediosagitally ............... 6
5. Posterior lealet of nose-leaf low, rounded or with three pointed projections (tridentate); two pedal
phalanges; tragus inconspicuous...................................................................................................... 43
– Posterior lealet of nose-leaf high, erect, pointed; three pedal phalanges; no tragus (family
Rhinolophidae, genus Rhinolophus) ................................................................................................ 53
6. Tail shorter than uropatagium; no postorbital processes; no premaxillae present; noseleaf very large,
comparatively simple; tragus very conspicuous, biid (family Megadermatidae) ............................. 7
– Tail as long as uropatagium; last vertebra Y- or T-shaped; postorbital process and premaxillae
present; noseleaf is formed by a longitudinal slit bordered by leshy outgrowths; tragus conspicuous,
not biid (family Nycteridae) ........................................................................................................... 35
7. Noseleaf less than 15 mm; pelage grey without yellow tinge; frontal shield of skull strongly concave,
minimum breadth of shield less than 6.5 mm ...................................Cardioderma cor (Peters, 1872)
– Noseleaf over 18 mm; pelage bluish-grey, generally with yellowish tinge; frontal shield of skull
comparatively lat, minimum breadth of shield over 6.5 mm .............................................................
........................................................................................ Lavia frons (E. Geoffroy St.-Hilaire, 1810)
8. Basal part of tail included in uropatagium; distal part of tail resting free on the dorsal side of the
uropatagium; ears very short, widely separated, not fused, backwards-pointing; tragus conspicuous
(family Emballonuridae) .................................................................................................................. 31
– Distal part of tail (ca half the width of the interfemoral membrane) not included in uropatagium
and not resting on the dorsal side of the uropatagium; ears short, mostly united to each other by a
cutaneous wall (family Molossidae) .............................................................................................. 108
9. Tongue simple, ixed to loor of mouth by posterior half, and without unfringed iliform papillae at
tip; muzzle not elongated ................................................................................................................. 10
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– Tongue more extensible, ixed to loor of mouth by its posterior third, its terminal
fourth or ifth covered above with unfringed iliform papillae (“brush-tipped”), muzzle
elongated .................................................................. Megaloglossus woermanni Pagenstecher, 1885
10. Facial pattern of white nose and eye patches....................................................................................11
– No facial pattern .............................................................................................................................. 12
11. White tufts at base of ears inconspicuous or absent; bony palate extending well beyond teeth;
rostrum, when viewed laterally, not upturned........... Scotonycteris bergmansi Hassanin et al., 2015
– White tufts at base of ears present, although sometimes inconspicuous; no postdental bony palate;
rostrum, when viewed laterally, upturned.................................Casinycteris argynnis Thomas, 1910
12. No white tufts present at base of ears, or tufts inconspicuous ......................................................... 13
– White tufts at base of ears present, although sometimes inconspicuous ......................................... 17
13. Facial axis of skull not noticeably delected against basicranial axis; forearm 54–90 mm, length
second digit 71–81 % of forearm; toes partially webbed; cheek teeth squarish or oblong; 6 + 2 or
7 + 2 palatal ridges (genus Myonycteris) ......................................................................................... 14
– Facial axis of skull more or less strongly delected against basicranial axis; forearm 75–130 mm,
length of second digit 64–71 % of forearm; toes not webbed; cheek teeth oblong; 7 + 1 palatal
ridges................................................................................................................................................ 16
14. Tibia dorsally furred; P4 and M1 squarish in outline; two lower incisors; M3 reduced but present;
forearm 68–88 mm (♂♂) and 67–90 mm (♀♀); weight 60–97 g (♂♂) and 60–100 g (♀♀); third
metacarpal 73–75 % of forearm length; tibia 42–46 % of forearm length ..........................................
............................................................................................. Myonycteris angolensis (Bocage, 1898)
– Tibia dorsally practically naked over distal quarter to third; P4 and M1 oblong in outline; one or two
lower incisors; M3 reduced or absent; forearm 54–76 mm; third metacarpal 67–73 % of forearm
length; tibia 36–43 % of forearm length .......................................................................................... 15
15. Interfemoral membrane only furred near legs; palatal ridge pattern 6 + 2 (sometimes 7 + 2); two
lower incisors; M3 absent; forearm 65–76 mm ..........................Myonycteris relicta Bergmans, 1980
– Interfemoral membrane wholly furred; palatal ridge pattern 7 + 2; one or two lower incisors;
M3 reduced but normally present; forearm 54–68 mm ........... Myonycteris torquata (Dobson, 1878)
16. Forearm 115–130 mm; M1 equal in length to M2 and M3 combined; tympanic forming a short tubular
bony auditory meatus; premaxillae spaced in front; colour generally tawny .....................................
…………… ......................................................................................... Eidolon helvum (Kerr, 1792)
– Forearm 79–107 mm; M1 shorter than M2 and M3 combined; tympanic not forming a bony auditory
meatus; premaxillae in contact or co-ossiied in front; colour generally not tawny nor yellowish ....
......................................................................................................................................................... 22
17. Cheekteeth 4/5 , 4/6 or 5/6; unusual weak dentition; interfemoral membrane greatly reduced; calcar
absent or rudimentary; forearm 47–53 mm ...................................Plerotes anchietae (Seabra, 1900)
– Cheekteeth 3/5; interfemoral membrane well developed; calcar present; forearm generally larger
(except for Micropteropus) .............................................................................................................. 18
18. Size very large, forearm 118–133 mm; rostrum very deep, end of muzzle truncated, ending in leshy
plate; no white shoulder tuft in ♀ .................................... Hypsignathus monstrosus H. Allen, 1862
– Size smaller, forearm 45–100 mm; rostrum shallow, end of muzzle simple; white shoulder tuft in ♀ ....
......................................................................................................................................................... 19
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19. Forearm 60–100 mm........................................................................................................................ 20
– Forearm 45–55 mm.......................................................................................................................... 21
20. Six prominent palatal ridges, not more than two being post-dental; the latter not differentiated from
inter-dental ridges; post-dental palate strongly concave posteriorly (genus Epomophorus) ........... 24
– Five, or 9–11, palatal ridges, with sometimes from 5 to 7 post-dental; latter strongly differentiated
from inter-dental ridges; post-dental palate lattened posteriorly (genus Epomops) ....................... 29
21. Rostrum broad; 5–6 palatal ridges, obviously divided by median groove (genus Micropteropus) . 30
– Rostrum slender; 12–13 palatal ridges, only posterior group narrowly divided ................................
........................................................................................... Nanonycteris veldkampii (Jentink, 1888)
22. Pm1 subequal in bulk to a lower incisor; premaxillae co-ossiied; braincase slightly delected; wings
from second toe; fur long and silky; forearm 79–83.5 mm .................................................................
............................................................................................. Myonycteris angolensis (Bocage, 1898)
– Pm1 much larger in bulk than a lower incisor; premaxillae in contact; braincase conspicuously
delected; forearm at least 82.0 mm; tibia furred or naked .............................................................. 23
23. Braincase moderately delected; width of last upper premolar about ⅓ of palate between fronts of last
upper premolars, clearly larger than half their lengths; wings from irst toe (occasionally between
irst and second); antitragal lobe distinct; fur short; tibiae dorsally practically naked; predominantly
in lowland areas; forearm 85–107 mm (♂♂) and 82–107 mm (♀♀); weight 111–168 g (♂♂) and
100–155 g (♀♀) . .......................................... Rousettus aegyptiacus (E. Geoffroy St.-Hilaire, 1810)
– Braincase strongly delected; cheekteeth excessively narrow; width of last upper premolar about 1/5
of that of the palate between fronts of last upper premolars, about half their lengths or slightly more;
wings from second toe (occasionally between irst and second); antitragal lobe obsolete; fur long
and coarse; tibiae dorsally furred; mostly above 1000 m; forearm 85–94 mm (♂♂) and 85–95 (♀♀);
weight 102–140 g (♂♂) and 94–162 g (♀♀) ...........................Stenonycteris lanosus Thomas, 1906
24. One conspicuous, thick postdental palatal ridge; total length of skull 44.4–57.3 mm (♂♂), 41.0–
55.5 mm (♀♀); forearm 72–95 mm (♂♂), 67–89 mm (♀♀) ……………………………………………………
………………………………………………………………………………
Epomophorus wahlbergi (Sundevall, 1846)
– Two postdental palatal ridges .......................................................................................................... 25
25. Size small to medium; total length of skull < 50 mm (♂♂) and < 46 mm (♀♀); forearm < 80 mm
(♂♂) and < 76 mm (♀♀) ................................................................................................................. 26
– Size larger; total length of skull > 51.8 mm (♂♂) and > 44 mm (♀♀); forearm > 80 mm (♂♂) and
> 75 mm (♀♀); fourth palal ridge mid-way between third and ifth ...................................................
............................................................................................... Epomophorus crypturus Peters, 1852
26. Two thick postdental palatal ridges in ♂♂, one postdental and one partially postdental in ♀♀; total
length of skull 47.1 mm (♂), 38.7 and >41.2 mm (♀♀); forearm 77 mm (♂♂), 68–74 mm (♀♀)
(otherwise known from Malawi and SW Tanzania only) …………………………………………………………
…………………………………………………………………Epomophorus anselli Bergmans & Van Strien, 2004
– Two thick postdental palatal ridges in both sexes; total length of skull < 48 mm (♂♂) and < 46 mm
(♀♀); forearm < 80 mm (♂♂) and < 76 mm (♀♀) ......................................................................... 27
27. Size smaller; total length of skull < 40 mm (♂♂) and < 36.5 mm (♀♀) ...........................................
........................................................................... Epomophorus minimus Claessen & De Vree, 1991
– Size medium; total length of skull > 40 mm (♂♂) and > 36.5 mm (♀♀) ....................................... 28
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28. Size larger; forearm 66.7–80.3 mm (♂♂) and 64.8–78.3 mm (♀♀); weight 54–99 g (♂♂)
and 51–81 g (♀♀); mainly woodlands. Distributed from NE Nigeria over S Sudan to
Eritrea and to NE Tanzania; furthermore known from few localities in DRC, Malawi and
SE Kenya ......................................................................... Epomophorus labiatus (Temminck, 1837)
– Size smaller; forearm 57.0–68.9 mm (♂♂) and 54.1–67.6 mm (♀♀); weight 32–58 g (♂♂) and
25–62 g (♀♀); mainly woodlands. Distributed in E Sudan, Ethiopia, Somalia, lower parts of Kenya
to S Malawi. ................................................................................Epomophorus minor Dobson, 1880
29. Four thick, prominent interdental and about 5–7 thinner postdental palate ridges ............................
......................................................................................................Epomops franqueti (Tomes, 1860)
– Three thick and prominent interdental palate ridges; postdental palate with two pairs of
prominent, triangular ridges at middle and one or a few thin and serrate ridges at palation
border ........................................................................................... Epomops dobsonii (Bocage, 1889)
30. Forearm 50–53 mm; total skull length 29–29.8 mm ..............Micropteropus pusillus (Peters, 1868)
– Forearm 59–63 mm; total skull length 33.2–33.4 mm ... Micropteropus intermedius Hayman, 1963
31. Six lower incisors; two or four upper incisors; forearm normally less than 52 mm ...........................
.................................................................................................................Coleura afra (Peters, 1852)
– Four lower incisors; two upper incisors (very minute, often absent); forearm more than 57 mm .. 32
32. Forearm more than 80 mm (83–96 mm); total skull length 30–33 mm; skull with shallow frontal
depression; no radio-metacarpal pouch; bullae complete; fur dark at base .........................................
...................................................................................................Saccolaimus peli (Temminck, 1853)
– Forearm less than 80 mm; total skull length 20–28 mm; skull with shallow (T. nudiventris) or deep
frontal depression; radio-metacarpal pouch present; bullae incomplete; fur light at base .............. 33
33. Forearm more than 70 mm, posterior part of body naked above and below .......................................
......................................................................................... Taphozous nudiventris Cretzschmar, 1830
– Forearm less than 70 mm; body fully haired ................................................................................... 34
34. Dorsally light brown, ventrally pale grey to dark greyish-brown; throat usually sepia-brown, darker
than belly; no gular patch in either sex; forearm 60–67 mm ...............................................................
..........................................................................Taphozous perforatus E. Geoffroy St.-Hilaire, 1818
– Dorsally grizzled grey, ventrally pure white or stained yellowish; throat white; large gular sac in
male; forearm 58–64 mm...............................Taphozous mauritianus E. Geoffroy St.-Hilaire, 1818
35. Second premolar of lower jaw large (equal to or larger than half of the irst lower premolar); I1–2 biid
(I2 exceptionally triid); marked constriction halfway along the height of the tragus (arge-group) 38
– Second lower premolar small (smaller than half of the irst lower premolar); I1–2 bi- or triid; no
constriction in the tragus .................................................................................................................. 36
36. I1–2 biid ............................................................................................................................................ 37
– I1–2 triid (hispida-group).................................................................................................................. 41
37. Tragus semilunate (macrotis-group) ................................................................................................ 42
– Tragus pyriform (free portion narrowest at base; its outer and inner margin evenly convex);
forearm length 42–52 mm; ear length 28–37 mm (thebaica-group) ………………………………………………
……………………………………………………… ............................ Nycteris thebaica E. Geoffroy St.-Hilaire, 1818
38. Forearm length more than 39 mm; skull length more than 19 mm ................................................. 39
– Forearm length less than 38 mm; skull length less than 19 mm ...................................................... 40
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European Journal of Taxonomy 382: 1–327 (2017)
39. Forearm length 47.3–50 mm; skull length 20.9–22.4 mm; ear length measuring 50–65 % of forearm
length ......................................................................................... Nycteris major (K. Andersen, 1912)
– Forearm length 39.5–46 mm; skull length 19.3–20.9 mm; ear length measuring 63–77 % of forearm
length ......................................................................................................Nycteris arge Thomas, 1903
40. Forearm length 36.5–37 mm; skull length 18–18.7 mm; tibia longer than half the forearm length ...
........................................................................................................Nycteris intermedia Aellen, 1959
– Forearm length 32–36.2 mm; skull length 15.9–16.6 mm; tibia shorter than half the forearm length
.....................................................................................................Nycteris nana (K. Andersen, 1912)
41. Forearm length 39–43 mm............................................................. Nycteris hispida (Schreber, 1774)
– Forearm length 57–66 mm....................................................................Nycteris grandis Peters, 1865
42. Forearm length 45–52 mm; ear length 22–34 mm .......................... Nycteris macrotis Dobson, 1876
– Forearm length 37–42 mm; ear length 29–34 mm .........................Nycteris parisii (de Beaux, 1924)
43. Posterior lealet of noseleaf simple in outline, elliptical or rounded, not tridentate; ears either separate
or united by low band; rostrum at least half as long as braincase; small upper premolar present;
forearm over 40 mm (family Hipposideridae, genera Doryrhina, Hipposideros, Macronycteris) .. 45
– Posterior lealet of noseleaf tridentate; ears always separate; rostrum sometimes less than half as
long as braincase; small upper premolar present or absent; forearm 31–54 mm ............................ 44
44. Rostrum less than half as long as braincase; ears low, 8 mm, rounded, rim-like; tridentate nose-leaf;
small, forearm 31–35 mm; tail not projecting beyond end of interfemoral membrane; two upper
premolars on each side; M3 not reduced (family Rhinonycteridae) …Cloeotis percivali Thomas, 1901
– Rostrum at least half as long as braincase; ears large, 18–20 mm with pointed tip; larger, forearm 44–
54 mm; tail projecting up to 7 mm beyond end of interfemoral membrane; one upper premolar on each
side; M3 greatly reduced (family Hipposideridae) ..... Asellia tridens (E. Geoffroy St.-Hilaire, 1813)
45. Ears long and narrow, pointed; cochleae greatly expanded, their width equal to at least 4 times the
distance between them ..................................................................................................................... 49
– Ears short, broad, rounded or triangular; cochleae not expanded, their width approximately equal to
the distance between them ............................................................................................................... 46
46. Ears rounded or broadly triangular; bluntly pointed upper incisors weak, the outer lobe obsolescent
or obsolete; crown area of outer lower incisors equal to or only slightly greater than that of the inner
lower incisors ................................................................................................................................... 50
– Ears triangular; pointed upper incisors strong, usually retaining much of the outer lobe; crown area
of the outer lower incisors much greater than that of the inner lower incisors................................ 47
47. Ears with a small projection at the antitragal fold; sphenoidal bridge narrow, not concealing lateral
apertures; pterygoid wings underdeveloped; forearm 58–66 mm ……………………………………… ………
………………………………………………………………………………………Hipposideros abae J.A. Allen, 1917
– Ears without antitragal modiication; sphenoidal bridge wide, partially concealing lateral apertures;
pterygoid wings expanded; forearm 78–114 mm ............................................................................ 48
48. Number of lateral lealets 3; smaller on average, forearm 93–106 mm (♂♂) and 84–101 mm (♀♀);
pelage shorter, not slightly wooly; predominantly in savannas ...........................................................
................................................................................................. Macronycteris vittatus (Peters, 1852)
– Number of lateral lealets 3 or 4; larger on average, forearm 100–124 mm (♂♂) and 95–116 (♀♀);
pelage longer, slightly wooly; predominantly in forests..............Macronycteris gigas Wagner, 1845
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VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
49. Smaller, length of forearm less than 74.0 mm; anteorbital foramen relatively large ..........................
................................................................................................ Doryrhina cyclops (Temminck, 1853)
– Larger, length of forearm exceeding 74.0 mm; anteorbital foramen relatively small .........................
.....................................................................................Hipposideros camerunensis Eisentraut, 1956
50. Posterior nose-leaf without a transverse supplementary structure developed from its posterior face
or with such a structure low, undeveloped and lacking a serrated upper edge ....................................
......................................................................................Hipposideros fuliginosus (Temminck, 1853)
– Posterior nose-leaf having a transverse supplementary structure with a serrated upper edge developed
from its posterior face ...................................................................................................................... 51
51. Anterior upper premolar small, slightly extruded from toothrow, or compressed between canine and
second upper premolar ..................................................................................................................... 52
– Anterior upper premolar minute, extruded from toothrow, canine and second upper premolar in
contact or nearly so ........................................................... Hipposideros beatus (K. Andersen, 1906)
52. Frontal sac opening in both sexes; forearm generally less than 48 mm (42–52 mm); tibia 18–22 mm;
dorsal pelage greyish or pale orange; median posterior narial compartiment of nasal swelling
relatively narrow, with wide lateral inlations ....................... Hipposideros caffer (Sundevall, 1846)
– Frontal sac opening in ♂♂ only; forearm generally over 48 mm (47–55 mm); tibia 18–24 mm;
dorsal pelage brownish or rufous; median posterior narial compartment of nasal swelling relatively
wide, with narrow lateral inlations ............................................. Hipposideros ruber (Noack, 1893)
53. First upper premolar placed between the upper canine and second premolar or slightly reduced and
partially displaced labially ............................................................................................................... 54
– First upper premolar missing or very reduced and external to toothrow; the canine and second
premolar touching each other or nearly so ...................................................................................... 62
54. Connecting process of nose-leaf rises to a distinct point ................................................................. 55
– Connecting process of nose-leaf low, subtriangular or obsolete ..................................................... 56
55. First phalange of fourth inger long, more than half as long as second, 24–28 % of metacarpal length;
sella wedge-shaped, top narrow; lancet subtriangular or hastate, tip rounded ....................................
......................................................................................................... Rhinolophus blasii Peters, 1867
– First phalange of fourth inger shorter, 19–23 % of metacarpal length; sella sides slightly concave;
lancet hastate, tip bluntly pointed ................................................ Rhinolophus landeri Martin, 1838
56. Connecting process of nose-leaf obsolete or poorly developed; ears large; forearm 50 mm or
more ................................................................................................................................................. 57
– Connecting process of nose-leaf low and rounded or subtriangular; forearm 55 mm or less ......... 60
57. Eight or nine internal ear folds ........................................................................................................ 58
– 11 or 12 internal ear folds ................................................................................................................ 59
58. Lancet tip truncated, only moderately exceeding height of sella; ear with eight folds; larger,
forearm 55–62 mm; skull with braincase conspicuously constricted behind the mastoid process;
infraorbital bridge longer and more slender; lower molars comparatively small; ear length 32–
40 mm ..............................................................................Rhinolophus ruwenzorii J. Eric Hill, 1942
– Lancet tip bluntly pointed, conspicuously exceeding height of sella; ear with nine folds;
smaller, forearm 54 mm; skull with braincase not conspicuously constricted behind the mastoid
process; infraorbital bridge shorter and stouter; lower molars comparatively large; ear length
28.5 mm ............................................................................................. Rhinolophus hilli Aellen, 1973
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European Journal of Taxonomy 382: 1–327 (2017)
59. 11 internal ear folds; forearm length 54.5 mm; ear length 34.5 mm ...................................................
......................................................................... Rhinolophus kahuzi Fahr & Kerbis Peterhans, 2013
– 12 internal ear folds; forearm length 49.7–51.5 mm; ear length 24.2–29 mm ....................................
........................................................................ Rhinolophus willardi Kerbis Peterhans & Fahr, 2013
60. Forearm length 49–54 mm; skull length 22.4–23.3 mm ...... Rhinolophus alcyone Temminck, 1853
– Forearm length 40–45 mm; skull length 17.6 mm or less ............................................................... 61
61. Sella comparatively narrow (1.2–1.3 mm prior to preservation); lancet subtriangular, almost hastate,
sides concave, tip bluntly pointed; horseshoe 6.0–7.4 mm .........Rhinolophus swinnyi Gough, 1908
– Sella comparatively broad (1.5–1.7 mm after preservation); lancet subtriangular or hastate, tip
somewhat rounded; horseshoe 6.7–9.0 mm .................... Rhinolophus simulator K. Andersen, 1904
62. Face and edges of sella naked; forearm length 50–56 mm; lancet hastate, tip rounded; horseshoe 6.6–
9.6 mm, narrower as muzzle; nasal swellings very low; frontal depression very shallow; supraorbital
ridges weak; interpterygoid groove absent or very indistinct ..............................................................
........................................................................................... Rhinolophus clivosus Cretzschmar, 1828
– Face and edges of sella with long hairs ........................................................................................... 63
63. Forearm length 60–67 mm; tibia 26–31 mm; ear length 26–36 mm; baculum > 3 mm; anterior
medial nasal swellings longer than broad ............................ Rhinolophus hildebrandtii Peters, 1878
– Forearm length 53–63 mm; tibia 22–25 mm; ear length 21–38 mm; baculum < 3 mm......................
......................................................................................... Rhinolophus eloquens K. Andersen, 1905
– Forearm length 47–60 mm; tibia 19–24 mm; ear length 19–28 mm; anterior medial nasal swellings
broader than long ...................................................................Rhinolophus fumigatus Rüppell, 1842
64. Second phalange of third digit nearly 3 times as long as the irst (family Miniopteridae; genus
Miniopterus) .................................................................................................................................... 75
– Second phalange of the third digit shorter (family Vespertilionidae) .............................................. 65
65. Ear slightly but evidently funnel-formed; very high braincase (subfamily Kerivoulinae; genus
Kerivoula) ........................................................................................................................................ 76
– Ears largely open at their base and not funnel-formed; normal braincase....................................... 66
66. Six Pm and M in upper and lower jaw (subfamily Myotinae, genus Myotis) ................................. 80
– Less than 6 Pm and M ..................................................................................................................... 67
67. One upper incisor and M1 and M2 with W-pattern distorted or nearly absent (subfamily Scotophilinae;
genus Scotophilus) ........................................................................................................................... 82
– Two upper incisors or 1 upper incisor and M1 and M2 with W-pattern normal (subfamily
Vespertilioninae) .............................................................................................................................. 68
68. One upper incisor ............................................................................................................................. 85
– Two upper incisors ........................................................................................................................... 69
69. Five Pm and M in upper and lower jaw (genera Neoromicia / Pipistrellus / Hypsugo) .................. 99
– Four Pm and M in upper and 5 in lower jaw ................................................................................... 70
70. Ears very enlarged, about 15 mm and more (genus Laephotis)....................................................... 74
– Ears normally much less enlarged ................................................................................................... 71
71. Very short and broad rostrum; the height of the braincase (with the bullae) more than ⅔ of the
condylo-basal length; lower lip with leshy lobes at angle of mouth (genus Glauconycteris) ........ 87
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VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
– Height of the braincase (bullae included) much less than the distance from incisors to posterior
zygoma root ..................................................................................................................................... 72
72. Very shortened wings; the third digit not longer than head and body length; skull much lattened
(genus Mimetillus) ........................................................................................................................... 73
– Normal wings; skull not especially lattened (genus Neoromicia) .................................................. 94
73. Pelage dark brown; wing membranes dark; occurring in forest habitats.............................................
................................................................................................. Mimetillus moloneyi (Thomas, 1891)
– Pelage pale brown; wing membranes lighter; occurring in savanna habitats ......................................
....................................................................................................... Mimetillus thomasi Hinton, 1920
74. On average larger; greatest skull length 14.1–15.5 mm; ear length 17–22 mm; forearm length 30–
39 mm; palatal length 6.2–7.3 mm ............................................... Laephotis botswanae Setzer, 1971
– On average smaller; greatest skull length 13.7–13.8 mm; ear length 15–18 mm; forearm length
32–36 mm; palatal length 6.1–6.2 mm ...................................... Laephotis angolensis Monard, 1935
75. Forearm length 45–52 mm; greatest skull length 16.4–17.8 mm; hind foot (including claw) 7–11 mm;
total length 92–128 mm; weight 8–19 g ..................................... Miniopterus inlatus Thomas, 1903
– Forearm length 35–42 mm; greatest skull length 11.1–14.5 mm; hind foot (including claw) 5–10 mm;
total length 82–97 mm; weight 5–10 g ............................................ Miniopterus minor Peters, 1867
– Forearm length 42–47 mm; greatest skull length 13.6–15.5 mm; hind foot (including claw) 6–11 mm;
total length 104–115 mm; weight 6–11 g .......................... Miniopterus natalensis (A. Smith, 1833)
– Forearm length 41–45 mm; greatest skull length 13.3–15.5 mm; hind foot (including claw) 7–11 mm;
total length 88–103 mm; weight 7–11 g ........... Miniopterus fraterculus Thomas & Schwann, 1906
76. Comb-like fringe of hairs on interfemoral membrane well developed............................................ 77
– Comb-like fringe of hairs on interfemoral membrane absent or scarcely developed ...................... 78
77. Dorsal pelage bright reddish-brown with conspicuous frosting (highlighted tips); ventral pelage
cream, dirty-white or white; size on average larger, forearm length 28–40 mm; skull longer and more
robust, greatest length 14.7–16.5 mm; outer lower incisors usually with three cusps, sometimes four;
inner upper incisors uni-, bi- or tricuspid ......................................Kerivoula argentata Tomes, 1861
– Dorsal pelage dark brown to pale grey with (in)conspicuous frosting, but not bright reddish-brown with
conspicuous frosting; ventral pelage sepia brown, grey to whitish; size on average smaller, forearm
length 26–34 mm; skull shorter and more delicate, greatest length 11.2–13.5 mm; outer lower incisors
with two or three cusps; inner upper incisors uni- or bicuspid .... Kerivoula lanosa (A. Smith, 1847)
78. Size smaller; forearm length 25–30 mm; dorsal pelage pale reddish-brown, fawn-brown or greyishbrown, usually darker at base, without frosting; outer lower incisors unicuspid; inner upper incisors
uni- or bicuspid ............................................................................ Kerivoula phalaena Thomas, 1912
– Size larger; forearm length 30 mm or more; dorsal pelage dark, frosted; inner upper incisors
bicuspid ............................................................................................................................................ 79
79. Dorsal pelage blackish or dark brown; forearm length 32–36 mm; lower incisors
unicuspid .......................................................................................... Kerivoula smithii Thomas, 1880
– Dorsal pelage dark brown with terminal third bright rusty-brown or chocolate brown; forearm length
30–33 mm; lower incisors tricuspid .............................................. Kerivoula cuprosa Thomas, 1912
80. With dichromatic (black and orange) wing membranes; forearm length 52–60 mm; sparse fringe
of bristle-like hairs along margin of interfemoral membrane between tips of calcars and tip of
tail .....................................................................................................Myotis welwitschii (Gray, 1866)
119
European Journal of Taxonomy 382: 1–327 (2017)
– Wing membranes uniformly dark; size smaller; no fringe of bristle-like hairs along margin of
interfemoral membrane .................................................................................................................... 81
81. Larger, forearm length 47–53 mm; wing membrane uniformly dark or inconspicuously particoloured,
black on most of the membrane and orange to dark red along the ingerbones; ventral and dorsal
hairs tricoloured ............................................................................ Myotis tricolor (Temminck, 1832)
– Smaller, forearm length 33–42 mm; wing membrane uniformly blackish-brown; ventral hairs
bicoloured; dorsal hairs bi- or tricoloured ............................................Myotis bocagii (Peters, 1870)
82. Very large, forearm length 78–88 mm ...................................... Scotophilus nigrita (Schreber, 1774)
– Smaller, forearm not over 61 mm .................................................................................................... 83
83. Ventral pelage dark brown to reddish-orange; habitat rainforest.......Scotophilus nux Thomas, 1904
– Ventral pelage pure white, yellowish, yellow, orange-yellow; habitat savannas and rainforestsavanna mosaics............................................................................................................................... 84
84. Ventral pelage pure white to medium brown, never yellowish; dorsal pelage pale to medium
brown, never tinged with yellow or green; forearm length 44–58 mm; skull total length 16–
18 mm ..........................................................................Scotophilus leucogaster (Cretzschmar, 1826)
– Ventral pelage pale yellow, bright yellow or orange-yellow, not tinged with brown; dorsal pelage
sepia brown, greenish-brown, greyish-brown or reddish-brown; forearm 47–60 mm; skull total
length 18–21 mm ................................................................... Scotophilus dinganii (A. Smith, 1833)
85. Penis short (5.6–6.2 mm); upper canine with anterior surface rounded, not grooved; anterior lower
premolar ca half height of posterior premolar ......................Nycticeinops schlieffenii (Peters, 1859)
– Penis extremely long (ca 9–16 mm); upper canine with anterior surface lat and grooved; anterior lower
premolar at least two-thirds height of posterior premolar, usually subequal (genus Scotoecus) ..... 86
86. Wing membranes dark-brown, slightly translucent ..........................Scotoecus hindei Thomas, 1901
– Wing membranes light, semi-translucent, becoming brownish around edges and near body .............
................................................................................................Scotoecus albofuscus (Thomas, 1890)
87. Wing and interfemoral membranes conspicuously veined; wings pale; colour of dorsal hairs creamybuff to yellowish-fawn (tricoloured or inconspicuously bicoloured); forearm length 38–45 mm ......
............................................................................................ Glauconycteris variegata (Tomes, 1861)
– Wing and interfemoral membranes not conspicuously veined ........................................................ 88
88. Dorsal body colour black and white; ive to seven dorsal markings (spots, patches and stripes); ears
subquadrangular; inner upper incisors unicuspid; forearm length 45–48 mm ....................................
..............................................................................................Glauconycteris superba Hayman, 1939
– Dorsal body colour not black and white; ears rounded ................................................................... 89
89. No body pattern; dorsal body colour umber-, sepia-, or reddish-brown; dorsal hairs tricoloured; inner
upper incisors strongly bicuspid, with unequal cusps; forearm length 34–38 mm .............................
..................................................................................Glauconycteris curryae Eger & Schlitter, 2001
– Markings restricted to lank stripes or shoulder spots; dorsal hairs bi- or tricoloured; inner upper
incisors generally bicuspid (except for argentata), with subequal or unequal cusps; forearm length
44 mm or less ................................................................................................................................... 90
90. Wing and interfemoral membranes pale brown; dorsal hairs pale golden-fawn or darker and
greyer ................................................................................Glauconycteris argentata (Dobson, 1875)
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– Wing and interfemoral membranes dark brown; dorsal hairs medium to dark greyish or dark sepia
brown ............................................................................................................................................... 91
91. Inner upper incisors strongly bicuspid, with subequal or unequal cusps; dorsal hairs tricoloured; no
body patterns or at most a shoulder spot ......................................................................................... 92
– Inner upper incisors weakly bicuspid, with unequal cusps; dorsal hairs bicoloured; body pattern
includes lank stripe and/or shoulder spot ....................................................................................... 93
92. No light lateral markings on body (perhaps rarely a white shoulder spot); forehead region of skull
weakly concave ....................................................................... Glauconycteris beatrix Thomas, 1901
– White shoulder spot is generally present; forehead region of skull moderately concave....................
........................................................................................Glauconycteris humeralis J.A. Allen, 1917
93. Dorsal hairs sepia brown; wings not reticulated ...........Glauconycteris alboguttata J.A. Allen, 1917
– Dorsal hairs medium to dark greyish or brownish; wings faintly reticulated......................................
.................................................................................................. Glauconycteris poensis (Gray, 1842)
94. Wings pale or white translucent; mid-ventral pelage cream, whitish or pale grey .......................... 98
– Wings dark; mid-ventral pelage whitish or pale greyish-brown to medium brown ........................ 95
95. Mid-ventral pelage white, whitish, cream or buff; forehead region of skull straight to weakly concave;
inner upper incisors weakly bicuspid (unicuspid when worn); forearm length 28–38 mm; occipital
helmet usually well developed............................................... Neoromicia capensis (A. Smith, 1829)
– Mid-ventral pelage pale (greyish-)brown to (medium-)brown; forehead region of skull weakly to
moderately concave (exceptionally almost straight); occipital helmet absent ................................ 96
96. Size large; forearm length 33–38 mm; greatest length of skull 13.0–14.1 mm; dorsal hairs unicoloured;
mid-ventral pelage greyish-brown to medium brown; inner upper incisors unicuspid; no anterior
upper premolar; forehead region of skull weakly concave ......Neoromicia brunnea (Thomas, 1880)
– Size smaller; forearm length less than 33 mm; dorsal hairs not unicoloured; inner upper incisors
usually unicuspid; anterior upper premolar extremely rarely present, in which case minute ......... 97
97. Dorsal and ventral hairs faintly bicoloured; mid-ventral colour pale brown or brown; forehead region
of skull weakly concave to almost straight; forearm length 22–32 mm ..............................................
................................................................................................Neoromicia somalica (Thomas, 1901)
– Dorsal hairs bi- or tricoloured; mid-ventral colour pale greyish-brown or brown; forehead region of skull
weakly to moderately concave; forearm length 27–33 mm....Neoromicia zuluensis (Roberts, 1924)
98. Dorsal colour dark brown, in strong contrast to pale wings; dorsal hairs usually unicoloured; ventral
hairs bicoloured; inner upper incisors weakly bicuspid; forearm length 28–33 mm ..........................
............................................................................................... Neoromicia tenuipinnis (Peters, 1872)
– Dorsal colour light brown, not in strong contrast to pale wings; dorsal hairs bi- or tricoloured; ventral
hairs bicoloured; inner upper incisors unicuspid; forearm length 29–38 mm .....................................
..................................................................................................Neoromicia rendalli (Thomas, 1889)
99. Outer upper incisor greatly reduced; entire underside pure white; dorsal pelage grey to sepia brown
[lower molars nyctalodont; forearm length 29–37 mm; anterior upper premolar medium sized,
conspicuous, within toothrow or slightly displaced; dorsal and ventral hairs bicoloured] .................
.................................................................................................Pipistrellus rueppellii (Fischer, 1829)
– Outer upper incisor not greatly reduced; underside light, dark or grey-tipped.............................. 100
121
European Journal of Taxonomy 382: 1–327 (2017)
100. Inner upper incisors unicuspid; both dorsal and ventral hairs bicoloured; dorsal pelage greyishbrown, reddish-brown, dark brown or blackish; anterior upper premolar medium-size to minute,
usually visible, within toothrow or fully displaced lingually; no white border on hind border of
wing; forearm length 29–38 mm ................................... Pipistrellus hesperidus (Temminck, 1840)
–
Inner upper incisors bicuspid ....................................................................................................... 101
101. Dorsal and ventral hairs bicoloured ............................................................................................. 102
–
Dorsal hairs unicoloured; ventral hairs usually unicoloured ....................................................... 105
102. Outer edge of tragus angular (hatchet-shaped); dorsal pelage chocolate brown or sepia brown,
sometimes with gold sheen; anterior upper premolar medium-sized, slightly displaced lingually;
no white border on hind border of wing; forearm length 25–35 mm ................................................
....................................................................................................... Neoromicia nana (Peters, 1852)
–
Outer edge of tragus curved (no sharp angle).............................................................................. 103
103. Skull larger, very weakly concave, greatest length 13.7–14.1 mm; wings without white hind border;
forearm length 32–36 mm; mid-ventral pelage slightly reddish-brown tinged with beige; anterior
upper premolar minute to medium-sized, usually visible, occasionally absent ................................
...................................................................................................... Hypsugo eisentrauti (Hill, 1968)
–
Skull smaller, weakly concave, greatest length less than 13.7 mm; wings with or without white hind
border ........................................................................................................................................... 104
104. Mid-ventral pelage pale brown, cream or white; anterior upper premolar minute, not visible above
gum; greatest length of skull 11.2–13.7 mm; forearm length 29–34 mm .........................................
................................................................................................... Hypsugo anchietae (Seabra, 1900)
–
Mid-ventral pelage dark to medium rust-brown, sometimes speckled; anterior upper premolar
absent; greatest length of skull 10.7–11.8 mm; forearm length 23–35 mm ......................................
............................................................................................Neoromicia guineensis (Bocage, 1889)
105. Size larger, forearm 33–38 mm; greatest length of skull 13.8–14.7 mm; mid-ventral pelage pale
brown, usually yellowish; anterior upper premolar minute, sometimes not visible, sometimes
missing; forehead region of skull weakly concave, sometimes almost straight ................................
........................................................................................... Pipistrellus grandidieri (Dobson, 1876)
–
Size smaller, forearm 33 mm or less; greatest length of skull 13.6 mm or less .......................... 106
106. Anterior upper premolar medium-sized to well-developed, visible, in toothrow or slightly lingually
displaced; dorsal pelage reddish-brown; white border on wing edge usually missing; forearm length
24–29 mm ................................................................................. Pipistrellus nanulus Thomas, 1904
–
Anterior upper premolar minute, barely visible above gum or not at all .................................... 107
107. Size larger, forearm length 28–33 mm; greatest length of skull 12.1–13.6 mm; mid-ventral pelage
pale to dark brown or reddish-brown........................................Hypsugo crassulus (Thomas, 1904)
–
Size smaller, forearm length 23–26 mm; greatest length of skull 10.3–11.4 mm; mid-ventral pelage
dark umber-brown .................................................................Hypsugo musciculus (Thomas, 1913)
108. A prominent vertical projection on the zygomatic arch; ears very large, 38–40 mm, conjoined on
extended snout; size very large, forearm length 62–73 mm; conspicuous bicolored dorsal pelage
pattern, with pale collar separating dark anterior and posterior regions of body ..............................
.......................................................................................... Otomops martiensseni (Matschie, 1897)
–
Not combining these characters ................................................................................................... 109
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109. Ears widely separated; ears and wing membranes translucent or whitish; underside of body entirely
white; 4 post-canine upper cheek teeth, M3 much reduced; one lower incisor on each side, with
concave crowns lengthened antero-posteriorly; upper incisors in contact with canines; no palatal
emargination (genus Myopterus) ..................................................................................................113
–
Ears not widely separated, sometimes conjoined, never translucent; 4 or 5 post-canine upper cheek
teeth (irst upper premolar often minute, sometimes deciduous); one to three (generally two) lower
incisors; upper incisors not in contact with canines .....................................................................110
110. Inner margins of ears meeting or almost meeting to form a V-shaped valley; third ridge of posterior
upper molar (M3) more than half length of second or equal in length; two lower incisors on each
side; anterior palatal emargination wide (genus Tadarida) ......................................................... 133
–
Inner margins of ears generally joined by band of skin (uncertain in niangarae, where they might
be well separated); anterior palatal emargination narrow or closed .............................................111
111. Third ridge of posterior upper molar (M3) more than half length of second or equal in length (genus
Chaerephon) .................................................................................................................................114
–
Third ridge of posterior upper molar (M3) less than half length of second or absent (sometimes
equal in condylurus) (genus Mops)...............................................................................................112
112. Anterior palatal emargination closed (subgenus Mops) .............................................................. 124
–
Anterior palatal emargination narrow (subgenus Xiphonycteris) ................................................ 130
113. Size smaller, forearm length 33–37 mm; skull length 17–19 mm; pelage dark brown above, without
lateral bands ...............................................................................Myopterus whitleyi (Scharff, 1900)
–
Size larger, forearm length 48–56 mm; skull length 21.3–25.7 mm; dorsal pelage slaty-grey with
two contrasting lateral bands of dirty white .................... Myopterus daubentonii Desmarest, 1820
114. Ears separated by a lobe or lappet ................................................................................................115
–
Ears joined by a band....................................................................................................................116
115. Ears long, separated by a large bulbous lobe projecting well over the forehead; ventral lank stripe
absent; wing membrane greyish-black; basisphenoid pits deep and large; rostrum uniquely with
prominent nasal swellings; forearm length 38 mm ......... Chaerephon gallagheri (Harrison, 1975)
–
Ears separated by a lat lappet over a small, backward-opening pocket; white or whitish ventral
lank stripe present; wing membrane dark brown to blackish; depth of basisphenoid pits shallow to
moderate, size small to medium; forearm length 39–46 mm ...........................................................
..............................................................................................Chaerephon major (Trouessart, 1897)
116. Size smaller; forearm length less than or equal to 41 mm............................................................117
–
Size larger; forearm length larger than 41 mm .............................................................................119
117. Dorsal pelage pale grey, pale rusty-brown or medium greyish-brown; wing membrane
usually white, sometimes pale brown or greyish-brown; ♂♂ with long bicoloured interaural
crest ....................................................................................... Chaerephon chapini J.A. Allen, 1917
–
Dorsal pelage almost black, brown, greyish-brown or reddish-brown; wing membrane white or
blackish-brown; ♂♂ with short unicoloured interaural crest .......................................................118
118. Size smaller; forearm length 32.8–37.8 mm; condylobasal length 14.4–15.9 mm; zygomatic width
9.3–10.3 mm .......................................................... Chaerephon leucogaster (A. Grandidier, 1869)
–
Size larger; forearm length 38.3–41.0 mm; condylobasal length 15.3–16.4 mm; zygomatic width
10.4–10.9 mm ................................................................ Chaerephon pumilus (Cretzschmar, 1826)
123
European Journal of Taxonomy 382: 1–327 (2017)
119. Ventral lank stripe present .......................................................................................................... 120
–
Ventral lank stripe absent............................................................................................................ 121
120. Flank stripe whitish; wing membranes pale grey; shallow basisphenoid pits of medium size;
anterior palatal emargination narrow; forearm length 41–48 mm; uniquely with pair of tail-glands
opening ventrally as a slit on each side of base of tail.......Chaerephon bemmeleni (Jentink, 1879)
–
Flank stripe white; wing membranes dorsally whitish or blackish, ventrally white; shallow to
medium basisphenoid pits of small to medium size; anterior palatal emargination closed; forearm
length 41–51 mm; no such glands ...........................................Chaerephon nigeriae Thomas, 1913
121. Anterior palatal emargination closed; size of basisphenoid pits large......................................... 122
–
Anterior palatal emarganation narrow; size of basisphenoid pits small to medium .................... 123
122. Size larger; forearm length 48–53 mm; basisphenoid pits deep; wing membrane blackish-brown;
sometimes with greyish spots dorsally ........................Chaerephon aloysiisabaudiae (Festa, 1907)
–
Size smaller; forearm length 42–46 mm; basisphenoid pits moderately deep; wing membrane
mostly dark brown, lighter towards tip; no spots or stripes. Chaerephon russatus J.A. Allen, 1917
123. Wing membrane light reddish-brown or light grey; forearm length 43–48 mm; dorsal pelage
somewhat frosted, occasionally with white spots ................ Chaerephon ansorgei (Thomas, 1913)
–
Wing membrane dark brown or light reddish-brown; forearm length 46–51 mm; white or whitish
stripes and/or rows of spots on crown of most individuals, and sometimes on shoulders and
lanks ................................................................................... Chaerephon bivittatus (Heuglin, 1861)
124. Size smaller; forearm length 51 mm or less ................................................................................ 125
–
Size larger; forearm length 51 mm or more................................................................................. 127
125. ♂♂ with paired glands between penis and anus; forearm length 41–46 mm; phalanges of 3rd (16–
19 mm) and 4th (13–16 mm) ingers short; ventral pelage white or very pale; crown darker than
back; whitish or pale grey lank stripe present, but faint; basisphenoid pits deep and large .............
.................................................................................... Mops (Mops) demonstrator (Thomas, 1903)
–
♂♂ without paired glands between penis and anus; phalanges of 3rd and 4th ingers medium to
long .............................................................................................................................................. 126
126. Forearm length 45–51 mm; phalanges of 3rd (20–24 mm) and 4th (17–20 mm) ingers long; ventral
pelage with little or no white; crown same colour as back; white ventral lank stripe present or
absent; basisphenoid pits shallow and small ................ Mops (Mops) condylurus (A. Smith, 1833)
–
Forearm length 44–48 mm; phalanges of 3rd (19–21 mm) and 4th (15–17 mm) ingers of medium
length; ventral pelage predominantly white or cream; crown darker than back; ventral lank stripe
absent; basisphenoid pits moderately deep and medium to large in size...........................................
..........................................................................Mops (Mops) niveiventer Cabrera & Ruxton, 1926
127. Size largest; forearm length 59–67 mm; white or pale ventral lank stripe present; dorsal pelage
colour variable; frosted and lecked; basisphenoid pits moderately deep and medium to large in
size ....................................................................................... Mops (Mops) midas (Sundevall, 1843)
–
Size smaller; forearm 58 mm or less ........................................................................................... 128
128. Inner margin of ears uncertain, possibly well separated; ventral lank stripe probably absent; wing
membrane dark brown; forearm length approximately 52 mm .........................................................
.........................................................................................Mops (Mops) niangarae J.A. Allen, 1917
–
Inner margin of ears joined by a band ......................................................................................... 129
124
VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
129. Forearm length 51–55 mm; wing membrane dark brown; dorsal pelage medium sepia or yellowishbrown, or pale rusty-brown....................................................Mops (Mops) trevori J.A. Allen, 1917
–
Forearm length 54–58 mm; wing membrane blackish-brown; dorsal pelage dark brown to almost
black; not frosted or lecked ..............................................Mops (Mops) congicus J.A. Allen, 1917
130. Size smaller, forearm length 31 mm or less................................................................................. 131
–
Size larger, forearm length 34 mm or more ................................................................................. 132
131. Two lower incisors on each side; lower canines without greatly enlarged cingula; upper incisors
procumbent (inclined towards the lips); wing membrane blackish-brown, pale brown to whitish,
ventral side paler; forearm length 27–31 mm ....... Mops (Xiphonycteris) nanulus J.A. Allen, 1917
–
Usually one lower incisor on each side, rarely two; lower canines with greatly enlarged cingula,
especially in ♂♂; wing membrane blackish; forearm length 27–30 mm ..........................................
............................................................................. Mops (Xiphonycteris) spurrelli (Dollman, 1911)
132. Ventral lank stripe present, but contrast weak; wing membrane blackish; ventral pelage dark;
occipital helmet prominent; wing insertion higher on body; basisphenoid pits of moderately depth
and small to medium in size; forearm length 35–42 mm ……………………………………………………………
……………………………………………………………………Mops (Xiphonycteris) brachypterus (Peters, 1852)
–
Ventral lank stripe absent; wing membrane various shades of brown; ventral pelage pale; occipital
helmet moderately prominent; wing insertion lower on body; basisphenoid pits moderate or shallow
deep and medium to large in size; forearm length 34–41 mm...........................................................
.............................................................................. Mops (Xiphonycteris) thersites (Thomas, 1903)
133. Little white on ventral surface; forearm length 60–67 mm; both sides of wing membrane dark
brown to almost black; basisphenoid pits medium in size; plantar pad absent .................................
................................................................................................... Tadarida ventralis (Heuglin, 1861)
–
Clear white, cream or yellowish ventral lank band present; forearm length 56–61 mm; dorsal
side of wing membrane brown, ventral side whitish; basisphenoid pits large in size; plantar pad
present ..................................................................................... Tadarida fulminans (Thomas, 1903)
125
European Journal of Taxonomy 382: 1–327 (2017)
Catalogue of voucher specimens
This list gives an overview of all of the voucher specimens examined by the various contributors or for
which information was found in the literature.
For the museum acronyms, see page 246.
For all of the specimens the following information is provided (as far as it was available):
Taxon name;
Country;
Locality;
The number of individuals (inds) of the species that were collected at that locality, with details on their
sex (♂(♂): male(s), ♀(♀): female(s), ?(?): unknown), age (ad = adult; emb = embryo; imm = immature;
juv = juvenile; sad = subadult), and conservation type (AC = alcohol and cranium; AL = alcohol, skull
not removed; AS = alcohol, skull missing; MS = mounted skin and skull; OT = other, undeined type;
SA = skull and alcohol; SB = skin, skeleton and skull; SC = skin and cranium; SI = skull in skin;
SK = skull only; SN = complete skeleton; SO = skin only; SS = skin and skull), UN = unknown;
Museum acronym and registration number(s) in alphabetical and/or numeric order. If more than two
consecutive number are present, the irst and the last are connected by “to”;
The collector(s) (leg. = legit) and the dates on which the specimens were collected. A question mark in
the date ield indicates that the specimens were collected on multiple dates of which some could not be
determined.
Chiroptera Blumenbach, 1779
BURUNDI: 2 ?? [2 UN], Kinazi, between 5 Jan. 1953 and 8 Jan. 1953, leg. P. Basilewsky (RMCA:
21939, 21940).
DEMOCRATIC REPUBLIC OF THE CONGO: 59 ?? [1 AL, 58 UN], unknown locality, unknown date,
leg. Prescott Musaba, unknown date, leg. unknown collector (RMCA: a1.097-M-3221, UNIKIS: CH1085
to CH1088, CH1090 to CH1093, CH1095, CH1096, CH1098, CH2000 to CH2004, CH2006 to CH2017,
CH2019, CH2020, CH2022 to CH2028, CH2030 to CH2032, CHYO106 to CHYO116, CR09-828,
CR09-839, CR09-842, CR09-845, CR09-863, CRT813/G72, CRT815/GE67); 4 ?? [4 UN], AlbertvilleMoba (road?) [= Kalemie-Moba (road?)], 19 Nov. 1953, 26 Feb. 1954, between 1 Apr. 1954 and
30 Apr. 1954, leg. H. Bomans (RMCA: 23317 to 23319, 23321); 1 ♀ [UN], Avakubi, 7 Jan. 1912, leg.
Cuthbert Christy (RMCA: 1606a); 1 ? [UN], Bambesa, prior to 28 Jun. 1937, leg. J.M. Vrijdagh (RMCA:
13877); 2 ?? [2 UN], Banana, 12 Apr. 1952, leg. Is. Mesmaekers (RMCA: 20761, 20762); 7 ?? [7 UN],
Beno, prior to 3 Jan. 1950, between 1 Jan. 1949 and 31 Dec. 1949, leg. P. Bayet (RMCA: 19834 to 19837,
19854, 19855, 19474); 2 ♀♀, 1 ♂ [3 SS], Bitale, 12 Jun. 1970, 14 Jun. 1970, 21 Jun. 1970, leg. Alexandre
Prigogine (RMCA: 36231, 36230, 36233); 1 ? [UN], Boende, between 1 Jan. 1966 and 31 Jan. 1966,
leg. P. Lootens (RMCA: 34161); 1 ? [UN], Bokuma, prior to 5 Mar. 1953, leg. P. Lootens (RMCA:
21675); 1 ♂ [SO], Bokungu, 30 Jun. 1948, leg. L. Dupuis (RMCA: 18641); 1 ♀ [SO], Bolobo, prior to
18 Jan. 1958, leg. Ngwe (RMCA: 26912); 2 ?? [2 UN], Boma, between 1 Feb. 1955 and 28 Feb. 1955,
leg. Is. Mesmaekers (RMCA: 23310, 23311); 1 ? [UN], Bosobolo, prior to 23 Mar. 1951, leg. Vachaudez
(RMCA: 20343); 1 ? [SK], Bukavu, between 1 Jan. 1952 and 31 Dec. 1952, leg. P.G. Vercammen
(RMCA: 82.001-M-0020); 1 ? [UN], Buta, prior to 9 Nov. 1938, leg. Jozef Hutsebaut (RMCA: 14904);
2 ♂♂ [2 SS], Byonga, 4 Jan. 1972, 16 Jan. 1973, leg. Alexandre Prigogine (RMCA: 38623, 38589);
1 ? [SS], Dilolo, 2 Feb. 1948, leg. Jozef Hutsebaut (RMCA: 18230); 1 ? [UN], Funda Biabo, between
1 Feb. 1914 and 28 Feb. 1914, leg. Charliers (RMCA: 5396); 5 ?? [5 UN], Ibembo, 20 Jul. 1950,
between 1 Nov. 1949 and 30 Nov. 1949, leg. Jozef Hutsebaut (RMCA: 20093, 19858 to 19861); 1 ♂
[SS], Idjwi, Island, 16 Sep. 1969, leg. Alexandre Prigogine (RMCA: 36219); 1 ? [UN], Irumu, between
126
VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
1 Nov. 1951 and 30 Nov. 1951, leg. Alexandre Fain (RMCA: 21463); 1 ♂ [SS], Kakanda Cave,
11 Sep. 1971, leg. Alexandre Prigogine (RMCA: 38566); 1 juv ♀, 1 ♂ [2 SS], Kamituga, 6 Apr. 1973,
26 Apr. 1973, leg. Alexandre Prigogine (RMCA: 38626, 38624); 1 ♀ [SS], Kamusimbi, 11 Feb. 1973,
leg. Alexandre Prigogine (RMCA: 38625); 4 ♀♀, 4 ♂♂ [8 SS], Kasebere, 14 Feb. 1970, 18 Jun. 1970,
25 Jun. 1970, 1 Dec. 1970, 2 Dec. 1970, 3 Dec. 1970, 24 Apr. 1971, leg. Alexandre Prigogine (RMCA:
38572, 36222, 36223, 36245, 38563, 38587, 36244, 38569); 1 ad ♂, 1 ? [1 AL, 1 UN], Kibumba,
10 Aug. 1970, leg. M.J. Celis and M. Lejeune (RMCA: 36209, 36210); 1 ? [UN], Kibwe Lya Mikako
Cave, 28 Dec. 1966, leg. M.J. Celis and M. Lejeune (RMCA: 35172); 2 ?? [1 AL, 1 UN], Kikwit,
unknown date, leg. Prescott Musaba, Moïse Bipoo, Richard Tamaru, Pauline Isude and Joseph Kussa,
between 10 Jun. 1995 and 28 Aug. 1995, leg. Herwig Leirs et al. (UNIKIS: KKT2845, RMCA: 97.021M-1175); 1 ? [UN], Kilo Mines [= Kilomines], prior to 27 Jun. 1913, leg. Thelie (RMCA: 1867); 1 ♂
[SS], Kiloboze [= Kilobono], 12 Jan. 1973, leg. Alexandre Prigogine (RMCA: 38627); 3 ♂♂ [3 UN],
Léopoldville [= Kinshasa], 13 May 1912, 17 May 1912, leg. Cuthbert Christy (RMCA: 1539e, 1539f,
1539d); 6 ?? [6 UN], Lubondai-Tshimbulu [= Lubondaie-Tshimbulu], prior to 9 Dec. 1955, leg.
S. Marks (RMCA: 23238 to 23240, 23395 to 23397); 3 ?? [3 AL], Lukolela, 1 Feb. 1931, leg. James Paul
Chapin (AMNH: 80958 to 80960); 1 juv ?, 1 ♂, 5 ?? [5 SK, 2 SO], Luluabourg [= Kananga], unknown
date, 16 Nov. 1965, 28 Aug. 1969, leg. Antoon Emeric Marcel De Roo (RMCA: 34416 to 34420, 33982,
38815); 1 ad ♀, 1 ad ♂, 5 ♀♀, 9 ♂♂ [14 SS, 2 SK], Maboya [= Maboyo], 8 Jan. 1970, 12 Oct. 1970,
16 Nov. 1970, 1 Dec. 1970, 8 Dec. 1970, 25 Dec. 1970, 9 Jan. 1971, 21 Jan. 1971, 22 Jan. 1971,
23 Jan. 1971, 31 Jan. 1971, 2 Feb. 1971, 14 Feb. 1971, 10 Mar. 1971, leg. Alexandre Prigogine (RMCA:
38573, 38562, 36247, 38564, 38580, 36246, 38570, 38582, 38576, 38581, 38565, 38575, 38578, 36221,
38571, 38579); 1 ? [UN], Mahagi Port, 29 Mar. 1953, leg. Célestin Hecq (RMCA: 21899); 2 ♂♂
[2 SS], Masibingi, 7 Mar. 1971, 20 Mar. 1971, leg. Alexandre Prigogine (RMCA: 38568, 38577); 2 ??
[2 UN], Moba, 19 Nov. 1953, leg. H. Bomans (RMCA: 22122, 22123); 2 ♀♀ [2 SS], Mule, 4 May 1970,
12 May 1970, leg. Alexandre Prigogine (RMCA: 36229, 36232); 1 ♀ [SS], Mwenga, 29 Nov. 1971, leg.
Alexandre Prigogine (RMCA: 38588); 2 ♂♂ [2 SS], Ndomo, 16 Mar. 1971, 18 Mar. 1971, leg. Alexandre
Prigogine (RMCA: 38574, 38567); 4 ♂♂ [4 SS], Ngula, 1 Mar. 1977, 8 Mar. 1977, 12 Mar. 1977, leg.
Tshinyenye (RMCA: 77.021-M-0019 to 77.021-M-0022); 1 ? [UN], Niambasha, prior to 25 Aug. 1959,
leg. Uwe Rahm (RMCA: 28034b); 1 ? [UN], Pawa, prior to 23 Jan. 1939, leg. Degotte (RMCA: 14989);
1 ? [UN], Popokabaka, between 1 Nov. 1955 and 30 Nov. 1955, leg. René Close (RMCA: 23390); 1 ♀,
1 ♂ [2 UN], Rubi-Tele, 19 Aug. 2013, leg. Guy-Crispin Gembu Tungaluna, Prescott Musaba and André
Malekani (UNIKIS: RBTL510, RBTL512); 1 ? [UN], Sombe, between 1 Jan. 1948 and 31 Dec. 1948,
leg. Dewit (RMCA: 19074); 1 ♂ [SO], St. Jos. (Mission) [= Mikalaya], 9 Sep. 1924, leg. Richard
Callewaert (AMNH: 86139); 10 ♀♀, 10 ♂♂, 2 ?? [19 SK, 3 UN], St. Joseph de Luluabourg Mission
[= Mikalaya], 15 Feb. 1923, 17 Feb. 1923, 19 Feb. 1923, 26 Feb. 1923, 11 Mar. 1923, 12 Mar. 1923,
27 Mar. 1923, 5 Apr. 1923, 11 Apr. 1923, 5 May 1923, 8 May 1923, 19 May 1923, 22 May 1923,
31 May 1923, 1 Jun. 1923, 2 Jun. 1923, 19 Jun. 1923, 27 Jun. 1923, 28 Jun. 1923, 15 Nov. 1923, leg.
Richard Callewaert (RMCA: 7177, 7118, 7114, 7183, 7180, 7107, 7173, 7119, 7185, 7176, 7175, 7178,
7120, 7179, 7181, 7170, 7174, 7113, 7184, 7117, 7172, 7171); 1 ♂ [UN], Stanleyville [= Kisangani],
2 Jul. 1912, leg. Cuthbert Christy (RMCA: 1419); 1 ? [UN], Tshuapa district, between 1 Jan. 1960
and 31 Dec. 1964, leg. P. Lootens (RMCA: 34170); 3 ?? [3 UN], Yoko, unknown date, leg. unknown
collector (UNIKIS: LK01, LK03, LK04).
RWANDA: 1 ♀ [SS], Astrida [= Butare], 24 Dec. 1961, leg. unknown collector (RMCA: 38816).
127
European Journal of Taxonomy 382: 1–327 (2017)
Pteropodiformi Hutcheon & Kirsch, 2006
Pteropodidae Gray, 1821
Eidolinae Almeida, Giannini & Simmons, 2016
Eidolon Rainesque, 1815
Eidolon helvum (Kerr, 1792)
BURUNDI: 8 ♀♀ [4 AL, 4 SB], Buhoro, 21 Dec. 1970, leg. P.L. Niort (MHNG: 1498.003 to 1498.010);
5 ♂♂, 2 ?? [2 SS, 2 AL, 1 UN, 2 SB], Bujumbura, 1 Aug. 1976, leg. Jacques Verschuren, 3 Jun. 1971,
4 Jun. 1971, 12 Jul. 1973, leg. P.L. Niort, between 1 Jul. 1982 and 31 Jul. 1982, leg. Vissens (RBINS:
19908, MHNG: 1498.031, 1498.032, 1498.011, 1498.012, RMCA: 82.023-M-0001, 82.023-M-0002);
3 ♂♂ [1 SA, 2 SK], Kayanza, 27 Mar. 1979, leg. Heinz Stephan (SMF: 57500 to 57502); 18 ♂♂
[18 SB], Mabaye [= Mabayi], 10 Dec. 1971, leg. P.L. Niort (MHNG: 1498.013 to 1498.030).
DEMOCRATIC REPUBLIC OF THE CONGO: 2 ♀♀, 4 ♂♂, 5 ?? [6 SS, 2 AL, 2 SO, 1 UN], unknown
locality, unknown date, prior to 31 Dec. 1930, leg. unknown collector, prior to 17 Mar. 1953, leg. Leonet,
prior to 28 Dec. 1955, leg. P. Dyleff, prior to 8 Dec. 1939, leg. Schwetz (RBINS: 181E, 6278, RMCA:
21690, 23402, 23403, RBINS: 9225 to 9228, RMCA: 15774, 15775); 8 ad ♀♀, 22 ad ♂♂, 6 juv ♀♀,
6 juv ♂♂, 3 ♀♀, 3 ♂♂, 3 ?? [46 SS, 5 AL], Avakubi, 20 Nov. 1909, 9 Sep. 1913, 1 Oct. 1913, 14 May
1914, 16 May 1914, 5 Jun. 1914, leg. Herbert Lang, James Paul Chapin and The American Museum
Congo Expedition, 1 Oct. 1913, leg. J.A. Howard (AMNH: 48704, KU: 125194, AMNH: 48705, 48706,
48659, 48660, 48663 to 48667, 48669 to 48676, 48678 to 48685, 48687 to 48697, 50363, 51383, 51384,
RMCA: 12371, 12372, 12375, AMNH: 48707 to 48711, 48702, 48703); 3 ♂♂ [1 SS, 2 SO], Boende,
23 May 1939, 4 Jun. 1939, leg. R.P. Wynants, 6 Aug. 1947, leg. Huypens (RMCA: 15868, 15867,
18236); 2 ♂♂ [1 SS, 1 SO], Bokalakala, 8 May 1959, 13 Oct. 1959, leg. N’Kele (RMCA: 28541,
27946); 2 ♀♀, 1 ♂ [3 SS], Bokuma, 16 Oct. 1950, leg. P. Lootens (RMCA: 20394 to 20396); 1 ? [SS],
Bokungu, prior to 25 Jul. 1949, leg. L. Dupuis (RMCA: 18816); 1 juv ?, 2 ♀♀, 1 ? [3 SS, 1 SO],
Bolobo, unknown date, prior to 20 Aug. 1958, leg. N’Kele (RMCA: 27416, 27276 to 27278); 1 ? [AL],
Boma, between 1 Aug. 1928 and 31 Aug. 1928, leg. Gerard (RMCA: 16184); 1 ♀ [UN], Bomane
[= Bomaneh], 24 May 2010, leg. Guy-Crispin Gembu Tungaluna, Prescott Musaba and José Akaibe
(UNIKIS: CRT2279); 3 ?? [3 SO], Bukavu, 16 Aug. 1952, leg. P.G. Vercammen (RMCA: 23174 to
23176); 13 ?? [9 SS, 4 SO], Buta, 14 Aug. 1938, 20 Jun. 1949, prior to 9 Jan. 1937, prior to 28 Dec. 1938,
prior to 10 Feb. 1930, leg. Jozef Hutsebaut (RMCA: 14943, 18838 to 18841, 10127 to 10131, 14944,
14945, 13697); 6 ♀♀, 10 ♂♂, 17 ?? [11 SS, 18 AL, 3 SO, 1 UN], Butembo, 31 Jan. 1956, 1 Feb. 1956,
11 Feb. 1956, 3 May 1956, 22 Jul. 1958, 29 Jul. 1958, 27 Aug. 1958, prior to 28 Dec. 1955, prior to
26 Jan. 1955, prior to 25 Jan. 1956, prior to 23 Oct. 1959, prior to 17 Apr. 1956, between 1 Aug. 1955
and 31 Aug. 1955, between 1 Jul. 1955 and 31 Jul. 1955, between 1 Apr. 1957 and 30 Apr. 1957, between
1 Jan. 1956 and 31 Mar. 1956, leg. P. Dyleff (RMCA: 23554, 23555, 23677 to 23681, 23681b, 27494,
27488, 27480, 23739, 23551, 23552, 23682 to 23689, 26421, 26422, 23313, 23135, 23556 to 23558,
28321, 23448, 22812, 23404); 1 ? [SS], Dilolo, between 1 Jan. 1932 and 31 Dec. 1932, leg. Delvaux
(RMCA: 12024); 4 ♂♂ [4 SS], Dimbelenge, 23 Jun. 1980, 27 Jun. 1980, 28 Jun. 1980, leg. Betu
Nsankulu (RMCA: 80.042-M-0001 to 80.042-M-0004); 1 ♂ [SS], Eala, 2 Jan. 1921, leg. Henri
Schouteden (RMCA: 6523); 10 ♀♀, 2 ♂♂ [12 AL], Eidolon Land, 4 Oct. 2009, leg. Alexandre
Hassanin, Nicolas Nesi, Célestin Pongombo, Peter Vallo and Didier Tshikung (MNHN: ZM-2011-714
to ZM-2011-725); 1 ad+foetus ♀ [AL], Equateur Province, between 1 Jan. 1921 and 31 Dec. 1921, leg.
Verlaine (RMCA: 38240); 1 ? [UN], Franz-Josef Falls, unknown date, leg. unknown collector (RMCA:
15074); 1 ♀ [UN], Goma, between 1 Dec. 1987 and 31 Dec. 1987, leg. J. Schoorl (ZMA: MAM.24139);
2 ad+foetus ♀♀, 3 ♀♀, 1 ♂, 4 ?? [6 SS, 2 SA, 1 SK, 1 UN], Ibembo, unknown date, 13 Aug. 1919,
30 Aug. 1919, 16 Sep. 1919, 22 Sep. 1919, leg. Jean Lebrun, 20 Feb. 1950, 21 Feb. 1950, 21 Dec. 1951,
leg. Jozef Hutsebaut, prior to 17 Jul. 1924, leg. J. Rodhain (RMCA: 4941a, 4925, 4919, 19700 to 19702,
20660, 4936, 4930, 7641); 1 ♂ [SS], Ikengo, 16 Feb. 1921, leg. Henri Schouteden (RMCA: 6512); 1 ♀,
128
VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
1 ♂ [2 SS], Inkongo [= Inkongo-Kakese], 27 Jan. 1923, leg. Wilson (RMCA: 6307, 6308); 1 ? [SN],
Irebu, unknown date, leg. J. Camp (USNM: A49408); 1 imm ♂, 1 ♀, 1 ♂ [2 SK, 1 UN], Isato,
23 Dec. 1947, leg. unknown collector, prior to 25 Nov. 1947, leg. A. Henrion (RBINS: 7716, 7717,
13110); 1 ♂ [SK], Ituri Forest, unknown date, leg. unknown collector (BMNH: 1907.1.2.8); 1 ? [SS],
Iyonda, 2 May 1952, leg. Karl Michahelles (RMCA: 21773); 1 ♀, 1 ♂ [1 SS, 1 SO], Kabambaie
[River], 16 Oct. 1921, leg. Henri Schouteden (RMCA: 6507, 6508); 1 ♂ [SS], Kabobo, Mount,
18 Feb. 1957, leg. Alexandre Prigogine (RMCA: 27113); 2 ♀♀, 1 ♂ [3 SS], Kakanda Cave, 22 Mar. 1966,
leg. Alexandre Prigogine (RMCA: 34247 to 34249); 1 ♀ [SS], Kakonda, 8 Oct. 1942, leg. F.L. Hendricks
(AMNH: 118868); 1 ♂ [SS], Kamituga, 17 Mar. 1965, leg. Alexandre Prigogine (RMCA: 33295); 1 ♂
[SS], Kasende, 6 Jul. 1934, leg. R.S. Pierrepont (MCZ: 31737); 1 ♂ [SS], Katana, 18 Mar. 1956, leg.
James Paul Chapin (AMNH: 180892); 1 ? [AL], Katanga, prior to 31 Aug. 1900, leg. Charles Lemaire
(RMCA: 425b); 2 ♂♂ [2 SS], Keseki, prior to 23 Jul. 1946, leg. N’Kele (RMCA: 17399, 17401); 1 ♀,
3 ♂♂, 14 ?? [18 UN], Kikwit, unknown date, leg. unknown collector, 25 Jan. 1996, 28 Jan. 1996, leg.
Robert (Bob) Swanepoel (TM: 46061, 46062, 46065 to 46072, 46075, 46077 to 46079, 45592, 45593,
45629, 45594); 1 ♂ [UN], Kimvuadu, 19 Feb. 1939, leg. C. Vleeschouwers (RMCA: 16084); 1 ♀, 1 ♂
[1 SS, 1 SO], Kisanga, 27 Oct. 1948, leg. J. de Wilde (RBINS: 12778, 12779); 1 ♀, 1 ♂, 2 ?? [2 AL,
2 UN], Kisangani, 7 Sep. 2012, leg. Prescott Musaba, Tembele and Marc Boketshu, 9 Jun. 2014, leg.
Prescott Musaba, prior to 5 Aug. 1988, leg. Luc De Vos, Kimbembi and Alain Vandelanoote
(UNIKIS: YK037CH, K13-591, RMCA: 88.025-M-0002, 88.025-M-0003); 1 ? [SO], Kiseny [=
Kasenye], prior to 23 Jul. 1946, leg. R. Van Saceghem (RMCA: 17189); 8 ?? [8 SS], Kitundu Kinda,
22 Sep. 1914, leg. de Baillet Latour (RMCA: 4225, 4226, 4228, 4229, 4232 to 4235); 1 ♂ [SS], Kitutu,
14 Mar. 1960, leg. Alexandre Prigogine (RMCA: 28885); 1 ♀, 1 ♂ [2 SS], Kitwabaluzi, 19 Apr. 1951,
leg. Alexandre Prigogine (RMCA: 20689, 20690); 2 ad ?? [2 SS], Kivu [= Kivu region], 11 Sep. 1935,
leg. unknown collector (MNHN: ZM-MO-1939-198, ZM-MO-1939-199); 1 ♂ [SK], Kongolo Island,
between 1 Apr. 1982 and 30 Apr. 1982, leg. Marc Colyn (RMCA: 90.042-M-0271); 5 ♀♀, 1 ♂ [5 SS,
1 SO], Kuleponge [= Kulepongi], 26 Jun. 1938, leg. J.M. Vrijdagh (RMCA: 14791 to 14795, 15322);
49 ♀♀, 17 ♂♂, 2 ?? [62 SS, 1 SK, 4 SO, 1 UN], Kunungu, unknown date, prior to 6 Aug. 1925, prior
to 25 Mar. 1929, prior to 22 Oct. 1930, between 1 Jan. 1922 and 31 Dec. 1922, leg. Ngwe, 9 Apr. 1921,
leg. Henri Schouteden, prior to 22 Oct. 1930, leg. N’Kele (RMCA: 8176, 6484, 6485, 6487, 6491 to
6497, 6516 to 6519, 7741 to 7748, 7756 to 7766, 10562 to 10574, 10618, 10639 to 10642, 9706, 9712,
8152 to 8165); 3 ♀♀ [3 SS], Kwamouth, 24 May 1921, leg. Henri Schouteden (RMCA: 6488 to 6490);
2 ♀♀, 1 ♂ [3 SS], Lake Tumba [= Tumba, Lake], 15 Nov. 1954, leg. James Paul Chapin (AMNH:
180894 to 180896); 2 ♀♀ [2 AL], Lake Tumba, Island in, 14 Jun. 1956, 17 Jun. 1965, leg. Alvin
Novick (MCZ: 48181, 48182); 1 ? [SS], Lemba Plain, prior to 3 May 1930, leg. Van Delft (RMCA:
10262); 1 ♀ [SS], Leopold 2nd, Lake [= Mai-Ndombe, Lake], unknown date, leg. unknown collector
(BMNH: 1927.3.1.47); 2 ♀♀, 3 ♂♂, 2 ?? [2 SK, 3 SO, 2 UN], Léopoldville [= Kinshasa], unknown
date, prior to 25 Nov. 1947, leg. A. Henrion, 12 Nov. 1945, leg. Jean Jacques Deheyn, 16 Jun. 1946,
between 1 Jan. 1946 and 31 Dec. 1946, leg. M. Henrion, between 1 Feb. 1942 and 28 Feb. 1942, leg.
Becquet (RBINS: 12271, RMCA: 17616a, 17797, 17955, 17956, 17356a, RBINS: 12771); 5 ♀♀, 8 ♂♂
[12 SS, 1 SO], Luebo, 13 Dec. 1921, 15 Dec. 1921, leg. Henri Schouteden (RMCA: 6483a, 6483b, 6501
to 6506, 6513 to 6515, 6509, 6510); 1 ♀ [SS], Luira River, upper, unknown date, leg. S.A. Neave
(BMNH: 1909.1.3.1); 1 ♀, 3 ♂♂ [4 SS], Lukolela, 1 Dec. 1930, leg. James Paul Chapin, 13 Aug. 1930,
17 Aug. 1930, 18 Aug. 1930, leg. Franklin Edson (AMNH: 86879, 86758 to 86760); 3 ♂♂ [3 SS], Lulua
[= Kananga], 30 Apr. 1976, leg. Betu Nsankulu (RMCA: 76.054-M-0001 to 76.054-M-0003); 1 ♀,
2 ♂♂ [3 SS], Luluabourg [= Kananga], unknown date, leg. unknown collector, unknown date, leg.
Richard Callewaert (BMNH: 1926.11.1.23, 1926.7.6.45, 1926.7.6.46); 7 ♀♀, 4 ♂♂ [8 SS, 1 SK, 2 SO],
Lusanga Mwanza, prior to 4 Aug. 1937, leg. Dartevelle (RMCA: 13936 to 13943, 13944a, 13944b,
13945); 2 ♀♀, 2 ♂♂, 1 ? [1 SS, 3 UN, 1 SB], Lwiro [= Luiro], unknown date, 11 May 1956, between
1 Aug. 1955 and 31 Aug. 1955, leg. J.J. Laarman, 20 Apr. 1955, leg. James Paul Chapin, 29 Jul. 2002,
leg. Michael H. Huhndorf (RMNH: MAM.16356, MAM.16384, AMNH: 180893, FMNH: 173291,
129
European Journal of Taxonomy 382: 1–327 (2017)
RMNH: MAM.16357); 1 ♀ [AL], Mabali, between 1 May 1956 and 30 Jun. 1956, leg. Alvin Novick
(MCZ: 48183); 1 ♀ [SK], Masako, 24 Nov. 1987, leg. Ndey B. Ifuta (RMCA: 93.079-M-0279); 1 ad ♀,
3 juv ♀♀, 1 ? [1 SS, 4 UN], Medje, 12 May 1914, leg. Cuthbert Christy, 6 Jun. 1910, 20 May 1914,
22 May 1914, leg. Herbert Lang, James Paul Chapin and The American Museum Congo Expedition
(RMCA: 4330, AMNH: 48699 to 48701, 48698); 1 ad ♂ [SS], Mondombe, 26 Dec. 1952, leg. L. Dupuis
(RMCA: 21715); 2 ♀♀, 1 ♂ [3 SS], Mongende, 11 Apr. 1921, leg. Henri Schouteden (RMCA: 6520 to
6522); 1 ad ♂ [SS], Mouth of Congo, between 1 Jan. 1892 and 31 Jan. 1892, leg. J.M. Lewis (BMNH:
1892.5.7.1); 1 ? [AL], Mpe, prior to 26 Jan. 1914, leg. Maes (RMCA: 2663); 8 ?? [8 SS], Muezi
Lupungu, 1 Oct. 1914, leg. de Baillet Latour (RMCA: 4227, 4230, 4231, 4236a, 4236b, 4237 to 4239);
2 ?? [2 UN], Mulungu, unknown date, prior to 1 Feb. 1940, leg. F.L. Hendricks (RMCA: 15870, 15871);
1 ? [SS], Mulungu Miti [= Mulungu], 6 Jan. 1947, leg. F.L. Hendricks (RMCA: 19649); 1 imm ♀ [SS],
Ndama, 21 Jul. 1954, leg. C.F. de Wilde (RBINS: 17295); 2 ?? [2 AL], New Beni [= Beni], between
1 Oct. 1926 and 31 Oct. 1926, leg. James Paul Chapin, Dewitt L. Sage, Frank P. Mathews and The
Ruwenzori-Kivu Expedition (AMNH: 82360, 82361); 3 ♀♀, 3 ♂♂ [6 SS], Ngerere Lepi, 5 Jan. 1943,
6 Jan. 1943, leg. J. de Wilde (RBINS: 12773, 12774, 12776, 12777, 12772, 12775); 4 ?? [4 UN], NiamNiam, unknown date, leg. unknown collector, 17 Jun. 1884, leg. Bohndorff (RBINS: 181c, 181, 181b,
181j); 2 ♀♀, 1 ♂ [3 SO], Nyamunyunye, between 1 Jan. 1951 and 31 Dec. 1951, leg. F.L. Hendricks
(RMCA: 74.044-M-0004 to 74.044-M-0006); 16 ?? [13 AL, 3 UN], Oshwe, prior to 12 Mar. 1914, leg.
Maes (RMCA: 2773 to 2788); 3 ♂♂ [3 UN], Semmio [= Zemio], unknown date, leg. Bohndorff (ZMB:
10206, 7427, 7428); 5 ?? [5 AL], Shabunda, prior to 10 Jul. 1939, leg. Braun (RMCA: 15381 to 15385);
18 ♀♀, 6 ♂♂, 1 ? [23 SS, 1 SK, 1 SB], St. Jos. (Misson) [= Mikalaya], unknown date, 19 Jun. 1924,
25 Jun. 1924, 3 Nov. 1924, leg. Richard Callewaert (AMNH: 86260, 86229 to 86250, 86253, 86254);
2 ♀♀, 1 ♂ [2 SS, 1 UN], St. Joseph de Luluabourg Mission [= Mikalaya], 20 Jul. 1922, 3 Jun. 1923,
7 Jun. 1923, leg. Richard Callewaert (RMCA: 7494, 7123, 7116); 1 ♀ [SS], Tandala, 18 Jul. 1979, leg.
C. Brian Robbins (USNM: 537699); 1 ♂ [UN], TASOK (The American School of Kinshasa)
[= Kinshasa, 10 km W], 15 Apr. 1980, leg. V. Wallach (ZMA: MAM.21145); 1 ♀ [SS], Tondu,
23 Feb. 1921, leg. Henri Schouteden (RMCA: 6511); 4 ♀♀ [4 SS], Tshikapa, 26 Nov. 1921, leg. Henri
Schouteden (RMCA: 6486, 6498 to 6500); 1 ♂ [SS], Uele [River], prior to 18 Feb. 1922, leg. The
Antwerp Zoo (RMCA: 6065); 1 juv ?, 1 ♀, 4 ♂♂ [3 SS, 1 SO, 2 UN], Umangi [Mission], prior to
31 Jul. 1899, leg. Charles de la Kethulle de Ryhove (RMCA: 182 to 187); 1 ♀ [UN], Yaekela, 3 May
2010, leg. Guy-Crispin Gembu Tungaluna, Prescott Musaba and José Akaibe (UNIKIS: CRT2387); 1 ♂
[SS], Yangambi, 28 Dec. 1947, leg. unknown collector (RBINS: 6985); 1 ? [SS], Yokamba Elongo,
28 Jun. 1954, leg. Stevenart (RMCA: 22411); 4 ?? [4 SS], Zambi, unknown date, leg. R. Van Saceghem
(RMCA: 4861 to 4864).
RWANDA: 17 ?? [1 SS, 14 AL, 2 SB], Astrida [= Butare], 15 May 1962, leg. Van Parijs, 17 Apr. 1968,
leg. P.L. Niort, 20 Jan. 1951, leg. Aureliaan (RMCA: 31176 to 31189, MHNG: 1498.001, 1498.002,
RMCA: 20491); 1 ad ♀, 5 ad ♂♂ [6 UN], Bugarura Island, between 27 Jun. 1953 and 29 Jun. 1953,
leg. unknown collector (RBINS: 10537 to 10542); 1 ad ♀ [AL], Cyili, unknown date, leg. Benny Baeten
and Luc Janssens (RMCA: 85.006-M-0527); 2 ad ♀♀, 5 ad ♂♂, 4 sad ♀♀, 1 sad ♂, 1 juv ♀ [13 AL],
Gafunzo, unknown date, leg. Benny Baeten and Luc Janssens (RMCA: 85.006-M-0520, 85.006-M-0522
to 85.006-M-0525, 85.006-M-0528, 85.006-M-0530 to 85.006-M-0536); 1 ♀, 2 ?? [3 UN], Kissenji
[= Gisenyi], unknown date, leg. Stegmann-Stein (ZMB: 67137 to 67139); 3 ♀♀, 1 ♂, 9 ?? [13 UN],
Mugarura, unknown date, leg. Stegmann-Stein, unknown date, leg. H. Schubolz (ZMB: 66523, 67037
to 67041, 67043, 67146 to 67149, 67152, 67153); 4 ♀♀, 1 ♂ [4 SS, 1 SO], Nyarutovu, 19 Sep. 1954,
20 Sep. 1954, 10 Aug. 1955, 8 Oct. 1955, 10 Oct. 1955, leg. Alexandre Fain (RMCA: 23139, 23646,
MHNG: 921.026, 921.027, RMCA: 23645); 2 ad ♂♂, 1 sad ♂ [3 AL], Shyanda, unknown date, leg.
Benny Baeten and Luc Janssens (RMCA: 85.006-M-0521, 85.006-M-0526, 85.006-M-0529).
130
VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
Rousettinae K. Andersen, 1912
BURUNDI: 2 juv ?? [1 AL, 1 UN], Bujumbura, 22 Apr. 1982, 26 Apr. 1982, leg. Danny Meirte (RMCA:
82.016-M-0020, 82.016-M-0082).
DEMOCRATIC REPUBLIC OF THE CONGO: 8 foetus ?? [1 AL, 5 SK, 2 SO], unknown locality,
unknown date, leg. Tshinyenye, prior to 1 Aug. 1957, leg. Heinz Stephan, prior to 18 Aug. 1965, leg.
Antoon Emeric Marcel De Roo (RMCA: 77.021-M-0017, 77.021-M-0018, 27420 to 27424, 33602);
1 ♂ [SS], Bengale, 28 Dec. 1976, leg. Tshinyenye (RMCA: 77.012-M-0008); 1 ♀ [SS], Boma,
23 Nov. 1920, leg. Henri Schouteden (RMCA: 7294); 2 ?? [2 SS], Bukavu, prior to 30 Jun. 1999, leg.
Camerman (RMCA: 99.011-M-0002, 99.011-M-0003); 2 juv ??, 2 ♂♂, 1 ? [2 SS, 3 AL], Butembo,
17 Oct. 1971, between 21 May 1975 and 30 Jun. 1975, leg. M. Lejeune, 31 Dec. 1969, leg. M.J. Celis
and M. Lejeune, prior to 18 Apr. 1958, leg. P. Dyleff (RMCA: 38242, 36196, 75.042-M-0007, 27042,
27043); 1 ♀ [SS], Byonga, 5 Apr. 1962, leg. Alexandre Prigogine (RMCA: 31242); 2 ?? [2 SS], Dilolo,
9 Feb. 1948, leg. Jozef Hutsebaut (RMCA: 18228, 18229); 2 ?? [2 AL], Epulu, unknown date, leg.
Luc Bijnens (RMCA: 90.031-M-0005, 90.031-M-0006); 2 sad ♂♂, 1 juv ♀, 1 ♀ [4 SK], Idjwi Island,
19 Jul. 1965, 23 Aug. 1969, 26 Aug. 1969, between 25 Jul. 1969 and 25 Aug. 1969, leg. Alexandre
Prigogine (RMCA: 33626, 36217, 36218, 36216); 1 ♂ [SS], Kakanda Cave, 21 Aug. 1964, leg.
Alexandre Prigogine (RMCA: 33064); 1 ♀ [SS], Kamueshi [= Kamuesha], 3 Apr. 1979, leg. Betu
Nsankulu (RMCA: 79.032-M-0001); 1 ♂ [SS], Kasebere, 13 Jan. 1970, leg. Alexandre Prigogine
(RMCA: 36220); 1 juv ? [AL], Kilinga, 15 Oct. 1954, leg. Gaston-François de Witte (RMCA: 37114);
8 ♂♂ [3 SS, 5 SO], Kimpoko [= Kimpoko-Nsele], 15 Oct. 1976, 6 Sep. 1978, 7 Sep. 1978, 20 Jan. 1980,
leg. Tshinyenye (RMCA: 77.012-M-0009, 80.038-M-0001, 80.038-M-0002, 79.003-M-0002 to 79.003M-0006); 1 ♀ [SS], Kingabwa, 31 Dec. 1975, leg. Tshinyenye (RMCA: 76.010-M-0001); 1 foetus ?
[AL], Luluabourg [= Kananga], unknown date, leg. Jan Deheegher (RMCA: 31560); 1 ♀ [SS], Make,
30 Oct. 1992, leg. Tshinyenye (RMCA: 92.148-M-0002); 1 imm ? [AL], Moba, 26 Nov. 1953, leg.
H. Bomans (RMCA: 22124); 1 ♀, 5 ?? [6 SO], Mudina Ndemba [= Ndemba], 18 Mar. 1975, leg.
Betu Nsankulu (RMCA: 75.061-M-0001 to 75.061-M-0006); 1 ♂ [SS], Mulungu, 8 Oct. 1965, leg.
Alexandre Prigogine (RMCA: 33968); 1 juv ? [AL], Mutwanga, 8 Nov. 1954, leg. Gaston-François
de Witte (RMCA: 37111); 1 ♀, 3 ♂♂ [4 SS], Ngula, 1 Mar. 1977, 5 Mar. 1977, 6 Mar. 1977, leg.
Tshinyenye (RMCA: 77.021-M-0007, 77.021-M-0008, 77.021-M-0005, 77.021-M-0006); 3 ♀♀, 8 ♂♂
[11 SS], Nkiene, 28 Sep. 1980, 29 Sep. 1980, 30 Sep. 1980, leg. Tshinyenye (RMCA: 80.053-M-0002
to 80.053-M-0012); 1 ? [SS], Ogondjo, 24 Apr. 1942, leg. J.M. Vrijdagh (RMCA: 17758); 1 juv ? [AL],
Uvira, unknown date, leg. Georges Marlier (RMCA: 79.011-M-0002); 1 ? [AL], Wafanya [= Wafania],
29 Aug. 1991, leg. P. Lootens (RMCA: 91.076-M-0060).
Epomophorini Gray, 1866
Epomophorus Bennett, 1836
BURUNDI: 1 ad ♂ [AL], unknown locality, 13 Nov. 1978, leg. F.C. Roest (RMNH: MAM.45841).
DEMOCRATIC REPUBLIC OF THE CONGO: 1 ♀ [AL], unknown locality, 13 Nov. 1978, leg.
F.C. Roest, 3 Dec. 1985, leg. unknown collector (RMNH: MAM.45692); 1 ♀ [SA], Katwé, 1 Oct. 2009, leg.
Alexandre Hassanin, Nicolas Nesi, Célestin Pongombo, Peter Vallo and Didier Tshikung (MNHN: ZM2011-788); 1 ? [UN], Kilinga, unknown date, leg. unknown collector (INPBC: V1257.1); 4 ♀♀ [4 SS],
Kipangaribwe River, 18 Jul. 1947, 31 Dec. 1947, 6 May 1949, leg. unknown collector (RBINS: 10671
to 10673, 10676); 5 ♀♀, 1 ♂ [3 SA, 3 AL], Kitombongo [= Kintobongo], 24 Sep. 2009, leg. Alexandre
Hassanin, Nicolas Nesi, Célestin Pongombo, Peter Vallo and Didier Tshikung (MNHN: ZM-2011789 to ZM-2011-794); 1 emb ♀, 1 emb ? [1 SA, 1 AL], Lubumbashi, 28 Sep. 2009, leg. Alexandre
Hassanin, Nicolas Nesi, Célestin Pongombo, Peter Vallo and Didier Tshikung (MNHN: ZM-2011-795,
131
European Journal of Taxonomy 382: 1–327 (2017)
ZM-2011-796); 2 ♀♀ [2 SA], Mukulakulu, 22 Sep. 2009, leg. Alexandre Hassanin, Nicolas Nesi,
Célestin Pongombo, Peter Vallo and Didier Tshikung (MNHN: ZM-2011-797, ZM-2011-798); 1 ? [UN],
Ndote, unknown date, leg. unknown collector (INPBC: W1282); 1 imm ? [SS], Panda [= Pande],
15 Dec. 1955, leg. De Smet (RMCA: 26210); 1 ? [UN], Rutshuru, unknown date, leg. unknown
collector (INPBC: V1634).
RWANDA: 1 ? [AL], unknown locality, 27 Oct. 1979, leg. unknown collector (RMNH: MAM.45687).
Epomophorus anselli Bergmans & Van Strien, 2004
DEMOCRATIC REPUBLIC OF THE CONGO: 1 ♂ [SA], Kitombongo [= Kintobongo], 24 Sep. 2009,
leg. Alexandre Hassanin, Nicolas Nesi, Célestin Pongombo, Peter Vallo and Didier Tshikung
(MNHN: ZM-2011-726).
Epomophorus crypturus Peters, 1852
DEMOCRATIC REPUBLIC OF THE CONGO: 1 ? [SK], unknown locality, prior to 8 Jan. 1940, leg.
unknown collector (RBINS: 6283); 1 ad ♀ [SK], Dikulwe River, unknown date, leg. Neuve (BMNH:
1907.12.13.2); 3 ad ♀♀, 1 ad ♂, 1 juv ♀ [1 SS, 4 SA], Elisabethville [= Lubumbashi], 28 Dec. 1955,
29 Dec. 1955, leg. M. Lips, between 1 Apr. 1926 and 30 Apr. 1926, leg. Henri Schouteden, prior to
7 Jul. 1936, leg. Richard (RMCA: 26268 to 26270, 16208, 13316); 1 ? [UN], Gandu, unknown date, leg.
Gaston Thierry (ZMB: 54663); 1 ♂ [SS], Ganza, 28 Jun. 1949, leg. Gaston-François de Witte (RBINS:
10665); 3 sad ♂♂, 1 juv ♂, 2 ♀♀ [3 SA, 3 AL], Jadotville [= Likasi], 1 Mar. 1957, 1 Dec. 1957,
22 Jun. 1958, 15 Oct. 1958, 13 Feb. 1959, 6 Dec. 1959, leg. Félix / Michel Anciaux de Faveaux (RMCA:
26254, 27591, 27916 to 27918, RBINS: 14595); 1 ad ♀ [AL], Kakanda Cave, unknown date, leg. the
Royal Museum for Central Africa (RMCA) (BMNH: 1955.136); 1 sad ♂, 1 ♀ [1 SK, 1 SB], Kakanda
Mutaka, 14 Jul. 1953, leg. petit Seminaire de St. Francois de Sales (RMCA: 21874, 21875); 1 ? [UN],
Kakontwa [= Kakontwe], 17 Jan. 1961, leg. Félix / Michel Anciaux de Faveaux (RBINS: 14597); 1 ad
♂, 1 sad ♀, 1 sad ♂, 2 ♀♀ [1 SS, 3 SA, 1 SK], Kakontwe, 14 Nov. 1959, leg. William Frank Harding
Ansell, 17 Jan. 1961, leg. unknown collector, 30 Jun. 1957, between 30 Jun. 1956 and 30 Jun. 1957, leg.
Félix / Michel Anciaux de Faveaux (RMCA: 29246, RBINS: 14596, MHNG: 1046.035, RMCA: 26716,
26776); 10 ♀♀, 3 ♂♂ [13 AL], 5 km from Kaloba , 3 Oct. 2009, leg. Alexandre Hassanin, Nicolas Nesi,
Célestin Pongombo, Peter Vallo and Didier Tshikung (MNHN: ZM-2011-743 to ZM-2011-755); 1 ad ♂,
1 juv ?, 2 imm ♀♀, 1 ? [1 SS, 1 AL, 1 SK, 2 SO], Kanzenze Mission, 14 Jul. 1947, leg. R.P. Lefébure,
28 Nov. 1948, between 1 Dec. 1948 and 31 Dec. 1948, leg. Narcisse Leleup (RBINS: 13107, BMNH:
1954.786, RMCA: 22485, 22486, 22491); 2 ad ♀♀ [1 SA, 1 UN], Kasaji, unknown date, leg. W.S. Fisher,
13 Mar. 1953, leg. Fisher (CM: 21806, RMCA: 21806); 2 ♀♀, 3 ♂♂ [5 SS], Kasapa, 22 Aug. 1969,
leg. J. Poelman, 22 Aug. 1969, leg. unknown collector (RBINS: 18779, 18780, 18841 to 18843); 1 ♀
[AL], Kasenga Mission [= Kasenga], 28 Dec. 1956, leg. Van Iseghem (RMCA: 26273); 1 ♀ [AL],
Kasombo, 20 Sep. 2009, leg. Alexandre Hassanin, Nicolas Nesi, Célestin Pongombo, Peter Vallo and
Didier Tshikung (MNHN: ZM-2011-756); 1 ad ♂, 1 sad ♀ [1 SA, 1 AL], Katanga s, unknown date, leg.
André Pilette (RMCA: 16210, 16248); 2 ♀♀ [2 AL], Katwé, 1 Oct. 2009, 2 Oct. 2009, leg. Alexandre
Hassanin, Nicolas Nesi, Célestin Pongombo, Peter Vallo and Didier Tshikung (MNHN: ZM-2011-757,
ZM-2011-758); 1 ♂ [SN], Kipangaribwe River, 12 Jan. 1948, leg. unknown collector (RBINS: 10667);
6 ♀♀, 1 ♂ [7 AL], Kitombongo [= Kintobongo], 24 Sep. 2009, 25 Sep. 2009, 26 Sep. 2009, leg.
Alexandre Hassanin, Nicolas Nesi, Célestin Pongombo, Peter Vallo and Didier Tshikung (MNHN: ZM2011-759 to ZM-2011-765); 1 ad ♂ [SO], Kolwezi, 1 Mar. 1958, leg. unknown collector (OSU: 3817);
1 ad ♀, 1 ad ♂, 1 sad ♂, 2 ♀♀, 7 ♂♂ [4 SA, 8 AL], Lubumbashi, 28 Sep. 2009, 12 Oct. 2009, leg.
Alexandre Hassanin, Nicolas Nesi, Célestin Pongombo, Peter Vallo and Didier Tshikung, 24 Mar. 1966,
leg. L. Poelman, 26 Mar. 1966, leg. Xavier Misonne (MNHN: ZM-2011-769 to ZM-2011-774, RBINS:
17207, RMCA: 34712, 34713, MNHN: ZM-2011-766 to ZM-2011-768); 1 sad ♀ [SA], Luluabourg
132
VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
[= Kananga], 1 Jan. 1963, leg. Jan Deheegher (RMCA: 31557); 1 ♂ [SS], Mabwe, 7 Aug. 1947, leg.
unknown collector (RBINS: 10670); 3 ♀♀ [3 AL], Mukulakulu, 22 Sep. 2009, leg. Alexandre Hassanin,
Nicolas Nesi, Célestin Pongombo, Peter Vallo and Didier Tshikung (MNHN: ZM-2011-775 to ZM2011-777); 2 ♀♀ [1 SA, 1 AL], cave 4 km from Muvule , 8 Oct. 2009, 9 Oct. 2009, leg. Alexandre
Hassanin, Nicolas Nesi, Célestin Pongombo, Peter Vallo and Didier Tshikung (MNHN: ZM-2011-778,
ZM-2011-779); 1 ad ♂ [SA], Panda Camp [= Likasi], 1 Apr. 1958, leg. Félix / Michel Anciaux de
Faveaux (RMCA: 27594); 13 ♀♀, 2 ♂♂, 1 ? [1 SA, 15 AL], forest N of Pandé , 11 Oct. 2009, leg.
Alexandre Hassanin, Nicolas Nesi, Célestin Pongombo, Peter Vallo and Didier Tshikung (MNHN: ZM2011-727 to ZM-2011-742); 1 juv ♀ [SA], Pempere Cave, 20 Oct. 1957, leg. Félix / Michel Anciaux de
Faveaux (RMCA: 27607); 1 ? [SO], Sakania Mission [= Sakania], 19 Oct. 1930, leg. J. De Riemaecker
(RMCA: 19418); 1 ♀ [AL], Upemba National Park, 25 Sep. 2009, leg. Alexandre Hassanin, Nicolas
Nesi, Célestin Pongombo, Peter Vallo and Didier Tshikung (MNHN: ZM-2011-780).
Epomophorus labiatus (Temminck, 1837)
BURUNDI: 1 ♂ [KB], Buhonga, 2 Jun. 1970, leg. P.L. Niort (MHNG: 1499.046); 6 ad ♀♀, 5 sad ♀♀,
8 juv ♀♀, 10 juv ♂♂, 25 ♀♀, 102 ♂♂, 32 ?? [13 SS, 5 SA, 142 AL, 1 SK, 1 SO, 1 UN, 2 SB, 18 KB,
5 NS], Bujumbura, unknown date, between 1 Aug. 1976 and 31 Aug. 1976, leg. Jacques Verschuren,
25 Oct. 1967, 29 Oct. 1967, 6 Nov. 1967, 11 Nov. 1967, 22 Nov. 1967, 1 Dec. 1967, 5 Dec. 1967,
14 Dec. 1967, 1 Jan. 1968, 12 Jan. 1968, 13 Jan. 1968, 18 Oct. 1968, 1 Nov. 1968, 9 Nov. 1968,
5 Dec. 1968, 6 Dec. 1968, 8 Mar. 1969, 20 May 1969, 12 Nov. 1969, 14 Nov. 1969, 15 Nov. 1969,
31 Dec. 1969, 5 Jan. 1970, 6 Jan. 1970, 7 Jan. 1970, 9 Jan. 1970, 2 Feb. 1970, 26 Feb. 1970, 10 Mar. 1970,
26 Nov. 1970, 28 Jan. 1971, 29 Jan. 1971, 10 Mar. 1971, 1 Jan. 1972, 5 Jan. 1972, 24 Jan. 1972,
31 Jan. 1972, 30 Mar. 1972, 15 Aug. 1972, 17 Sep. 1972, 3 Feb. 1973, 14 Mar. 1973, 19 Jul. 1973,
10 Feb. 1974, 11 Feb. 1974, 13 Feb. 1974, 17 Feb. 1974, between 6 Jun. 1968 and 7 Jun. 1968, between
30 Mar. 1972 and 31 Mar. 1972, between 26 Oct. 1967 and 27 Oct. 1967, leg. P.L. Niort, 21 Apr. 1982,
22 Apr. 1982, 23 Apr. 1982, 24 Apr. 1982, 25 Apr. 1982, 26 Apr. 1982, 27 Apr. 1982, 28 Apr. 1982,
29 Apr. 1982, between 1 Apr. 1982 and 30 Apr. 1982, leg. Danny Meirte (RBINS: 19909, MHNG:
1498.054, 1499.068, 1498.065, 1498.066, 1498.049, 1499.086, 1499.026 to 1499.028, 1499.058,
1499.087, 1498.040 to 1498.043, 1498.055, 1498.093 to 1498.097, 1498.056 to 1498.058, 1499.088,
1499.081, 1497.097 to 1498.099, 1498.052, 1499.075, 1498.100, 1499.089, 1499.076, 1498.061
to 1498.064, 1499.085, 1499.021, 1499.023, 1498.083, 1498.084, RMCA: 82.016-M-0003, 82.016M-0004, 82.016-M-0010 to 82.016-M-0019, 82.016-M-0021, 82.016-M-0022, MHNG: 1498.044 to
1498.048, RMCA: 82.016-M-0023 to 82.016-M-0042, 82.016-M-0044 to 82.016-M-0048, MHNG:
1499.071, RMCA: 82.016-M-0049 to 82.016-M-0067, 82.016-M-0070, 82.016-M-0071, 82.016-M0073 to 82.016-M-0079, MHNG: 1498.035, 1499.024, 1499.025, RMCA: 82.016-M-0080, 82.016M-0081, 82.016-M-0083 to 82.016-M-0093, 82.016-M-0125 to 82.016-M-0128, MHNG: 1499.050,
1499.051, RMCA: 82.016-M-0130 to 82.016-M-0144, MHNG: 1499.054, RMCA: 82.016-M-0153,
MHNG: 1499.057, RMCA: 82.016-M-0154, 82.016-M-0155, MHNG: 1498.037, 1498.038, 1497.098,
1499.072, 1499.070, 1499.005, 1499.006, 1499.069, 1498.050, 1498.051, 1498.068 to 1498.070,
1499.008, 1498.039, 1498.071, 1499.007, 1498.075, 1499.009, 1498.067, RMCA: 82.016-M-0156,
RBINS: 19909.1 to 19909.4, MHNG: 1498.036, 1499.073, 1498.059, 1498.060); 1 ♂ [AL], Bukara,
17 Jun. 1972, leg. P.L. Niort (MHNG: 1499.074); 1 ♂ [KB], Busiga, between 1 Jan. 1900 and 1 Jan. 1967,
leg. P.L. Niort (MHNG: 1499.001); 1 ♀ [SS], Cibitoke, 15 May 1978, leg. unknown collector (RBINS:
20272); 1 ♀ [SS], Gifuruzi [= Kifuruzi], 8 Apr. 1969, leg. P.L. Niort (MHNG: 1498.076); 1 ad ♀,
20 ♀♀, 16 ♂♂, 1 ? [3 SS, 23 AL, 2 SK, 1 SB, 1 SN, 7 KB, 1 AB], Gihanga, 18 Jan. 1970, 25 Jan. 1970,
8 Apr. 1970, 11 Apr. 1970, 21 May 1970, 1 Nov. 1970, 2 Nov. 1970, 15 Nov. 1970, 24 Jan. 1971,
29 Jan. 1971, 17 Feb. 1971, 23 Mar. 1971, 22 Nov. 1972, 18 May 1973, leg. P.L. Niort, 14 Mar. 1979,
leg. Heinz Stephan (MHNG: 1499.047, 1499.038, 1499.039, SMF: 57512, MHNG: 1499.049, 1499.055,
1499.010, 1499.011, 1499.082 to 1499.084, 1499.048, 1499.040 to 1499.045, 1499.077, 1499.063,
133
European Journal of Taxonomy 382: 1–327 (2017)
1499.052, 1499.053, 1497.099, 1499.012 to 1499.020, 1499.056, 1499.032 to 1499.034, 1499.036,
1499.037); 1 juv ♀ [SA], Giheta, 8 Mar. 1979, leg. Heinz Stephan (SMF: 57511); 5 ♂♂ [1 SS, 3 AL,
1 KB], Gitega, 29 Dec. 1967, 12 Mar. 1971, 14 Mar. 1971, 25 Aug. 1971, between 13 Mar. 1971 and
14 Mar. 1971, leg. P.L. Niort (MHNG: 1499.059 to 1499.061, 1498.053, 1499.062); 2 ♂♂ [1 SS,
1 AL], Kibimba Mission, 3 Jan. 1969, leg. P.L. Niort (MHNG: 1498.073, 1498.074); 37 ?? [37 UN],
Kigamba, 15 Jul. 1946, 15 Jul. 1976, leg. Jacques Verschuren (RBINS: 19900.01, 19902, 19900.02 to
19900.35, 19902.2); 2 ♀♀ [2 SK], Lake Cohoha [= Cohoha, Lake], 9 Mar. 1979, 11 Mar. 1979, leg.
Heinz Stephan (SMF: 57510, 57509); 7 ♀♀ [2 SS, 2 AL, 3 KB], Makamba, 9 Apr. 1969, 10 Apr. 1969,
23 Mar. 1970, leg. P.L. Niort (MHNG: 1498.080, 1498.081, 1499.030, 1499.031, 1498.077 to 1498.079);
4 ♀♀ [4 AL], Murore, 27 May 1971, leg. P.L. Niort (MHNG: 1499.064 to 1499.067); 2 ♂♂ [1 SS,
1 AL], Mweya, 19 Dec. 1972, leg. P.L. Niort (MHNG: 1499.078, 1499.079); 3 ad ♀♀, 3 ♀♀, 3 ♂♂
[2 SA, 6 SK, 1 CO], Resha, 28 Feb. 1979, 2 Mar. 1979, 3 Mar. 1979, 15 Mar. 1979, 17 Mar. 1979,
3 Apr. 1979, 16 Jun. 1979, leg. Heinz Stephan (SMF: 57504 to 57506, 57514, 57513, 57503, 57507,
57508, 57515); 1 ♀ [SB], Ruhinga Hill, 12 Mar. 1993, leg. Jay L. Udelhoven (FMNH: 155941); 1 ad ♂,
1 ♀ [1 SS, 1 KB], Rumonge, 3 Nov. 1969, leg. P.L. Niort, between 1 Feb. 1955 and 28 Feb. 1955, leg.
Alexandre Fain (MHNG: 1498.092, RMCA: 81.012-M-0042); 1 juv ♂, 3 ♀♀, 1 ♂, 1 ? [6 AL], Russissi,
bank [= Rusizi Delta], 27 Apr. 1982, leg. Danny Meirte (RMCA: 82.016-M-0145, 82.016-M-0147,
82.016-M-0149 to 82.016-M-0152); 9 ♀♀ [1 SS, 4 AL, 1 SB, 3 KB], Zimet Quarter, 13 Oct. 1969,
25 Oct. 1969, 29 Nov. 1969, 10 Dec. 1969, leg. P.L. Niort (MHNG: 1499.003, 1499.004, 1498.085,
1498.086, 1498.088 to 1498.091, 1499.002).
DEMOCRATIC REPUBLIC OF THE CONGO: 1 ? [SS], unknown locality, prior to 5 Dec. 1957, leg.
De Semeries (RMCA: 26682); 2 ?? [2 UN], Albert National Park [= Virunga National Park],
unknown date, leg. unknown collector (INPBC: V1839.2.1, V1839.2.2); 2 ad ♂♂ [1 SA, 1 AL],
Albertville [= Kalemie], unknown date, leg. the Royal Museum for Central Africa (RMCA), prior to
12 Jul. 1955, leg. service hygiene publique albertville (BMNH: 1955.1191, RMCA: 22947); 1 ? [SK],
Alima, 23 Sep. 1959, leg. P. Dyleff (RMCA: 28530); 3 ad ♀♀, 2 ad ♂♂, 1 juv ♂, 1 ♀, 2 ♂♂ [4 AL,
4 SK, 1 UN], Baraka, unknown date, 16 Apr. 1947, leg. the Belgian Hydrobiological Expedition to
Tanganyika, 16 Apr. 1947, leg. unknown collector (BMNH: 1953.1, 1953.2, 1953.4, 1853.1, 1953.3,
RBINS: 9061.1 to 9061.4); 1 ad ?, 2 juv ?? [3 SK], Beni, 2 Jun. 1914, leg. Bonnevie, between 20 Oct. 1932
and 20 Oct. 1933, between 1 Oct. 1932 and 31 Oct. 1932, leg. Henri Martin Hackars (RMCA: 3245,
12134, 12133); 1 juv ♂ [SA], Bobandana, 10 Jul. 1933, leg. unknown collector (MNHN: ZMMO-1975-1066); 2 ?? [2 UN], Bongeya River, unknown date, leg. unknown collector (INPBC: V264.1,
V264.3); 1 ♀ [SS], Bushanganya, 20 Oct. 1953, leg. De Schrijver (RMCA: 27939); 3 ad ♂♂, 1 sad ♂,
1 juv ♀, 1 juv ♂, 2 juv ??, 2 imm ??, 45 ♀♀, 21 ♂♂, 45 ?? [44 SS, 1 SA, 65 AL, 1 SK, 10 SO], Butembo,
unknown date, 25 Apr. 1954, 22 Apr. 1956, 28 Apr. 1956, 27 Jul. 1956, 25 Jul. 1957, 20 Aug. 1957,
26 Aug. 1957, 8 Sep. 1957, 8 Jan. 1958, 10 Jan. 1958, 5 Feb. 1958, 27 Feb. 1958, 6 Mar. 1958,
18 Mar. 1958, 4 Apr. 1958, 16 Apr. 1958, 23 Apr. 1958, 24 Apr. 1958, 28 Apr. 1958, 5 May 1958, 14 May
1958, 16 May 1958, 2 Jun. 1958, 4 Jun. 1958, 9 Jun. 1958, 16 Jun. 1958, 8 Jul. 1958, 10 Jul. 1958,
15 Jul. 1958, 22 Jul. 1958, 12 Aug. 1958, 13 Aug. 1958, 14 Aug. 1958, 16 Aug. 1958, 20 Aug. 1958,
17 Sep. 1958, 1 Oct. 1958, 18 Jan. 1961, prior to 9 Oct. 1957, prior to 29 Jul. 1957, prior to 26 May
1959, prior to 26 Jan. 1955, prior to 23 Oct. 1959, prior to 18 Apr. 1958, prior to 17 Dec. 1956, between
7 Jun. 1958 and 8 Jul. 1958, between 1 Jul. 1956 and 31 Jul. 1956, between 1 Jun. 1956 and 30 Jun. 1956,
between 1 Apr. 1957 and 30 Apr. 1957, between 1 Jan. 1931 and 31 Dec. 1931, leg. P. Dyleff, 31 Dec. 1969,
1 Jan. 1970, leg. M.J. Celis and M. Lejeune, 17 May 1975, 20 May 1975, 25 May 1975, 1 Jun. 1975,
7 Jun. 1975, leg. M. Lejeune, 13 Aug. 1956, leg. De Semeries, prior to 5 Jul. 1950, leg. Alexandre
Prigogine (RMCA: 27651 to 27654, 36193, 36195, 36201, 36203, 75.042-M-0003, 27781, 27357,
27484, 27490, 26681, 27779, 27780, 27380, 27489, 27482, 27350, 27379, 27382, 27485, 27493, 75.042M-0004, 27491, 27492, 30809, 27356, 27383, 75.042-M-0005, 26650, 27486, 23741, 27487, 27355,
27351, 22722, 75.042-M-0006, 26652, 26651, 27352, 26198, 23740, 27349, 27353, 27376, 36197,
134
VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
27481, 27385, 27358, 27354, 27381, 75.042-M-0001, 75.042-M-0002, 27386, 27483, 26894, 27377,
27387, 26423, 25848 to 25851, 24004 to 24013, 27378, 24014 to 24018, 27041, 27044 to 27050, 28322
to 28339, 22813 to 22815, 27886 to 27890, 26434, 19922, 26583 to 26586); 1 sad ♀, 3 ♀♀ [3 SS, 1 SK],
Cel I [= Bagbele], 1 Feb. 1950, 24 Feb. 1950, 17 May 1950, leg. H. de Saeger et al., 24 Feb. 1950, leg.
H. de Saeger (RBINS: 13506 to 13508, RMCA: 36546); 1 ad ♀, 4 ad ♂♂, 1 sad ♀, 1 juv ♀, 1 ♀, 3 ♂♂
[1 SS, 2 SA, 4 SK, 4 UN], Cel II, 1 Feb. 1951, 1 Mar. 1951, 31 Mar. 1952, 2 Apr. 1952, leg. H. de Saeger
et al., 1 Mar. 1951, 15 Jan. 1952, 31 Mar. 1952, 2 Apr. 1952, 3 Apr. 1952, leg. H. de Saeger (RBINS:
13511, 13510, RMCA: 36552, 36547, RBINS: 13516, RMCA: 36550, 36551, RBINS: 13513.1, 13513.2,
RMCA: 36548, 36549); 1 juv ♂, 1 ? [2 AL], Costermansville [= Bukavu], between 1 May 1950 and
31 May 1950, leg. Babilon, prior to 11 Jan. 1939, leg. Hautmann (RMCA: 20271, 14975); 6 ad ♀♀, 6
ad ♂♂, 1 ad? ♀, 1 sad ♀, 5 juv ♀♀, 1 juv ♂, 4 ♀♀, 1 ? [20 SS, 4 AL, 1 UN], Faradje, 1 Apr. 1952, leg.
H. de Saeger et al., 7 Apr. 1911, 2 Jul. 1911, 6 Nov. 1911, 22 Nov. 1911, 23 Dec. 1911, 2 Jan. 1912,
23 Feb. 1912, 16 Sep. 1912, 22 Oct. 1912, 5 Nov. 1912, 2 Dec. 1912, 17 Dec. 1912, leg. Herbert Lang,
James Paul Chapin and The American Museum Congo Expedition (RBINS: 13517, AMNH: 48747,
48750, 48743, 48745, 48746, RMCA: 12374, 12373, AMNH: 48749, 49204, 49200 to 49202, 48742,
48748, 48661, 48734 to 48741, 48732); 2 ♂♂ [2 UN], Fataki, between 1 Jan. 1948 and 6 Apr. 1964,
between 1 Jan. 1948 and 31 Dec. 1948, leg. Jacques Verschuren (RBINS: 18528.1, 18528.2); 3 ad ♂♂
[3 SA], Fomulac-Katana, 29 Sep. 1963, 30 Sep. 1963, 3 Oct. 1963, leg. P. Kunkel (SMF: 31792, 31793,
31795); 2 ad ♀♀, 1 ♀, 2 ♂♂, 6 ?? [5 SS, 5 SK, 1 SO], Gangala-na-Bodio, 1 Dec. 1956, between
1 Dec. 1956 and 31 Dec. 1956, leg. Max Poll, 22 Aug. 1945, 22 Nov. 1947, 9 Apr. 1948, leg. unknown
collector (RMCA: 25602, 25603, RBINS: 12936, 12934, 12935, 11658.1 to 11658.4, RMCA: 25604,
25605); 1 ad ♂, 1 ♀ [1 SA, 1 UN], Garamba National Park, 1 Jan. 1950, leg. H. de Saeger et al.,
between 1 Jan. 1950 and 31 Dec. 1952, leg. H. de Saeger (RBINS: 13518, RMCA: 36553); 1 ad ♀ [SS],
Ibebero, 14 Jul. 1964, leg. Svepel (ZMZ: 15596); 1 ad? ? [SS], Kabale, between 10 Aug. 1939 and
5 Sep. 1939, leg. Schwetz (RMCA: 15833); 1 ad ♂ [SO], Kakondo, between 1 Dec. 1946 and
31 Dec. 1946, leg. F.L. Hendricks (RMCA: 19654); 1 ad ♀, 1 sad ♀, 3 sad ?? [5 SK], Kaniki Kapangu,
12 Mar. 1966, 24 Dec. 1966, between 24 Mar. 1966 and 24 Dec. 1966, between 21 Dec. 1966 and
24 Dec. 1966, leg. Antoon Emeric Marcel De Roo (RMCA: 35276, 35282, 35284, 35285, 35283); 1 ?
[SO], Kasai o, prior to 16 Oct. 1968, leg. Antoon Emeric Marcel De Roo (RMCA: 35286); 1 ad ♀ [SA],
Kasaka, prior to 7 Dec. 1971, leg. Vanderborght (RMCA: 38155); 1 juv ♀, 1 ? [1 AL, 1 UN], Katanda,
unknown date, leg. unknown collector, 12 May 1954, leg. Gaston-François de Witte (INPBC: W976,
RMCA: 37109); 2 ad ♂♂, 1 sad ? [1 SS, 1 SK, 1 SO], Kere Kere [= Kerekere], between 1 Jan. 1950
and 27 Oct. 1950, leg. Turco (RMCA: 20094 to 20096); 1 ♀ [UN], Keyberg, prior to 5 Aug. 1952, leg.
unknown collector (RBINS: 13106); 1 ♀ [SS], Kilwezi, 6 Aug. 1948, leg. Gaston-François de Witte
(RBINS: 10677); 1 ad ♀ [SA], Kiseny [= Kasenye], between 1 Jan. 1926 and 31 Jan. 1926, leg. Henri
Schouteden (RMCA: 16206); 1 ♀ [SA], Kitombongo [= Kintobongo], 24 Sep. 2009, leg. Alexandre
Hassanin, Nicolas Nesi, Célestin Pongombo, Peter Vallo and Didier Tshikung (MNHN: ZM-2011-781);
2 ad ♀♀, 1 sad ♀, 1 sad ♂, 2 juv ♀♀, 6 ♀♀, 3 ♂♂, 1 ? [13 SA, 3 AL], Kivu region, 25 Jun. 1963,
28 Jun. 1963, 22 Jul. 1963, 12 Sep. 1963, 24 Sep. 1963, 29 Sep. 1963, 30 Sep. 1963, 1 Oct. 1963,
2 Oct. 1963, 23 Nov. 1963, 4 Jun. 1964, 24 Dec. 1964, between 1 Jul. 1963 and 31 Dec. 1963, leg.
P. Kunkel (SMF: 31798, 31804, 31808, 31806, 31814, 31813, 31805, 31802, 31815, 31794, 31796,
31799, 31797, 31807, 31800, 31801); 1 ? [AL], Kiyite, 7 Apr. 1969, leg. M.J. Celis and M. Lejeune
(RMCA: 36199); 1 ♀ [SS], Léopoldville [= Kinshasa], 16 Jul. 1942, leg. Colback (RMCA: 31127);
1 ♀ [AL], Luebo, 25 Jul. 1927, leg. James Paul Chapin (AMNH: 82375); 1 ♀ [SA], Luena, 23 Sep. 2009,
leg. Alexandre Hassanin, Nicolas Nesi, Célestin Pongombo, Peter Vallo and Didier Tshikung
(MNHN: ZM-2011-782); 1 juv ♂ [SA], Lukonzolwa, 20 Nov. 1911, leg. Louis Stappers (RMCA:
16490); 1 ad ♂ [SS], Lulimba, 9 Oct. 1953, leg. Alexandre Prigogine (RMCA: 22355); 1 ad ♀ [SA],
Lulimbi, prior to 13 Jun. 1977, leg. M. Lejeune (RMCA: 77.029-M-0001); 1 ♀ [SB], Lurhogole
Swamp area, 10 Jul. 2005, leg. Project Beatra (FMNH: 189290); 9 ad ♀♀, 7 ad ♂♂, 5 ♀♀, 3 ♂♂ [8 SS,
5 SA, 2 SK, 1 SO, 8 SB], Lwiro [= Luiro], 22 Jun. 1964, 10 Jul. 1964, 14 Jul. 1964, 15 Jul. 1964,
135
European Journal of Taxonomy 382: 1–327 (2017)
17 Jul. 1964, 27 Nov. 1964, leg. Svepel, 9 Jul. 2001, 13 Jul. 2001, 29 Jul. 2002, leg. Michael H. Huhndorf,
14 Oct. 1963, 28 Nov. 1964, leg. Fritz Dieterlen, 24 Mar. 1963, 25 Sep. 1963, 27 Sep. 1963, 20 Dec. 1965,
between 1 Aug. 1963 and 31 Aug. 1963, leg. P. Kunkel, 6 Apr. 1967, 28 Apr. 1967, leg. Alexandre
Prigogine (ZMZ: 15528, FMNH: 171628, 171629, ZMZ: 15534, 16678, SMNS: 7289, ZMZ: 15594,
14140, SMF: 31810, ZMZ: 15535, SMF: 31803, 31812, 31811, ZMZ: 15581, RMCA: 35070, SMNS:
7290, FMNH: 173292 to 173294, RMCA: 35071, FMNH: 171625 to 171627, SMF: 31809); 1 ? [AL],
Lwiro Katana [= Luiro], between 1 Aug. 1954 and 31 Oct. 1954, leg. Laurent (RMCA: 27505); 1 ad
♂ [SK], Mahagi Port, 5 May 1925, leg. Henri Schouteden (RMCA: 9004); 1 ♀ [AL], Moba, unknown
date, leg. H. Bomans (BMNH: 1957.431); 1 ad ♂, 1 sad ?, 1 ? [2 SS, 1 SK], Mount Wago [= Wago,
Mount], 18 Sep. 1952, leg. unknown collector, between 13 Sep. 1952 and 18 Sep. 1952, between
1 Jan. 1951 and 28 Mar. 1952, leg. Alexandre Fain (RBINS: 21667, RMCA: 20803, 23165); 1 ad ♀
[SA], Mukwande River, prior to 28 Dec. 1971, leg. Gaston-François de Witte (RMCA: 37108); 1 ?
[UN], Mulala, unknown date, leg. unknown collector (INPBC: W332); 1 ad ♂ [SS], Mulongo,
30 Oct. 1974, leg. Michael D. Gallagher (HZM: 32.7869); 1 ad ♀, 1 ♂, 4 ?? [2 SS, 4 AL], Mulungu,
12 Feb. 1947, prior to 1 Feb. 1940, between 1 Jan. 1948 and 31 Dec. 1948, leg. F.L. Hendricks (RMCA:
19656, 19651, 15875 to 15878); 1 sad ? [SK], Mutembu Kaniki Kapangu [= Kaniki Kapangu],
12 Mar. 1966, leg. Antoon Emeric Marcel De Roo (RMCA: 35279); 1 ad ♂, 1 sad ♀, 1 sad ♂, 1 juv ♀,
1 ♂, 3 ?? [2 SA, 1 AL, 2 SO, 3 UN], Mutsora, unknown date, 14 May 1952, leg. unknown collector,
19 Nov. 1952, 1 Sep. 1954, 7 Jan. 1955, 2 Feb. 1955, leg. Gaston-François de Witte (INPBC: W1240,
W1367, W1475, RMCA: 37110, RBINS: 17292, RMCA: 37106, 37113, 37112); 1 ? [UN], Ndotwe,
unknown date, leg. unknown collector (INPBC: V1439); 1 juv ♂ [SO], Ngokai, 21 Mar. 1953, leg.
Gaston-François de Witte (RMCA: 37107); 1 ad ♂ [SS], Nkene, 6 May 1959, leg. Uwe Rahm (RMCA:
28030); 1 juv ♂ [SS], Nyakaliba, 10 Feb. 1954, leg. Gaston-François de Witte (RBINS: 17293); 1 ♀
[UN], Panga, between 1 Sep. 1925 and 30 Sep. 1925, leg. Henri Schouteden (RMCA: 9139a); 1 juv ♂,
1 ♀, 1 ♂, 1 ? [4 SS], Parc National de Garamba [= Garamba National Park], 15 Jan. 1952,
4 Feb. 1952, 2 Apr. 1952, leg. H. de Saeger et al. (RBINS: 13512, 13519, 13514, 13515); 2 ad ♂♂, 2 sad
♀♀, 3 ♀♀, 1 ♂, 15 ?? [3 SS, 1 SA, 2 AL, 2 SK, 15 UN], Rutshuru, unknown date, 8 Jan. 1938,
10 Jan. 1938, leg. unknown collector, 1 Mar. 1937, 30 Mar. 1937, leg. J. Ghesquière, 1 Aug. 1925,
3 Aug. 1925, leg. Edmund Heller, 23 May 1927, leg. James Paul Chapin, Dewitt L. Sage, Frank
P. Mathews and The Ruwenzori-Kivu Expedition, between 1 Jun. 1935 and 30 Jun. 1935, leg. GastonFrançois de Witte (INPBC: V125, V1310, V1382.2, V1386.2, V1387, V1423, V1525.2, V1574, V1592,
V1641, V397, V398, V408, V411, W618, RMCA: 13825, FMNH: 26497, RBINS: 4787, AMNH: 82378,
FMNH: 30578, RMCA: 13867, RBINS: 4786, RMCA: 17082); 2 ad ♀♀, 1 ad ♂ [3 SK], Ruwenzori
SE, unknown date, leg. R.E. Dent (BMNH: 1906.12.4.10, 1906.12.4.7, 1906.12.4.9); 1 ad ♀ [SS],
Semmio [= Zemio], 1 Jan. 1884, leg. Bohndorff (RMNH: MAM.27276); 1 ♀ [SS], Topetope, 9 Jul. 1966,
leg. Alexandre Prigogine (RMCA: 34380); 1 ad ♀ [SO], Tshibinda, 16 Sep. 1950, leg. F.L. Hendricks
(RMCA: 20265); 3 ad ♀♀, 1 ad ?, 1 sad ♂, 1 juv ♀, 1 ♀, 3 ♂♂ [1 SS, 4 SA, 3 AL, 2 SO], Uvira,
unknown date, 17 Jun. 1958, leg. Georges Marlier, unknown date, leg. Waldo LaSalle Schmitt and
E. Baker, 1 Feb. 1910, leg. Graub, 12 Sep. 1951, leg. Narcisse Leleup, 23 Jan. 1962, 3 Feb. 1962, leg.
Kiss, 4 May 1950, leg. Laurent (RMCA: 79.011-M-0001, USNM: 301713, 301714, NMW: 4880,
RMCA: 22755, 22756, 27987, 31008, 31009, 79.010-M-0004); 1 ad ♂, 1 sad ? [1 SS, 1 SK], Mount
Wago , between 1 Jan. 1951 and 31 Dec. 1951, leg. Alexandre Fain (RMCA: 23149, 23150).
RWANDA: 3 ad ♂♂, 2 sad ♂♂, 1 juv ?, 7 ?? [7 SS, 2 SA, 4 AL], Astrida [= Butare], 3 May 1950,
10 Jan. 1951, leg. Aureliaan, 12 Oct. 1976, 14 Oct. 1976, leg. Frieder Meier, 14 Oct. 1955, between 1 May
1955 and 31 May 1955, leg. Alexandre Fain, 20 Feb. 1962, leg. Van Parijs, prior to 14 Apr. 1955, leg.
P.G. Vercammen (RMCA: 20492, MHNG: 1872.016 to 1872.019, RMCA: 23140, MHNG: 1872.020,
RMCA: 31044, 20175, 20176, MHNG: 921.029, 921.030, RMCA: 23170); 15 ad ♀♀, 17 ad ♂♂, 9 sad
♀♀, 8 sad ♂♂, 5 juv ♀♀, 3 juv ♂♂, 1 ♀ [50 SA, 8 AL], Butare, unknown date, leg. Benny Baeten,
11 Dec. 1984, 20 Dec. 1984, 23 Dec. 1984, 24 Dec. 1984, 25 Dec. 1984, 31 Dec. 1984, 1 Jan. 1985,
136
VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
2 Jan. 1985, 14 Jan. 1985, 15 Jan. 1985, leg. Benny Baeten and Luc Janssens, 7 Oct. 1981, 8 Oct. 1981,
9 Oct. 1981, 8 Nov. 1981, leg. Frits De Vree, Benny Baeten and Victor Van Cakenberghe (RMCA:
87.046-M-0350 to 87.046-M-0356, 87.046-M-0362, 87.046-M-0376 to 87.046-M-0378, 85.006M-0469, 85.006-M-0480, 85.006-M-0491 to 85.006-M-0494, 85.006-M-0490, 85.006-M-0470 to
85.006-M-0475, 85.006-M-0479, 85.006-M-0483, 85.006-M-0484, 85.006-M-0500, 85.006-M-0501,
85.006-M-0508, 85.006-M-0509, 85.006-M-0478, 85.006-M-0481, 85.006-M-0482, 85.006-M-0495 to
85.006-M-0499, 85.006-M-0506, 85.006-M-0507, 85.006-M-0505, 85.006-M-0476, 85.006-M-0502,
85.006-M-0503, 85.006-M-0519, 85.006-M-0466, 85.006-M-0467, 85.006-M-0477, 85.006-M-0504,
85.006-M-0468, 82.006-M-1076 to 82.006-M-1078, 82.006-M-1031 to 82.006-M-1033, 82.006-M1079); 1 ad ♀, 2 sad ♀♀, 4 ♀♀, 9 ♂♂, 3 ?? [3 SA, 16 SK], Butare INRS [= Butare], 17 Feb. 1971,
20 Feb. 1971, 22 Apr. 1971, leg. M. Lejeune, 23 Apr. 1971, leg. A. Elbl (RMCA: 74.020-M-0048 to
74.020-M-0054, 74.020-M-0056 to 74.020-M-0062, 74.020-M-0055, 74.020-M-0063, 35912 to 35914);
3 ad ♀♀, 1 ad ♂, 2 sad ♂♂, 1 juv ♀ [5 SA, 2 AL], Cyili, 23 Dec. 1984, 24 Dec. 1984, leg. Benny
Baeten and Luc Janssens (RMCA: 85.006-M-0485, 85.006-M-0511, 85.006-M-0512, 85.006-M-0486
to 85.006-M-0488, 85.006-M-0510); 1 ad ♀, 1 ad ♂, 1 sad ♂ [3 SA], Gafumba, 12 Jan. 1985, leg.
Benny Baeten and Luc Janssens (RMCA: 85.006-M-0489, 85.006-M-0513, 85.006-M-0514); 2 juv ♂♂
[2 SA], Gafunzo Kigaga, 5 Jan. 1985, 6 Jan. 1985, leg. Benny Baeten and Luc Janssens (RMCA:
85.006-M-0515, 85.006-M-0516); 4 ad ♀♀, 2 ad ♂♂ [6 SS], Gakoma (Mission), unknown date,
20 Sep. 1964, 21 Sep. 1964, 27 Sep. 1964, 28 Sep. 1964, leg. Svepel (ZMZ: 15540, 15544, 16677,
15530, 15542, 16676); 1 ad ♀, 1 ad ♂, 5 sad ♀♀, 2 sad ♂♂, 2 juv ♀♀, 1 imm ♀, 1 ♀ [6 SA, 7 AL],
Gisenyi, 21 Oct. 1980, leg. Danny Meirte, 21 Oct. 1981, leg. Frits De Vree, Benny Baeten and Victor
Van Cakenberghe, 28 Jun. 1980, leg. Hans H. de Iongh (RMCA: 80.055-M-0015 to 80.055-M-0017,
80.055-M-0025 to 80.055-M-0031, 82.006-M-1034, ZMA: MAM.21241, MAM.21242); 1 ad ♂ [SA],
Gitovu, 28 Apr. 1971, leg. A. Elbl (RMCA: 35911); 1 sad ♀ [AL], Karama, 13 Oct. 1981, leg. Frits
De Vree, Benny Baeten and Victor Van Cakenberghe (RMCA: 82.006-M-1035); 1 ad ♂, 2 sad ♀♀
[2 SA, 1 AL], Kayonza, 6 Nov. 1980, leg. Danny Meirte (RMCA: 80.055-M-0049 to 80.055-M-0051);
7 ad ♂♂, 2 sad ♀♀, 1 sad ♂, 1 juv ♀ [10 SA, 1 AL], Kibuye, 9 Oct. 1980, 1 Nov. 1980, leg. Danny
Meirte, 3 Dec. 1971, leg. Alena Vanderborght-Elbl, G. Mathys and A. Elbl (RMCA: 80.055-M-0019,
80.055-M-0020, 80.055-M-0042 to 80.055-M-0048, 73.058-M-0001, 80.055-M-0010); 3 ad ♀♀, 5 ad
♂♂, 2 sad ♀♀, 8 sad ♂♂ [8 SA, 10 AL], Kidaho, unknown date, leg. Benny Baeten (RMCA: 87.046M-0357 to 87.046-M-0361, 87.046-M-0363 to 87.046-M-0375); 5 ad ♀♀, 1 ad ♂, 1 sad ♀, 3 sad ♂♂,
1 juv ♀ [7 SA, 4 AL], Kigali, 19 Nov. 1971, leg. Alena Vanderborght-Elbl, G. Mathys and A. Elbl,
26 Oct. 1980, 27 Oct. 1980, leg. Danny Meirte (RMCA: 73.057-M-0011, 80.055-M-0033 to 80.055M-0035, 80.055-M-0018, 80.055-M-0036 to 80.055-M-0041); 1 ♀, 1 ♂ [2 AL], Kisenyi [= Gisenyi],
5 Jul. 1927, leg. James Paul Chapin, Dewitt L. Sage, Frank P. Mathews and The Ruwenzori-Kivu
Expedition (AMNH: 82376, 82377); 1 ad ♀, 1 juv ♂ [2 SA], Mabanza, 10 Oct. 1980, leg. Danny Meirte
(RMCA: 80.055-M-0011, 80.055-M-0021); 2 ad ♀♀, 1 sad ♂ [3 SA], Masaka - Kigali, 19 Nov. 1971,
leg. Alena Vanderborght-Elbl, G. Mathys and A. Elbl (RMCA: 73.057-M-0003 to 73.057-M-0005); 4 ad
♀♀, 1 ad ♂ [5 SS], Mimuli, 4 Nov. 1964, 5 Nov. 1964, 6 Nov. 1964, 10 Nov. 1964, leg. Svepel (ZMZ:
15539, 15541, 15536, 14141, 15527); 3 ♀♀ [3 SO], Mugusa, Mount, 12 Jun. 1968, leg. unknown
collector (RBINS: 16031 to 16033); 3 ad ♀♀, 1 ad ♂, 1 sad ♂, 2 emb ?? [5 SA, 2 AL], Muhasi [=
Muhazi], 20 Aug. 1971, 23 Aug. 1971, 25 Aug. 1971, leg. Alena Vanderborght-Elbl, G. Mathys and
A. Elbl (RMCA: 73.057-M-0006, 73.057-M-0001, 73.057-M-0007, 73.057-M-0002, 73.057-M-0008 to
73.057-M-0010); 2 ad ♀♀, 1 sad ♂, 1 juv ♂ [3 SA, 1 AL], Murunda, 14 Oct. 1980, 15 Oct. 1980, leg.
Danny Meirte (RMCA: 80.055-M-0022, 80.055-M-0012, 80.055-M-0023, 80.055-M-0024); 1 ♂ [SS],
Musha, 25 Mar. 1968, leg. Félix / Michel Anciaux de Faveaux (RBINS: 16023); 4 ad ♀♀, 1 ad ♂, 4 sad
♀♀, 13 sad ♂♂, 1 ? [10 SA, 13 AL], Nyamata, 11 Nov. 1981, 12 Nov. 1981, 13 Nov. 1981, 14 Nov. 1981,
15 Nov. 1981, 16 Nov. 1981, leg. Frits De Vree, Benny Baeten and Victor Van Cakenberghe (RMCA:
82.006-M-1093, 82.006-M-1101, 82.006-M-1062, 82.006-M-1102 to 82.006-M-1104, 82.006-M-1058,
82.006-M-1105, 82.006-M-1059, 82.006-M-1094 to 82.006-M-1096, 82.006-M-1055, 82.006-M-1060,
137
European Journal of Taxonomy 382: 1–327 (2017)
82.006-M-1097 to 82.006-M-1099, 82.006-M-1056, 82.006-M-1057, 82.006-M-1061, 82.006-M-1092,
82.006-M-1100, 82.006-M-1139); 9 ad ♀♀, 5 ad ♂♂, 5 sad ♀♀, 4 sad ♂♂, 4 juv ♀♀, 3 juv ♂♂,
1 imm ♀ [15 SA, 16 AL], Nyumba Cave, 3 Oct. 1981, 6 Oct. 1981, 7 Oct. 1981, leg. Frits De Vree,
Benny Baeten and Victor Van Cakenberghe (RMCA: 82.006-M-1037, 82.006-M-1039, 82.006-M-1040,
82.006-M-1042 to 82.006-M-1044, 82.006-M-1047, 82.006-M-1049, 82.006-M-1054, 82.006-M-1080
to 82.006-M-1090, 82.006-M-1140, 82.006-M-1036, 82.006-M-1038, 82.006-M-1041, 82.006-M1045, 82.006-M-1048, 82.006-M-1050 to 82.006-M-1053, 82.006-M-1091); 1 ad ♂, 1 juv ♀, 1 juv ♂
[3 SA], Nyundo, 17 Oct. 1980, 18 Oct. 1980, 23 Oct. 1980, leg. Danny Meirte (RMCA: 80.055-M0013, 80.055-M-0014, 80.055-M-0032); 1 sad ♀ [SA], Rubengera (Mission), between 1 Jan. 1931
and 31 Dec. 1931, leg. Lestrade (RMCA: 11462); 1 ad ♀, 2 ad ♂♂, 7 ♀♀, 1 ♂, 2 ?? [7 SS, 4 SA, 1 SK,
1 SO], Rubona, 18 Jan. 1966, 15 Oct. 1966, 22 Nov. 1966, 4 Dec. 1966, 10 Dec. 1966, 21 Dec. 1966,
27 Dec. 1966, 8 Jan. 1967, leg. Félix / Michel Anciaux de Faveaux, between 1 Jan. 1985 and 30 Jun. 1985,
leg. Théoneste (RBINS: 16028 to 16030, 15481.1, 15481.2, 16035, 16027, 16026, 16024, 16025,
16034, RMCA: 87.047-M-0132, 87.047-M-0133); 3 ad ♀♀, 11 ad ♂♂, 6 sad ♀♀, 20 sad ♂♂, 4 juv
♀♀, 2 juv ♂♂ [24 SA, 22 AL], Rukira, 19 Nov. 1981, 20 Nov. 1981, 21 Nov. 1981, 22 Nov. 1981,
23 Nov. 1981, 24 Nov. 1981, 25 Nov. 1981, 26 Nov. 1981, leg. Frits De Vree, Benny Baeten and Victor
Van Cakenberghe (RMCA: 82.006-M-1106, 82.006-M-1063, 82.006-M-1066, 82.006-M-1067, 82.006M-1107 to 82.006-M-1111, 82.006-M-1117 to 82.006-M-1120, 82.006-M-1064, 82.006-M-1070 to
82.006-M-1073, 82.006-M-1121 to 82.006-M-1125, 82.006-M-1065, 82.006-M-1074, 82.006-M-1112,
82.006-M-1113, 82.006-M-1126 to 82.006-M-1131, 82.006-M-1068, 82.006-M-1114, 82.006-M-1132
to 82.006-M-1135, 82.006-M-1075, 82.006-M-1115, 82.006-M-1116, 82.006-M-1136, 82.006-M-1069,
82.006-M-1137, 82.006-M-1138); 2 ad ♂♂ [1 SA, 1 AL], Shangi, 5 Jan. 1985, 6 Jan. 1985, leg. Benny
Baeten and Luc Janssens (RMCA: 85.006-M-0517, 85.006-M-0518).
Epomophorus minimus Claessen & De Vree, 1991
DEMOCRATIC REPUBLIC OF THE CONGO: 3 ♀♀, 1 ♂ [4 SA], Kitombongo [= Kintobongo],
24 Sep. 2009, 25 Sep. 2009, 26 Sep. 2009, leg. Alexandre Hassanin, Nicolas Nesi, Célestin Pongombo,
Peter Vallo and Didier Tshikung (MNHN: ZM-2011-783 to ZM-2011-786); 1 ♀ [SA], Luena,
21 Sep. 2009, leg. Alexandre Hassanin, Nicolas Nesi, Célestin Pongombo, Peter Vallo and Didier
Tshikung (MNHN: ZM-2011-787).
Epomophorus minor Dobson, 1880
DEMOCRATIC REPUBLIC OF THE CONGO: 1 ? [UN], Albert National Park [= Virunga
National Park], unknown date, leg. unknown collector (INPBC: V720); 1 imm ♂, 1 ♀ [2 SK],
Albertville [= Kalemie], 4 Mar. 1947, 5 Jun. 1947, leg. unknown collector (RBINS: 9065, 9066); 1 ?
[UN], Bunyenzi, unknown date, leg. unknown collector (INPBC: W1515); 1 juv ♂ [AL], Congo da
Lemba, between 13 Jan. 1900 and 22 Jan. 1900, leg. Paul Van Bellighen (RMCA: 408a); 1 ♀ [SK],
Gandajika, 10 Dec. 1951, leg. unknown collector (RBINS: 14599); 1 foetus ?, 1 ♀ [1 SS, 1 AL],
Kakonda, 8 Oct. 1942, leg. F.L. Hendricks (AMNH: 118866, 118867); 1 juv ♀, 1 juv ♂ [2 AL], Moba,
19 Nov. 1953, 23 Apr. 1954, leg. H. Bomans (RMCA: 26271, 26272); 1 ♀ [SK], Rutshuru, 1 Jul. 1937,
leg. unknown collector (RBINS: 4788).
Epomophorus wahlbergi (Sundevall, 1846)
BURUNDI: 1 ♀ [SB], Giserama, Colline, 31 Jul. 1991, leg. Julian C. Kerbis Peterhans (FMNH:
148972); 1 imm ♂ [SA], Kigamba, 15 Jul. 1946, leg. Jacques Verschuren (RBINS: 19928); 4 ♂♂
[3 AL, 1 SO], Mabaye [= Mabayi], 20 Dec. 1970, 10 Dec. 1971, leg. P.L. Niort (MHNG: 1499.092,
1499.093, 1499.090, 1499.091); 1 ♀ [SB], Mumushwizi Valley, 23 Aug. 1993, leg. Jay L. Udelhoven
(FMNH: 155942).
138
VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
DEMOCRATIC REPUBLIC OF THE CONGO: 1 ad ♂ [SA], unknown locality, 17 Oct. 1950, leg.
The Antwerp Zoo (RBINS: 6939); 3 ad ♀♀, 2 ad ♂♂, 1 ad ?, 1 sad ?, 1 juv ?, 1 ♂, 1 ? [2 SS, 3 SA,
2 AL, 3 SK], Banana, 1 Jan. 1889, leg. P. Hesse, 25 Jul. 1920, between 26 Jul. 1920 and 26 Aug. 1920,
leg. Henri Schouteden, 9 May 1910, leg. Abel Gruvel, prior to 3 Jun. 1936, leg. Schwetz (SMF: 2510,
RMCA: 6539, 6540, MNHN: ZM-MO-1911-1565, ZM-MO-1911-1565A, ZM-MO-1911-1565B, ZMMO-1911-1565C, RMCA: 6541, 13303, 13304); 1 ad ♀, 1 juv ♂ [2 SK], Banana or Netonna, unknown
date, prior to 31 Dec. 1889, leg. P. Hesse (SMF: 2511, 2512); 1 ♀ [UN], Bas Congo (former) Province
[= Kongo Central Province], prior to 31 Dec. 1897, leg. Alphonse Cabra (RMCA: 73); 1 sad ? [SK],
Biakabomba, between 1 Jan. 1931 and 31 Dec. 1931, leg. Betu Nsankulu (RMCA: 79.032-M-0019);
1 ? [SK], Bokungu, between 1 Jan. 1949 and 9 Sep. 1949, leg. L. Dupuis (RMCA: 18844b); 1 ad ♂,
1 sad ♀, 3 ♀♀, 1 ♂, 30 ?? [7 SS, 1 SA, 21 AL, 1 SK, 6 SO], Boma, 5 Jul. 1920, between 1 Oct. 1920
and 31 Oct. 1920, leg. Henri Schouteden, 7 Oct. 1913, leg. Styczynski, between 1 Apr. 1936 and
30 Sep. 1936, leg. Schwetz, prior to 25 Nov. 1937, between 1 Aug. 1928 and 31 Aug. 1928, leg. Gerard,
prior to 1 Dec. 1950, leg. Van Riel, prior to 26 May 1933, leg. Colsoulle, prior to 30 Mar. 1934, leg.
Institute of Tropical Medicine “Prince Leopold”, prior to 4 Aug. 1937, leg. Dartevelle (RMCA: 6532,
6538, 2484a, 2484b, 13705, 13706, 16261, 16204, 16205, 20177, 20178, 14182, 12125a to 12125d,
12127j to 12127m, 12251 to 12256, 13946, 13948 to 13950, 13952, 13953, 13955 to 13958); 1 juv ♀
[SK], Byonga, 22 Jan. 1965, leg. Alexandre Prigogine (RMCA: 33285); 1 ad ♂ [SS], Cape Lopez,
10 Feb. 1915, leg. Herbert Lang, James Paul Chapin and The American Museum Congo Expedition
(AMNH: 48662); 1 ? [AL], Congo da Lemba, prior to 22 Jan. 1900, leg. Paul Van Bellighen (RMCA:
409); 1 imm ♀ [UN], Gandajika, between 1 Oct. 1948 and 31 Oct. 1948, leg. P. de Francquen (RBINS:
13108); 1 ? [UN], Ibembo, unknown date, leg. unknown collector (RMCA: 20278); 1 ad ♂ [SK], Island
of Idjwi , 19 Jul. 1965, leg. Alexandre Prigogine (RMCA: 33625); 1 ♂ [SO], Kabanga, 12 Jan. 1952,
leg. L. Spaute (RBINS: 13329); 1 sad ♀ [SK], Kamueshi [= Kamuesha], between 1 Jan. 1931 and
31 Dec. 1931, leg. Betu Nsankulu (RMCA: 79.032-M-0018); 2 ?? [2 UN], Kananga, 31 Mar. 1924,
leg. P. Callewaert (NMBA: 3800, 7695); 3 ♀♀ [3 SK], Kanzenze Mission, 1 Oct. 1948, leg. unknown
collector (RBINS: 13280 to 13282); 2 sad ♀♀ [1 SA, 1 AL], Lubondai-Tshimbulu [= LubondaieTshimbulu], between 1 Jan. 1953 and 3 Feb. 1955, leg. S. Marks (RMCA: 22819, 22820); 1 sad ♂
[AL], Lubumbashi, 1 Apr. 1926, leg. J. Bequaert (RMCA: 16209); 3 ad ♀♀, 2 ad ♂♂, 1 ad ?, 1 juv ♂,
3 ♀♀, 11 ?? [14 SS, 3 SA, 1 AL, 2 SK, 1 SO], Luluabourg [= Kananga], 1 Jan. 1963, 18 Apr. 1963,
leg. Jan Deheegher, 21 Sep. 1963, 22 Sep. 1963, 1 Feb. 1964, 14 Feb. 1964, 22 Feb. 1964, 2 Mar. 1964,
18 Apr. 1964, 6 May 1964, 10 May 1964, 14 Dec. 1964, 21 Feb. 1965, 27 Feb. 1965, 16 Mar. 1965,
22 Mar. 1965, 3 Aug. 1968, between 1 Jan. 1964 and 31 Dec. 1964, leg. Antoon Emeric Marcel
De Roo, 26 Nov. 1922, leg. unknown collector (RMCA: 31554, AMNH: 207736, RMCA: 32651,
AMNH: 207737, 206843, RMCA: 33409, 33644, 31550, 32387, 32378, 33407, 32661, AMNH: 207738,
RMCA: 33410, 32662, MNHN: ZM-MO-1960-3859, ZM-MO-1960-3860, RMCA: 33408, 35609,
32650, 34405a); 1 ad ♀, 3 ad ♂♂, 1 ♀, 1 ♂ [1 SS, 4 SK, 1 SO], Moanda [= Muanda], 18 Aug. 1920,
21 Aug. 1920, between 19 Aug. 1920 and 21 Aug. 1920, leg. Henri Schouteden (RMCA: 6534, 7290,
6533, 6535, 6537, 6536); 1 ♀, 1 ? [2 SO], Mulongo, unknown date, leg. Gerard (RMCA: 10515,
10517); 1 ♂ [SS], Mungamba, 17 Jan. 1982, leg. Betu Nsankulu (RMCA: 83.025-M-0005); 3 ♀♀,
1 ♂, 1 ? [5 AL], 4 km from Muvule Cave, 8 Oct. 2009, 9 Oct. 2009, leg. Alexandre Hassanin, Nicolas
Nesi, Célestin Pongombo, Peter Vallo and Didier Tshikung (MNHN: ZM-2011-799 to ZM-2011-803);
2 juv ♂♂ [2 SK], Ndemba, 28 Mar. 1977, leg. Betu Nsankulu (RMCA: 77.038-M-0002, 77.038-M0003); 3 ♂♂ [3 AL], Nyabiondo, 29 Dec. 1990, 30 Dec. 1990, leg. J.P. Lubula Bulambo and Ngenge
Masumbuko Kamitongo (ZMA: MAM.24349 to MAM.24351); 2 ?? [2 UN], Rutshuru, unknown date,
leg. unknown collector (INPBC: V1346, V1636.2); 2 ♀♀, 3 ♂♂, 1 ? [5 SS, 1 SO], St. Jos. (Misson)
[= Mikalaya], 23 Nov. 1922, 22 Dec. 1923, 7 Aug. 1924, 30 Nov. 1924, leg. Richard Callewaert
(AMNH: 86259, 86255, 86256, 86258, 86251, 86252); 4 ?? [4 SS], St. Joseph de Luluabourg Mission
[= Mikalaya], 5 May 1922, leg. Richard Callewaert (RMCA: 7537 to 7540); 1 sad ?, 2 ♀♀, 2 ♂♂
[2 SS, 1 SK, 2 SO], Tshibashi (River), 27 Feb. 1976, between 1 Feb. 1976 and 27 Feb. 1976, leg.
139
European Journal of Taxonomy 382: 1–327 (2017)
Betu Nsankulu (RMCA: 76.015-M-0002 to 76.015-M-0006); 1 ? [AL], Tshikapa, between 1 Jan. 1921
and 31 Dec. 1921, leg. Henri Schouteden (RMCA: 16231); 2 ♀♀ [1 AL, 1 SB], Washiha Forest,
19 Jul. 2001, leg. Michael H. Huhndorf (FMNH: 171630, 171717).
RWANDA: 1 juv ♀ [AL], Astrida [= Butare], prior to 15 Jul. 1951, leg. The Antwerp Zoo (RMCA:
20496).
Epomops Gray, 1866
DEMOCRATIC REPUBLIC OF THE CONGO: 2 ♀♀ [1 SA, 1 AL], Kikwit, 27 Jun. 2015, 29 Jun. 2015,
leg. Prescott Musaba, Moïse Bipoo, Charle Andabhati, Benjamin Kirongozi and Kambale Karupao
(UNIKIS: KKT2847, KKT2860); 2 ♂♂ [2 AL], Rethy, 20 Oct. 2015, leg. Anne Laudisoit et al.
(UNIKIS: ITU703, ITU704).
Epomops dobsonii (Bocage, 1889)
DEMOCRATIC REPUBLIC OF THE CONGO: 1 imm ♀, 2 ♀♀ [1 AL, 2 UN], Elisabethville
[= Lubumbashi], 1 Jan. 1930, leg. unknown collector, 12 Jan. 1956, leg. Félix / Michel Anciaux de
Faveaux, between 1 Jan. 1930 and 31 Dec. 1930, leg. H.J. Bredo (RBINS: 6280, BMNH: 1957.432,
ZMA: MAM.20163); 1 imm ♂ [UN], Kambove, 6 Feb. 1907, leg. S.A. Neave (BMNH: 1907.12.13.1);
1 ♀, 2 ♂♂, 1 ? [4 SS], Kanzenze Mission, prior to 31 May 1939, between 1 May 1939 and 31 May
1939, leg. unknown collector, between 1 May 1939 and 31 May 1939, leg. R.P. Lefébure (RBINS:
1811a, 1811b, 1790, 1790B); 1 ♂ [SS], Kasende, 26 Jun. 1934, leg. R.S. Pierrepont (MCZ: 31738);
1 ? [SS], Katentania, between 1 Jan. 1926 and 31 Jan. 1926, leg. R. Van Saceghem (RMCA: 8243);
1 imm ?, 1 ♀, 5 ♂♂ [5 SS, 2 SN], Kipangaribwe River, 17 Dec. 1947, 3 Jan. 1948, 12 Jan. 1948,
14 Jan. 1948, 19 Jan. 1948, leg. Gaston-François de Witte, between 1 Jan. 1941 and 31 Dec. 1943, leg.
R. Grauvet (RBINS: 10668, 10666, 10664, 10669, 10674, 10675, 10678); 1 ? [AL], Mwera, prior to
26 Sep. 1956, leg. R.P. Th. de Caters (RMCA: 23945); 1 ? [AL], Vinamont Farm, 18 Nov. 1955, leg.
Félix / Michel Anciaux de Faveaux (RMCA: 26267).
RWANDA: 1 ad ♂ [SS], Rubona, 21 Dec. 1966, leg. Félix / Michel Anciaux de Faveaux (RBINS:
16037); 1 ad ♂ [SA], Shangi Schad [= Shangi], 5 Jan. 1985, leg. Benny Baeten and Luc Janssens
(RMCA: 85.006-M-0463).
Epomops franqueti (Tomes, 1860)
DEMOCRATIC REPUBLIC OF THE CONGO: 1 ♂, 3 ?? [1 SS, 2 AL, 1 SO], unknown locality,
unknown date, leg. Prescott Musaba, 12 Mar. 1956, leg. J.J. Laarman, between 1 Jan. 1985 and
31 Dec. 1986, leg. WHO and Marc Colyn, prior to 28 Aug. 1978, leg. Betu Nsankulu
(RMCA: a1.097-M-3172, RMNH: MAM.16345, RMCA: 88.011-M-0010, 78.020-M-0002); 1 ♀, 1 ♂
[2 AL], Aketi, 23 Jun. 2015, 25 Jun. 2015, leg. Vlir/Aketi (UNIKIS: AKETI616, AKETI617); 1 ad ♂,
1 juv ♀, 3 juv ♂♂, 3 ♀♀, 2 ?? [7 SS, 1 AL, 1 SO, 1 UN], Avakubi, 30 Sep. 1912, 1 Oct. 1912,
30 Oct. 1912, 31 Oct. 1912, leg. Cuthbert Christy, 24 Sep. 1913, 27 Mar. 1914, 14 Apr. 1914, 28 Apr. 1914,
13 May 1914, 23 May 1914, leg. Herbert Lang, James Paul Chapin and The American Museum Congo
Expedition (RMCA: 1421, AMNH: 48730, 48728, 48731, 48727, 48658, 48729, RMCA: 1611, 1599,
1600); 1 ♀ [UN], near Avakubi , between 8 Dec. 1974 and 15 Dec. 1974, leg. Michael D. Gallagher
(HZM: 31.7858); 1 juv ♂, 1 ♂ [2 SS], Bakwakenge [= Bakwa-Kenge], 20 Aug. 1979, 12 Oct. 1979,
leg. Betu Nsankulu (RMCA: 80.013-M-0007, 80.013-M-0010); 8 ♀♀, 2 ♂♂ [5 AL, 5 SK], Baliko,
11 Jun. 2012, 12 Jun. 2012, 13 Jun. 2012, 15 Jun. 2012, leg. William (“Bill”) T. Stanley, prior to
31 Jan. 2007, leg. Guy-Crispin Gembu Tungaluna (FMNH: 219582 to 219586, RMCA: a1.097-M-0417
to a1.097-M-0421); 1 foetus ?, 1 ♀ [1 UN, 1 SB], Bambesa, 3 Jun. 1938, leg. J.M. Vrijdagh (RMCA:
14797a, 14797b); 1 ? [SO], Bas Congo (former) Province [= Kongo Central Province], prior to
140
VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
18 Apr. 1952, leg. Richard (RMCA: 22716); 1 ♀ [SS], Befale, 1 Sep. 1927, leg. J. Ghesquière (RMCA:
9342); 1 ♂ [SS], Bena Mbala, 12 Jan. 1984, leg. Betu Nsankulu (RMCA: 84.035-M-0023); 1 juv ?, 1 ♀
[2 SS], Bene Longo, 23 Aug. 1978, 12 Oct. 1979, leg. Betu Nsankulu (RMCA: 80.013-M-0011, 79.022M-0003); 1 imm ♂ [UN], Beni, unknown date, leg. Rudolf Grauer (NMW: 4879); 1 ♂ [SS], Beyi Kalonji, 22 Feb. 1980, leg. Betu Nsankulu (RMCA: 80.020-M-0002); 2 ?? [2 SS], Boangi, 17 Apr. 1954,
leg. P. Verbuyt (RMCA: 22186, 22187); 6 ?? [5 AL, 1 SO], Boende, 25 Sep. 1939, leg. E.P. Wijnant,
between 1 Jan. 1967 and 31 Dec. 1967, between 1 Jan. 1966 and 31 Jan. 1966, leg. P. Lootens (RMCA:
17335, 34163, 38649, 38654 to 38656); 1 ♀, 1 ♂, 2 ?? [4 SS], Bokuma, 1 Oct. 1950, 2 Oct. 1950,
4 Oct. 1950, between 1 Oct. 1950 and 31 Oct. 1950, leg. P. Lootens (RMCA: 20220, 20219, 20221,
20393); 2 ?? [1 SS, 1 SO], Bokungu, prior to 20 Apr. 1950, between 1 Jan. 1949 and 9 Sep. 1949, leg.
L. Dupuis (RMCA: 18844a, 19659); 1 ? [SS], Boma, unknown date, leg. J. Rodhain (RMCA: 7640); 10
♀♀, 3 ♂♂ [3 SA, 10 AL], Bomane [= Bomaneh], 21 May 2010, 30 May 2010, leg. Guy-Crispin Gembu
Tungaluna, Prescott Musaba and José Akaibe (UNIKIS: CRT2131 to CRT2133, CRT2142, CRT2173,
CRT2176, CRT2180, CRT2183, CRT2198, CRT2199, CRT2063, CRT2065, CRT2066); 1 ♂ [AL],
Bombanga - Bumba, 7 Aug. 1986, leg. WHO and Marc Colyn (RMCA: 88.011-M-0011); 2 ?? [2 SO],
Bombo Lumene, 20 Aug. 1992, leg. Tshinyenye (RMCA: 92.119-M-0005, 92.119-M-0006); 1 ? [SO],
Bongale, 22 Mar. 1951, leg. P. Lootens (RMCA: 20559); 1 ♀ [AL], Bonzale, 13 Jul. 1979, leg. Frieder
Meier (MHNG: 1870.042); 1 juv ?, 14 ?? [14 SS, 1 SO], Buta, 11 Mar. 1935, 13 Jan. 1936, 19 Jan. 1936,
12 Jan. 1944, 20 Jun. 1949, prior to 3 Oct. 1938, prior to 20 Dec. 1927, prior to 16 Jun. 1937, prior to
15 Jul. 1935, between 1 Jun. 1934 and 30 Jun. 1934, between 1 Jan. 1932 and 31 Dec. 1932, leg. Jozef
Hutsebaut (RMCA: 12859, 17304, 13307, 13306, 18835, 11827, 12565 to 12569, 12858, 13854, 9388,
14841); 2 ♀♀, 5 ♂♂ [7 SS], Byonga, 24 Nov. 1962, 11 Jan. 1964, 12 Jan. 1964, 28 Jan. 1965,
22 Feb. 1965, 21 Sep. 1966, 23 Sep. 1966, leg. Alexandre Prigogine (RMCA: 32400, 32401, 34799,
33284, 34798, 31402, 33286); 1 ♂, 1 ? [2 SS], Dimbelenge, 27 Jun. 1980, 28 Jun. 1980, leg. Betu
Nsankulu (RMCA: 80.042-M-0006, 80.042-M-0005); 8 ♀♀, 7 ♂♂ [4 SA, 11 AL], Djabir, 8 May 2011,
9 May 2011, 12 May 2011, 13 May 2011, leg. Prescott Musaba and Jonathan Kosele (UNIKIS: DJB148,
DJB150, DJB151, DJB168, DJB002, DJB005, DJB021, DJB059, DJB066, DJB069, DJB072 to DJB075,
DJB171); 8 ♀♀, 7 ♂♂ [1 SS, 14 AL], Epulu, 16 Jun. 1958, leg. Station de domestication des okapis
Epulu, 25 Aug. 2015, 26 Aug. 2015, 27 Aug. 2015, 29 Aug. 2015, 30 Aug. 2015, 31 Aug. 2015, leg.
André Malekani (RMCA: 27751, UNIKIS: EPLU631, EPLU633, EPLU634, EPLU653, EPLU655 to
EPLU657, EPLU661, EPLU678, EPLU692, EPLU693, EPLU697, EPLU698, EPLU701); 1 ♀ [SB],
2 km W of Epulu, 29 Jun. 1991, leg. Julian C. Kerbis Peterhans (FMNH: 149391); 18 ?? [1 AL, 17 UN],
Faradje, unknown date, leg. unknown collector (UNIKIS: CRT801/G63, CRT802 to CRT805, CRT807,
CRT808/G62, CRT809 to CRT812, CRT814, CRT816, CRT817, CRT818/G64, CRT819/G61, CRT820,
CRT821/G70); 1 ad ♂ [SS], Fort Beni [= Vieux Beni], unknown date, leg. R.E. Dent (BMNH:
1906.12.4.59); 1 imm ♂ [UN], Gangala-na-Bodio, 31 May 1948, leg. unknown collector (RBINS:
11659); 1 ? [SS], Gemena, 26 Jun. 1959, leg. Jacobs (RMCA: 28087); 1 ? [SS], Ibembo, 17 Sep. 1950,
leg. Jozef Hutsebaut (RMCA: 20204); 1 ad ♂ [AL], Ikela, prior to 4 Jan. 1958, leg. P. Lootens (RMCA:
27016); 2 ♀♀, 3 ♂♂ [5 UN], Île Mai [= Mai Island], 20 Feb. 2012, 29 Mar. 2012, 31 Mar. 2012, leg.
Prescott Musaba, Tembele and Marc Boketshu (UNIKIS: CHIMA13, CHIMA31 to CHIMA33,
CHIMA39); 6 ♀♀, 4 ♂♂, 1 ? [4 AL, 7 UN], Île Mbiye [= Mbiye Island], 20 Feb. 2012, 14 Apr. 2012,
19 Apr. 2014, 20 Apr. 2014, between 31 Mar. 2012 and 13 Apr. 2012, leg. Prescott Musaba, 16 Jan. 2012,
18 Feb. 2012, 31 Mar. 2012, leg. Prescott Musaba, Tembele and Marc Boketshu (RMCA: a1.097-M-3178,
UNIKIS: CHIM06, CHIM08, CHIM05/14, RMCA: a1.097-M-3176, a1.097-M-3177,
UNIKIS: CHIM010/14, CHIM09/14, CHIM46, CHIM49, RMCA: a1.097-M-3179); 1 ♂ [SS], Inkongo
[= Inkongo-Kakese], unknown date, leg. unknown collector (BMNH: 1917.1.21.2); 1 juv ♂, 3 ♂♂, 1 ?
[2 SA, 3 AL], Irangi, 16 Mar. 1964, 17 Aug. 1964, leg. P. Kunkel, 16 Oct. 1990, 17 Oct. 1990, leg. Wim
Bergmans (SMF: 31835, ZMA: MAM.24190, SMF: 31834, ZMA: MAM.24191, MAM.24192); 4 ♀♀
[4 UN], Isai River, between 16 Dec. 1974 and 23 Dec. 1974, leg. Michael D. Gallagher (HZM: 20.7833,
23.7836, 29.7856, 32.7859); 1 ♀ [SS], Isoga, 17 Jan. 1964, leg. Alexandre Prigogine (RMCA: 32402);
141
European Journal of Taxonomy 382: 1–327 (2017)
3 ♀♀, 1 ♂ [3 SS, 1 AL], Ituri River, unknown date, leg. unknown collector, unknown date, leg. Howard
de Walden (BMNH: 1930.11.11.66 to 1930.11.11.69); 3 ♀♀ [3 UN], Ituri River, Bridge, between
8 Dec. 1974 and 15 Dec. 1975, between 8 Dec. 1974 and 15 Dec. 1974, leg. Michael D. Gallagher
(HZM: 16.7829, 25.7838, 22.7835); 1 ♂ [SS], Kagada, 20 Jul. 1952, leg. Alexandre Prigogine (RMCA:
21952); 1 juv ♂, 5 ♀♀, 9 ♂♂ [15 SS], Kamituga, 24 Jul. 1950, 4 Jul. 1954, 24 Feb. 1960, 25 May 1960,
9 Nov. 1962, 26 Nov. 1962, 26 Jul. 1963, 15 Nov. 1963, 11 Mar. 1966, 21 Mar. 1967, 25 Mar. 1967,
26 Mar. 1967, 27 Mar. 1967, 11 Nov. 1968, leg. Alexandre Prigogine (RMCA: 34209, 35512, 31663,
35069, 28886, 20431, 35066, 35068, 30556, 35072, 31662, 31403, 35067, 23424, 31346); 1 ♂ [SS],
Kamueshi [= Kamuesha], 3 Apr. 1979, leg. Betu Nsankulu (RMCA: 79.032-M-0016); 1 ♀, 5 ♂♂, 3 ??
[8 SS, 1 SK], Kasai, 13 Dec. 1981, 17 Jan. 1982, 10 Feb. 1982, 17 Feb. 1982, prior to 12 Aug. 1983, leg.
Betu Nsankulu (RMCA: 83.025-M-0012, 83.025-M-0017, 83.025-M-0018, 83.025-M-0014, 83.025-M0016, 83.025-M-0013, 83.025-M-0011, 83.025-M-0015, 83.025-M-0024); 1 ♀ [SS], Katabayi,
10 Apr. 1979, leg. Betu Nsankulu (RMCA: 79.032-M-0017); 1 ? [SS], Katende Falls, 2 Aug. 1957, leg.
Vaillemans (RMCA: 26554); 2 juv ??, 1 ♀ [3 AL], Katshungu, 30 Jan. 1950, leg. Laurent (RMCA:
79.010-M-0001 to 79.010-M-0003); 1 ♂ [SS], Keseki, prior to 23 Jul. 1946, leg. N’Kele (RMCA:
17400); 3 ♀♀ [3 SA], Kikole-Pecheur, 17 May 1980, leg. M. Mutashia (SMF: 58070 to 58072);
1 imm ? [UN], Kimpoko [= Kimpoko-Nsele], 22 Jun. 1979, leg. Tsh Mutashiya (NMW: 25774); 1 ♂
[SS], Kinda, 17 Aug. 1978, leg. Betu Nsankulu (RMCA: 79.022-M-0004); 1 ♂ [SS], Kingulube,
4 Dec. 1968, leg. Alexandre Prigogine (RMCA: 35506); 1 juv ♂, 5 ♀♀, 1 ? [1 SS, 1 AL, 1 SO, 4 UN],
Kinshasa, unknown date, leg. Matashiya-Tshinyenye, 2 Jul. 1909, leg. Herbert Lang, James Paul Chapin
and The American Museum Congo Expedition, 21 Apr. 1964, leg. Jo Van Orshoven, between 15 Jan. 1975
and 26 Jan. 1975, leg. Michael D. Gallagher (FMNH: 125133, AMNH: 48712, ZMA: MAM.11169,
HZM: 26.7853, 27.7854, 33.7860, 8.7807); 2 ?? [2 UN], Kinshasa City Province, between 1 Apr. 1999
and 30 Apr. 1999, leg. unknown collector (SMF: 93241, 93242); 1 ad ♀ [UN], Kinzambi Mission,
26 Jul. 1995, leg. Herwig Leirs et al. (MSB: 274829); 3 ♀♀, 3 ♂♂, 1 ? [5 AL, 2 UN], Kisangani,
22 May 1986, between 1 Apr. 1986 and 30 Apr. 1986, leg. Ndey B. Ifuta, between 1 Sep. 1985 and
30 Sep. 1985, leg. Marc Colyn, between 6 Jan. 1975 and 14 Jan. 1975, leg. Michael D. Gallagher
(RMCA: 87.010-M-0001, 87.010-M-0011, 87.010-M-0012, 86.024-M-0003, 86.024-M-0004, HZM:
12.7811, 13.7812); 3 ♀♀ [3 SS], Kitutu, 6 Feb. 1964, 14 Feb. 1964, 1 Mar. 1964, leg. Alexandre
Prigogine (RMCA: 32403 to 32405); 1 ? [SA], Kivu region, 16 Mar. 1964, leg. P. Kunkel (SMF: 31833);
1 ? [SA], Koloka, between 1 Jun. 1911 and 30 Jun. 1911, leg. H. Schubolz (SMF: 6357); 4 ♀♀, 7 ♂♂,
1 ? [7 SA, 5 AL], Kona, 12 May 2010, 13 May 2010, 15 May 2010, 16 May 2010, 17 May 2010, leg.
Guy-Crispin Gembu Tungaluna, Prescott Musaba and José Akaibe (UNIKIS: CRT1917, CRT1921,
CRT1929, CRT1939, CRT1940, CRT1978, CRT1984, CRT1999, CRT2000, CRT2011, CRT2012,
CRT2014); 1 ? [SK], Kongolo Island, between 1 Dec. 1977 and 31 Dec. 1977, leg. Marc Colyn (RMCA:
90.042-M-0229); 1 juv ♀, 1 ♂ [2 SS], Kunungu, prior to 25 Mar. 1929, between 1 Jan. 1922 and
31 Dec. 1922, leg. Ngwe (RMCA: 7773, 9711); 1 ♀, 1 ♂ [2 AL], Lai Lay [= Lailai], 28 Apr. 1992, leg.
Ngenge Masumbuko Kamitongo (ZMA: MAM.24562, MAM.24563); 6 ♂♂, 1 ? [1 SS, 1 AL, 2 SK,
2 SO, 1 UN], Léopoldville [= Kinshasa], unknown date, leg. Vandenbranden, 11 Apr. 1912, 16 Apr. 1912,
leg. Cuthbert Christy, 12 Nov. 1945, leg. Jean Jacques Deheyn, 16 Jun. 1946, leg. M. Henrion, 18 May
1945, leg. Lepersonne, between 1 Feb. 1942 and 28 Feb. 1942, leg. Becquet (RMCA: 5381, 1538f,
17616b, 1538b, 17798, 17336, 17356b); 1 ♀ [SA], Lieki, 1 May 2010, leg. Guy-Crispin Gembu
Tungaluna, Prescott Musaba and José Akaibe (UNIKIS: CRT2036); 1 ad? ♂, 18 ?? [19 SS], Lovanium
University [= Kinshasa], 1 Apr. 1962, 5 Apr. 1962, 7 May 1962, 26 May 1962, 5 Jun. 1962, 12 Jun. 1962,
leg. De Bont, 10 Jan. 1962, 24 Sep. 1962, 24 Nov. 1962, 5 Jan. 1963, 3 May 1963, 7 May 1963,
16 Jul. 1963, 12 Apr. 1964, 19 Apr. 1964, 21 Apr. 1964, between 1 Apr. 1963 and 20 Mar. 1964, leg.
De Bont and Jo Van Orshoven (RMCA: 31194, 32600, 32598, 31196, 32590, 32599, 32597, 32593,
32595, 31193, 32592, 32589, 31190, 31191, 31195, 31192, 32591, 32594, 32596); 1 ♀ [SS], Luebo,
19 Dec. 1921, leg. Henri Schouteden (RMCA: 6646); 2 ♂♂ [1 SS, 1 AL], Lukolela, 25 Jul. 1930, leg.
Franklin Edson, 25 Jul. 1930, leg. James Paul Chapin (AMNH: 86765, 86906); 1 juv ♂, 1 juv ?, 2 ♀♀
142
VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
[4 SS], Lukonga, 7 Oct. 1978, 2 Aug. 1979, 6 Aug. 1979, leg. Betu Nsankulu (RMCA: 80.013-M-0008,
80.013-M-0009, 80.013-M-0012, 79.022-M-0002); 1 ♂, 1 ? [2 SS], Lulua [= Kananga], 2 May 1976,
leg. Betu Nsankulu (RMCA: 77.015-M-0001, 77.015-M-0002); 1 ad ♂, 4 ♀♀, 3 ♂♂, 32 ?? [27 SS,
12 AL, 1 SO], Luluabourg [= Kananga], unknown date, leg. Richard Callewaert, unknown date,
18 Apr. 1963, leg. Jan Deheegher, 22 Sep. 1963, 11 Jan. 1964, 4 May 1964, 10 May 1964, 2 Sep. 1964,
14 Nov. 1964, 14 Feb. 1965, 20 Feb. 1965, 21 Feb. 1965, 22 Feb. 1965, 25 Feb. 1965, 27 Feb. 1965,
4 Mar. 1965, 6 Mar. 1965, 8 Mar. 1965, 19 Mar. 1965, 12 Apr. 1965, 30 Apr. 1965, 8 May 1965, 22 May
1965, 15 Sep. 1969, between 1 Nov. 1962 and 30 Nov. 1962, between 1 Jan. 1964 and 31 Dec. 1964, leg.
Antoon Emeric Marcel De Roo (BMNH: 1926.7.6.55, 1926.7.6.56, RMCA: 31553, 31555, 31556,
31558, 31559, 32653, 32654, AMNH: 207739, RMCA: 33404, 33647, 33390, 33646, 35796, 31551,
31552, 33403, 32690, 33391 to 33393, 33648, 32663 to 32665, 33394 to 33396, 33405, 33397 to 33399,
32652, 33400 to 33402, 33406, 34405b, 31548); 1 ♀ [AL], Lwana River, 12 Jun. 1992, leg. Ngenge
Masumbuko Kamitongo (ZMA: MAM.24561); 1 ♂ [UN], Malandje [= Malandji], unknown date, leg.
Mechow (ZMB: 10031); 4 ♀♀, 3 ♂♂ [1 SS, 2 SK, 1 SO, 3 UN], Mambaka, unknown date, 17 Jan. 1913,
20 Jan. 1913, 21 Jan. 1913, 22 Jan. 1913, 2 Jan. 1914, 20 Jan. 1914, leg. Cuthbert Christy (BMNH:
1914.4.8.2, RMCA: 1601, 3449, 1602, 3448, 1603, 1604); 1 juv ♀, 1 juv ♂, 2 ♀♀ [4 AL], Mandima,
24 Jul. 1976, 29 Jul. 1976, 1 Aug. 1976, leg. Frieder Meier (MHNG: 1872.039, 1872.036 to 1872.038);
3 ad ♀♀, 1 ad ♂, 4 sad ♂♂, 4 juv ♀♀, 7 juv ♂♂, 53 ♀♀, 45 ♂♂, 5 ?? [3 SA, 23 AL, 96 SK], Masako,
22 May 1986, 24 Nov. 1987, 25 Nov. 1987, 26 Nov. 1987, 27 Nov. 1987, 30 Apr. 1989, 25 Aug. 1989,
26 Aug. 1989, 27 Aug. 1989, 28 Aug. 1989, 30 Aug. 1989, 14 Sep. 1989, 21 Nov. 1989, 12 Dec. 1989,
17 Dec. 1989, 22 Dec. 1989, 28 Dec. 1989, 7 Jan. 1990, 21 Jan. 1990, 25 Jan. 1990, 26 Jan. 1990,
27 Jan. 1990, 28 Jan. 1990, 29 Jan. 1990, 10 Feb. 1990, 13 Feb. 1990, 14 Feb. 1990, 16 Feb. 1990,
17 Feb. 1990, 27 Feb. 1990, 28 Feb. 1990, 17 Mar. 1990, 23 Mar. 1990, 25 Mar. 1990, 27 Oct. 1990,
30 Oct. 1990, between 1 Oct. 1989 and 31 Oct. 1989, leg. Ndey B. Ifuta, 12 Jun. 2015, leg. Prescott
Musaba, Moïse Bipoo, Charle Andabhati, Benjamin Kirongozi and Kambale Karupao (RMCA: 93.079M-0162 to 93.079-M-0164, UNIKIS: CHMS068, RMCA: 93.079-M-0136 to 93.079-M-0139, 93.079M-0190 to 93.079-M-0192, 93.079-M-0271, 93.079-M-0193, 93.079-M-0194, 93.079-M-0168, 93.079M-0169, 93.079-M-0140, 93.079-M-0155 to 93.079-M-0161, 93.079-M-0129 to 93.079-M-0135,
87.010-M-0004, 87.010-M-0005, 93.079-M-0141 to 93.079-M-0144, 93.079-M-0262, 93.079-M-0001
to 93.079-M-0008, 93.079-M-0200 to 93.079-M-0209, 93.079-M-0263, 93.079-M-0314, 93.079-M0184, 93.079-M-0195, 93.079-M-0172 to 93.079-M-0174, 93.079-M-0009 to 93.079-M-0014, 93.079M-0210 to 93.079-M-0212, 93.079-M-0185, 93.079-M-0175, 93.079-M-0176, 93.079-M-0015 to
93.079-M-0020, 93.079-M-0213, 93.079-M-0264 to 93.079-M-0266, 93.079-M-0186, 93.079-M-0165,
93.079-M-0177 to 93.079-M-0180, 93.079-M-0196, 93.079-M-0021 to 93.079-M-0025, 93.079-M0187, 93.079-M-0188, 93.079-M-0154, 93.079-M-0166, 93.079-M-0167, 93.079-M-0199, 93.079-M0181, 93.079-M-0145 to 93.079-M-0150, 93.079-M-0189, 93.079-M-0171, 93.079-M-0182, 93.079M-0183, 93.079-M-0197, 93.079-M-0198, 93.079-M-0151 to 93.079-M-0153, 93.079-M-0126 to
93.079-M-0128); 1 ? [AL], Masako km 15 Buta, 22 May 1986, leg. Ndey B. Ifuta (RMCA: 87.010-M0003); 1 ? [SK], Mawambi, 8 Aug. 1913, leg. Cuthbert Christy (RMCA: 3450); 1 ad ♀ [SA],
Mbwambala, 7 Jul. 1995, leg. Herwig Leirs et al. (RMCA: 97.021-M-0748); 3 ad ♀♀, 1 ad ♂, 3 juv
♀♀, 2 juv ♂♂, 2 ♀♀, 1 ♂, 3 ?? [3 SS, 12 UN], Medje, unknown date, 25 Apr. 1914, leg. Cuthbert
Christy, 21 Mar. 1910, 12 Sep. 1910, 14 Sep. 1910, 15 Sep. 1910, 2 Oct. 1910, 11 Oct. 1910, 11 Mar. 1914,
19 Mar. 1914, 22 Mar. 1914, 7 Apr. 1914, leg. Herbert Lang, James Paul Chapin and The American
Museum Congo Expedition (BMNH: 1919.5.8.9, AMNH: 48726, 49205, RMCA: 12398, AMNH:
48713, 48719, 48721, 48722, 48716, 48714, 48715, 48723, RMCA: 4331, AMNH: 51309, RMCA:
12358); 1 embryo ♀, 1 embryo ♂, 1 embryo ? [2 SS, 1 AL], Mistandunga, 23 Jul. 1930, leg. Franklin
Edson, 23 Jul. 1930, leg. James Paul Chapin (AMNH: 86766, 86767, 86907); 1 imm ♀ [UN], Moera,
unknown date, leg. Rudolf Grauer (NMW: 4878); 1 ? [UN], Mongbwalu, unknown date, leg. unknown
collector (RMCA: 15566); 1 ♂ [SO], Mulongo, unknown date, leg. Gerard (RMCA: 10514); 1 ♀, 1 ?
[1 SS, 1 UN], Mutsora, unknown date, leg. unknown collector, 4 Apr. 1951, leg. R. Christiaens
143
European Journal of Taxonomy 382: 1–327 (2017)
(INPBC: W703, RBINS: 13327); 1 imm ♀, 1 ? [1 SS, 1 UN], Mutwanga, unknown date, leg. unknown
collector, 15 Jul. 1946, leg. J. de Wilde (INPBC: W1283, RBINS: 13326); 1 ? [SS], Nganza River,
27 May 1966, leg. Antoon Emeric Marcel De Roo (RMCA: 35277); 1 ad ♀ [SS], Niangara, 7 Oct. 1910,
leg. Herbert Lang, James Paul Chapin and The American Museum Congo Expedition (AMNH: 48717);
1 ad ♀, 2 ad ♂♂ [1 SS, 1 SO, 1 OT], Niapu, 2 Dec. 1913, 4 Dec. 1913, leg. Herbert Lang, James Paul
Chapin and The American Museum Congo Expedition (AMNH: 49196, 48724, 51308); 1 ♀ [SO],
Nshamba, 13 Dec. 1969, leg. Antoon Emeric Marcel De Roo (RMCA: 35798); 7 ♀♀, 6 ♂♂ [1 AL,
3 SK, 9 UN], Obenge, 4 Feb. 2013, 5 Feb. 2013, 7 Feb. 2013, leg. Prescott Musaba and André Malekani
(UNIKIS: TLL351 to TLL353, TLL355, TLL360, TLL362, TLL372, TLL373, TLL378 to TLL380,
TLL394, TLL416); 1 ? [AL], Omaniumdu, between 5 Jul. 1959 and 8 Jul. 1959, leg. Laurent (RMCA:
34353); 2 ♀♀ [2 SS], Panga, between 1 Sep. 1925 and 30 Sep. 1925, leg. Henri Schouteden (RMCA:
9139b, 9140); 2 ?? [2 SS], Paulis [= Isiro], between 1 Jan. 1947 and 31 Dec. 1947, leg. Abbeloos
(RMCA: 17927, 17928); 1 ♂ [UN], Pidigala Nord River, 23 Apr. 1952, leg. H. de Saeger et al. (RBINS:
13501); 2 ♀♀ [1 SK, 1 UN], Pili Pili [= Pilipili], 19 Dec. 1912, 19 Dec. 1913, leg. Cuthbert Christy
(RMCA: 1605, 3447); 2 ♀♀ [2 SK], Quatorze, 28 Jun. 2013, 29 Jun. 2013, leg. William («Bill»)
T. Stanley (FMNH: 222414, 222415); 29 ♀♀, 23 ♂♂ [52 AL], Rubi-Tele, 17 Aug. 2013, 18 Aug. 2013,
19 Aug. 2013, 20 Aug. 2013, 21 Aug. 2013, 22 Aug. 2013, 23 Aug. 2013, 24 Aug. 2013, 25 Aug. 2013,
26 Aug. 2013, 27 Aug. 2013, 28 Aug. 2013, leg. Guy-Crispin Gembu Tungaluna, Prescott Musaba and
André Malekani (UNIKIS: RBTL483 to RBTL485, RBTL496, RBTL500, RBTL501, RBTL506 to
RBTL509, RBTL511, RBTL513, RBTL517 to RBTL519, RBTL523 to RBTL527, RBTL531 to
RBTL533, RBTL535 to RBTL539, RBTL543 to RBTL546, RBTL552, RBTL553, RBTL559 to
RBTL561, RBTL565 to RBTL572, RBTL574, RBTL573, RBTL578 to RBTL582); 2 ?? [1 AL, 1 UN],
Rutshuru, unknown date, leg. unknown collector, 27 May 1955, leg. Gaston-François de Witte
(INPBC: V1626, RMCA: 37115); 1 ? [SS], Semliki River, between 1 Aug. 1932 and 15 Aug. 1932, leg.
Henri Martin Hackars and Georges Marlier (RMCA: 12165); 1 imm ?, 1 foetus ?, 5 ♀♀, 7 ♂♂ [8 SS,
4 AL, 1 SK, 1 UN], Stanleyville [= Kisangani], unknown date, 26 May 1912, 26 Jun. 1912, 2 Jul. 1912,
20 Jul. 1912, prior to 12 Mar. 1913, leg. Cuthbert Christy, 4 May 1915, 1 Jul. 1915, leg. Herbert Lang,
James Paul Chapin and The American Museum Congo Expedition (BMNH: 1914.4.8.1, AMNH: 49206,
RMCA: 1414, 1417, 1415, 1411 to 1413, 1416, 1418, 1420, AMNH: 49203, RMCA: 1530a, 1531); 1 ♀,
1 ♂ [2 AL], Tandala, 12 Jul. 1979, leg. Frieder Meier, 19 Jul. 1979, leg. C. Brian Robbins (MHNG:
1870.040, USNM: 463474); 1 juv ♀, 1 ♀, 1 ♂ [2 SS, 1 UN], Teturi, 6 Feb. 1947, 30 Jun. 1947,
15 Mar. 1948, leg. unknown collector (RBINS: 7044, 7053, 7052); 1 ♂ [UN], Tingasi, unknown date,
leg. Emin Pascha / Pasha / Bey (BMNH: 1887.12.1.26); 1 ♀ [SO], Tshibasha, 19 Sep. 1969, leg. Antoon
Emeric Marcel De Roo (RMCA: 35797); 1 ♀ [SS], Tshibashi (River), 27 Feb. 1976, leg. Betu Nsankulu
(RMCA: 76.015-M-0001); 1 ♀ [SS], Tshibungu, 27 Nov. 1978, leg. Betu Nsankulu (RMCA: 79.022M-0005); 2 ?? [2 SK], Tshopo, between 1 Jun. 1982 and 30 Jun. 1982, leg. Marc Colyn (RMCA: 90.042M-0230, 90.042-M-0231); 1 ? [AL], Tshoppo Falls [= Tshopo Falls], prior to 12 Mar. 1913, leg.
Cuthbert Christy (RMCA: 1530b); 1 ? [AL], Tshuapa District, between 1 Jan. 1960 and 31 Dec. 1964,
leg. P. Lootens (RMCA: 34184); 13 ♀♀, 14 ♂♂ [16 AL, 6 SK, 5 UN], Uma, 15 Aug. 2012, 16 Aug. 2012,
17 Aug. 2012, 18 Aug. 2012, 10 Apr. 2014, 11 Apr. 2014, 12 Apr. 2014, leg. Guy-Crispin Gembu
Tungaluna, Prescott Musaba and André Malekani, 16 Aug. 2012, 18 Aug. 2012, leg. Prescott Musaba
(UNIKIS: UMA535, UMA542, UMA570, UMA572, UMA574 to UMA576, UMA587, UMA606,
UMA608, UMA613, UMA614, UMA631, UM001 to UM003, UM009, UM012, UM019, UM023,
RMCA: a1.097-M-3173, a1.097-M-3175, UNIKIS: UM011, UM015, UM017, RMCA: a1.097-M-3174,
UNIKIS: UM020); 1 juv ♀ [SS], Vankerckhovenville [= Nzoro], 17 Apr. 1912, leg. Herbert Lang,
James Paul Chapin and The American Museum Congo Expedition (AMNH: 48720); 12 ♀♀ [12 UN],
Weko Forest, between 27 Dec. 1974 and 3 Jan. 1975, leg. Michael D. Gallagher (HZM: 10.7809,
11.7810, 15.7828, 17.7830 to 19.7832, 21.7834, 30.7857, 35.7862 to 37.7864, 9.7808); 2 ♀♀, 1 ♂
[1 SK, 2 UN], Yaekela, 3 May 2010, 4 May 2010, 6 May 2010, leg. Guy-Crispin Gembu Tungaluna,
Prescott Musaba and José Akaibe (UNIKIS: CRT2374, CRT2395, CRT1867); 3 ♀♀, 2 ♂♂ [2 SS, 3 AL],
144
VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
Yalosemba, 12 Jun. 1979, 13 Jun. 1979, 15 Jun. 1979, leg. C. Brian Robbins (USNM: 463471, 537696,
463472, 537697, 463473); 1 ad ♂, 1 ♂ [1 SS, 1 AL], Yalusaka, 10 Mar. 1958, between 1 Mar. 1957 and
31 Mar. 1957, leg. F. Carbone (RMCA: 27174, 31717); 1 ♂ [AL], Yambombo, 20 Jun. 1979, leg. Frieder
Meier (MHNG: 1870.041); 5 ♀♀, 1 ♂ [6 UN], Yangambi, between 4 Jan. 1975 and 6 Jan. 1975, leg.
Michael D. Gallagher (HZM: 14.7827, 24.7837, 34.7861, 38.7855, 38.7865, 7.7805); 12 ♀♀, 1 ♂, 1 ?
[14 UN], Yoko, unknown date, 18 May 2009, 25 May 2009, 26 May 2009, 23 Sep. 2009, 25 Sep. 2009,
leg. unknown collector, 4 Sep. 2012, 5 Sep. 2012, 7 Sep. 2012, leg. Prescott Musaba, Tembele and Marc
Boketshu, 5 Apr. 2014, leg. Prescott Musaba (UNIKIS: YK034CH, GE241/G16, GE244, GE279/G19,
GE285/G17, GE289, GE290, YK007CH, YK008CH, YK023CH, YK026CH, YK027CH, YK031,
YK033CH).
Hypsignathus H. Allen, 1862
Hypsignathus monstrosus H. Allen, 1862
DEMOCRATIC REPUBLIC OF THE CONGO: 1 foetus ?, 1 ? [1 SK, 1 UN], Alipago, 29 May 1938,
leg. J.M. Vrijdagh (RMCA: 14796a, 14796b); 1 ♀ [UN], Angu, near, 22 Apr. 1906, leg. unknown
collector (BMNH: 1907.7.8.23); 2 juv ??, 2 ♀♀ [4 SS], Arebi Bondo Mabe, 19 Jul. 1925, leg. Henri
Schouteden (RMCA: 8498 to 8501); 1 ad ♀, 9 ad ♂♂, 2 juv ♀♀, 1 juv ♂, 2 imm ♀♀, 3 imm ♂♂, 4 ♂♂,
4 ?? [19 SS, 5 AL, 2 UN], Avakubi, unknown date, 3 Mar. 1914, 11 May 1914, 27 May 1914, leg.
unknown collector, 6 May 1910, 23 Dec. 1913, 24 Feb. 1914, 26 Feb. 1914, 27 Feb. 1914, 28 Feb. 1914,
3 Mar. 1914, 5 Mar. 1914, 6 Mar. 1914, 7 May 1914, 11 May 1914, 12 May 1914, 15 May 1914, 18 May
1914, 27 May 1914, 28 May 1914, 28 Feb. 1915, leg. Herbert Lang, James Paul Chapin and The
American Museum Congo Expedition, 6 Mar. 1914, leg. J.A. Howard (NMW: 18282, RMCA: 12.376,
12376, AMNH: 48646, 48645, 48647 to 48649, 48636, 49279, 48656, 48637, 48639, 48650, MCZ:
17363, AMNH: 48657, 48638, 48651, 48640, MCZ: 17362, AMNH: 48655, 48641, 48653, KU: 125195,
AMNH: 49208, 48643); 1 ♂ [SS], Bafwabaka, 2 May 1910, leg. Herbert Lang (FMNH: 43873); 1 ad
♂, 1 juv ♀, 2 juv ♂♂, 1 ♂, 1 ? [5 SS, 1 AL], Bafwaboka [= Bafwabaka], 2 Mar. 1910, 2 May 1910,
leg. Herbert Lang, James Paul Chapin and The American Museum Congo Expedition (AMNH: 48654,
48631, 48632, 48635, 49280, RMCA: 12360); 2 ♀♀, 1 ♂ [1 AL, 2 SK], Baliko, 10 Jun. 2012,
12 Jun. 2012, leg. William (“Bill”) T. Stanley, prior to 31 Jan. 2007, leg. Guy-Crispin Gembu Tungaluna
(FMNH: 219587, 219588, RMCA: a1.097-M-0413); 1 ♀ [SS], Bamanya, 16 Apr. 1952, leg. Leonet
(RMCA: 21689); 1 juv ?, 2 ♀♀ [3 UN], Bambesa, 29 May 1938, 27 Aug. 1938, leg. J.M. Vrijdagh
(RMCA: 14677, 14678, 14796); 1 ad ♂ [SK], Bambu [= Kilomines], 1 Jan. 1948, leg. unknown
collector (RBINS: 7037); 1 ? [UN], Baraka, unknown date, leg. Rudolf Grauer (ZMB: 67118); 1 ad ♀
[SK], Bela, 14 Apr. 1947, leg. unknown collector (RBINS: 7040); 1 ♂ [SS], Beyi - Kalonji, 18 Feb. 1980,
leg. Betu Nsankulu (RMCA: 80.020-M-0003); 1 ? [UN], Boende, prior to 16 Mar. 1940, leg. R.P. Wynants
(RMCA: 15949); 1 ? [SS], Bokuma, 22 Oct. 1950, leg. P. Lootens (RMCA: 20397); 1 ♂, 2 ?? [3 SS],
Bokungu, 2 Dec. 1948, prior to 25 Jul. 1949, leg. L. Dupuis (RMCA: 18693, 18817, 18818); 6 ?? [3 SS,
3 SO], Bolafa, between 1 May 1956 and 31 May 1956, leg. Stevenart (RMCA: 25841 to 25846); 1 ?
[SS], Bongale, 15 Mar. 1951, leg. P. Lootens (RMCA: 20560); 1 ? [SS], Bukavu, prior to 30 Jun. 1999,
leg. Camerman (RMCA: 99.011-M-0001); 9 ?? [7 SS, 2 UN], Buta, 12 Jun. 1936, 10 May 1940,
2 Aug. 1940, prior to 19 Jan. 1923, prior to 15 Jul. 1935, between 1 Dec. 1934 and 31 Dec. 1934, between
1 Jun. 1934 and 30 Jun. 1934, leg. Jozef Hutsebaut (RMCA: 17321, 13379, 13380, 17320, 12563, 12564,
12671, 12857, 6273); 1 ? [AL], Butembo, between 1 Apr. 1956 and 30 Apr. 1956, leg. P. Dyleff (RMCA:
23956); 1 ♂ [SS], Byonga, 10 Jan. 1972, leg. Alexandre Prigogine (RMCA: 38586); 1 ? [AL], Congo
da Lemba, unknown date, leg. Paul Van Bellighen (RMCA: 492); 1 imm ?, 2 ♀♀, 3 ♂♂, 2 ?? [2 SS,
2 AL, 2 SO, 2 UN], Eala, unknown date, between 1 Jan. 1921 and 31 Dec. 1921, leg. Verlaine, 4 Jan. 1921,
5 Jan. 1921, leg. Henri Schouteden (RMCA: 16422, 6526 to 6528, 6531, 6524, 6524a, 16215); 7 ♀♀,
1 ♂, 9 ?? [7 SS, 6 AL, 1 SK, 3 SO], Epulu, unknown date, leg. Luc Bijnens, 4 Mar. 1957, 9 May 1957,
17 May 1957, 26 May 1958, leg. Station de domestication des okapis Epulu, 21 Sep. 1956, 24 Sep. 1956,
145
European Journal of Taxonomy 382: 1–327 (2017)
between 1 Sep. 1956 and 30 Sep. 1956, leg. Max Poll, 4 Mar. 1957, leg. Heinz Stephan (RMCA: 90.031M-0002, 26610, 25597 to 25599, 25596, 27750, 26609, 27417, 27418, 26611, 26612, 27645 to 27649);
1 ♂ [AL], Epulu, 6 km S, 2 km W, 6 Jun. 1981, leg. John A. Hart (FMNH: 138136); 1 ? [SN], Equateur
Province, between 1 Jan. 1921 and 31 Dec. 1921, leg. Verlaine (RMCA: 38239); 1 ad? ♀ [SS], Eringete
[= Eringite], 19 Sep. 1949, leg. unknown collector (RBINS: 12945); 1 juv ?, 1 ♂, 3 ?? [3 SS, 1 AL,
1 SK], Ibembo, unknown date, leg. J. Rodhain, 10 Sep. 1949, 10 Feb. 1950, 20 Dec. 1951, between
1 Dec. 1952 and 31 Dec. 1952, leg. Jozef Hutsebaut (RMCA: 7642, 19620, 19527, 20744, 21714); 1 ?
[SO], Ikengo, 15 Feb. 1921, leg. Henri Schouteden (RMCA: 6529); 1 ad ? [AL], Inongo, 1 Oct. 1945,
leg. Gustave Gustin (RMCA: 18124); 1 ♂ [AL], Irebu, unknown date, leg. J. Camp (USNM: 61684);
2 ♀♀ [2 SS], Kamituga, 2 Aug. 1954, 10 Jun. 1964, leg. Alexandre Prigogine (RMCA: 32855, 23422);
1 ♀ [AL], Karambi, 3 Apr. 1992, leg. unknown collector (ZMA: MAM.24557); 4 ?? [4 SK], Kasai,
prior to 11 Jul. 1905, leg. La Compagnie du Kassai (RMCA: 738, 738a to 738c); 1 ? [AL], Kasongo,
between 1 Sep. 1959 and 31 Jan. 1960, leg. Benoit (RMCA: 31106); 1 ♂ [SK], Kasongo Baza
[= Kasonga-Banze], between 1 Feb. 1960 and 28 Feb. 1960, leg. Faber (RMCA: 30820); 9 ?? [9 UN],
Kikwit, unknown date, leg. unknown collector (TM: 45595, 45628, 45630, 46063, 46064, 46073,
46074, 46080, 46081); 1 ad+foetus ♀ [AL], Kilo mines [= Kilomines], prior to 15 Dec. 1930, leg. du
Soleil (RMCA: 10775); 1 ♂ [SS], Kilomamensa, unknown date, leg. André Pilette (RMCA: 6143); 1 ?
[UN], Kindu, unknown date, leg. Rudolf Grauer (ZMB: A.4809); 1 ♀, 1 ♂ [1 SS, 1 UN], Kisala
[= Kisala-Kiama], 11 Mar. 1957, leg. J.J. Laarman, 16 Oct. 1920, leg. Henri Schouteden
(RMNH: MAM.16346, RMCA: 6530); 1 ♀ [SS], Kitangua, 13 Dec. 1981, leg. Betu Nsankulu (RMCA:
83.025-M-0007); 1 ♀ [UN], Kona, 12 May 2010, leg. Guy-Crispin Gembu Tungaluna, Prescott Musaba
and José Akaibe (UNIKIS: CRT1916); 1 ♀ [SS], Koteli [= Kotili], 2 Jan. 1925, leg. Henri Schouteden
(RMCA: 8784); 2 ♀♀ [2 SS], Kunungu, prior to 19 Nov. 1937, prior to 1 Aug. 1939, leg. N’Kele
(RMCA: 15509, 14169); 2 ?? [2 SS], Lesse, 4 Nov. 1913, leg. Bonnevie (RMCA: 3236, 3237); 1 ? [UN],
Libenge, 10 Apr. 1938, leg. Leontovitch (RMCA: 14686); 2 ?? [1 SS, 1 SK], Lisala, prior to 19 Jan. 1929,
leg. Babilon (RMCA: 9574, 9575); 1 ♂ [UN], Lobi, 19 Feb. 1906, leg. unknown collector (BMNH:
1907.7.8.21); 1 ? [SS], Lovanium University [= Kinshasa], 9 May 1963, leg. De Bont and Jo Van
Orshoven (RMCA: 32588); 1 ♀ [SS], Luebo, 26 Aug. 1921, leg. Henri Schouteden (RMCA: 6525);
1 imm ♀ [UN], Lukoi River, unknown date, leg. D. Fraser (BMNH: 1927.3.1.17); 1 juv ♂, 2 ♀♀,
3 ♂♂, 2 ?? [6 SS, 2 AL], Lukolela, 6 Aug. 1930, 28 Aug. 1930, 29 Aug. 1930, 31 Aug. 1930, leg.
Franklin Edson, 31 Jul. 1931, leg. Lecocq, 6 Aug. 1930, leg. James Paul Chapin, between 1 Aug. 1923
and 31 Aug. 1923, leg. J. Ghesquière (AMNH: 86764, 86763, RMCA: 11736, AMNH: 86761, 86762,
86946, 86947, RMCA: 7038); 1 ♂ [AL], Lukolela, 4 mi from, 31 Aug. 1930, leg. James Paul Chapin
(AMNH: 80957); 1 ♀, 1 ? [1 SO, 1 UN], Luluabourg [= Kananga], unknown date, leg. Richard
Callewaert, 3 Feb. 1966, leg. Antoon Emeric Marcel De Roo (BMNH: 1926.7.6.48, RMCA: 35275);
2 ♀♀, 3 ♂♂ [1 SS, 4 AL], Mabali, 9 Nov. 1954, leg. James Paul Chapin, between 1 May 1956 and
30 Jun. 1956, leg. Alvin Novick (AMNH: 185262 to 185264, MCZ: 48184, 48185); 2 ♀♀ [2 SK],
Masako, 23 May 1989, leg. Ndey B. Ifuta (RMCA: 93.079-M-0367, 93.079-M-0368); 1 ? [AL], Mpe,
prior to 26 Jan. 1914, leg. Maes (RMCA: 2662); 1 ♀, 1 ? [2 SS], Ndwa, prior to 1 Aug. 1939, between
1 Jan. 1938 and 31 Dec. 1939, leg. Ngwe (RMCA: 15421, 15422); 2 ?? [2 AL], Nepoko Bomili, prior
to 12 Dec. 1946, leg. M. Henrion (RMCA: 18137, 18137a); 1 imm ♂, 3 ?? [1 SS, 3 AL], New Beni
[= Beni], 25 Oct. 1926, leg. James Paul Chapin, between 1 Oct. 1926 and 31 Oct. 1926, leg. James Paul
Chapin, Dewitt L. Sage, Frank P. Mathews and The Ruwenzori-Kivu Expedition (AMNH: 82529, 82357
to 82359); 1 ? [SS], Paulis [= Isiro], 4 May 1951, leg. Abbeloos (RMCA: 20510); 1 ad ♂, 1 ♂ [1 SS,
1 UN], Penge, 21 Apr. 1914, leg. Herbert Lang, James Paul Chapin and The American Museum Congo
Expedition, 21 Apr. 1914, leg. unknown collector (AMNH: 48642, RMCA: 48642); 1 ♀ [AL], RubiTele, 19 Aug. 2013, leg. Guy-Crispin Gembu Tungaluna, Prescott Musaba and André Malekani
(UNIKIS: RBTL514); 2 ad ♀♀, 2 juv ♀♀, 1 ♀ [4 SS, 1 AL], Stanleyville [= Kisangani], 18 Aug. 1909,
16 Dec. 1914, leg. Herbert Lang, James Paul Chapin and The American Museum Congo Expedition,
18 Aug. 1909, leg. unknown collector (AMNH: 48652, 48629, 48630, 49199, MCZ: 17364); 2 ♀♀
146
VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
[2 UN], Tingasi, 29 Oct. 1883, leg. unknown collector (BMNH: 1887.12.1.24, 1887.12.1.25); 1 ♂ [SK],
Tondu, 19 Feb. 1921, leg. Henri Schouteden (RMCA: 7784); 1 ad ♀ [SK], Tschuapa [= Tshuapa],
18 Oct. 1966, leg. unknown collector (RBINS: 15188); 1 ♀ [SS], Uele [River], between 1 Dec. 1927
and 31 Dec. 1927, leg. Jozef Hutsebaut (MCZ: 23935); 1 ♂ [UN], Yaekela, 3 May 2010, leg. GuyCrispin Gembu Tungaluna, Prescott Musaba and José Akaibe (UNIKIS: CRT2388); 2 ad ♀♀, 2 ad ♂♂,
1 juv ♂ [5 SS], Yalokele, 26 Jun. 1953, 15 Aug. 1953, 28 Aug. 1953, leg. L. Dupuis (RMCA: 21912,
21913, 21889, 21890, 21911); 1 emb ♀, 1 ♀ [1 SS, 1 AL], Yalosemba, 21 Jun. 1979, leg. C. Brian
Robbins (USNM: 463475, 537698); 2 ♀♀, 2 ♂♂ [2 SS, 1 SO, 1 UN], Yangambi, 1 Jan. 1938, leg.
unknown collector, 18 Sep. 1939, leg. J.M. Vrijdagh, 25 Oct. 1947, leg. Rodolphe Germain, 7 Aug. 1951,
leg. L. Toussaint (RBINS: 5941, RMCA: 17238, RBINS: 6984, 12944); 2 ♀♀, 5 ♂♂ [2 AL, 5 UN],
Yoko, 24 Sep. 2009, leg. unknown collector, 4 Sep. 2012, 7 Sep. 2012, leg. Prescott Musaba, 4 Sep. 2012,
5 Sep. 2012, 7 Sep. 2012, leg. Prescott Musaba, Tembele and Marc Boketshu (UNIKIS: GE283,
RMCA: a1.097-M-3164, UNIKIS: YK006, YK028CH, YK029CH, RMCA: a1.097-M-3165,
UNIKIS: YK032CH); 1 ♀ [UN], Zemio, unknown date, leg. unknown collector (RBINS: 184); 1 ad ♀,
1 ad ♂, 1 ad? ?, 1 ? [4 SS], Zobia, prior to 18 Apr. 1946, leg. unknown collector (RBINS: 6963 to 6966).
Micropteropus Matschie, 1899
DEMOCRATIC REPUBLIC OF THE CONGO: 10 ♀♀, 9 ♂♂ [1 SK, 18 UN], Kikwit, 27 Jun. 2015,
28 Jun. 2015, 29 Jun. 2015, leg. Prescott Musaba, Moïse Bipoo, Charle Andabhati, Benjamin Kirongozi
and Kambale Karupao (UNIKIS: KKT2848 to KKT2859, KKT2861 to KKT2866, KKT2877); 3 ♀♀,
1 ♂ [4 UN], Rethy, 22 Oct. 2015, 23 Oct. 2015, leg. Anne Laudisoit et al. (UNIKIS: ITU706, ITU710,
ITU712, ITU714).
Micropteropus intermedius Hayman, 1963
DEMOCRATIC REPUBLIC OF THE CONGO: 1 ♀ [SA], Banana or Netonna, unknown date, leg.
P. Hesse (SMF: 2509); 1 ♀, 2 ?? [2 SS, 1 SA], Luluabourg [= Kananga], 15 Mar. 1964, 31 Dec. 1964,
leg. Antoon Emeric Marcel De Roo (RMCA: 32380, 34406, 34409); 1 ♂ [SA], Thysville Cave b 13 a
[= Banjan Caves], 13 Jul. 1949, leg. Narcisse Leleup (RMCA: 22661).
Micropteropus pusillus (Peters, 1868)
BURUNDI: 3 sad ♂♂ [2 SA, 1 SK], Resha, 16 Mar. 1979, 18 Mar. 1979, 19 Mar. 1979, leg. unknown
collector (SMF: 57493 to 57495).
DEMOCRATIC REPUBLIC OF THE CONGO: 1 ad ♀, 1 ad ♂, 1 juv ♀, 3 juv ??, 1 ♀, 1 ♂, 9 ?? [1 SS,
2 SA, 8 AL, 4 SK, 2 UN], unknown locality, unknown date, leg. unknown collector, unknown date, leg.
Tshinyenye, unknown date, prior to 31 Dec. 1889, leg. P. Hesse, between 3 Feb. 1980 and 23 Mar. 1980,
leg. Betu Nsankulu, prior to 18 Aug. 1965, leg. Antoon Emeric Marcel De Roo, prior to 4 Jul. 1958, leg.
E. Jans (BMNH: 1990.223, MNHN: ZM-MO-1996-2470, ZM-MO-1996-2471, RMCA: 80.026-M0010, SMF: 11744, 11746, 11747, 2505 to 2508, RMCA: 80.020-M-0004, 33562, 33564, 33583, SMF:
11750, RMCA: 27375); 2 ♀♀, 1 ? [3 SS], Bakwakenge [= Bakwa-Kenge], 9 Sep. 1978, 12 Oct. 1979,
leg. Betu Nsankulu (RMCA: 80.013-M-0004, 80.013-M-0005, 79.022-M-0001); 1 ? [UN], Bas Congo
(former) Province [= Kongo Central Province], unknown date, leg. Alphonse Cabra (RMCA: 67);
1 ♂ [SS], Bengale, 28 Dec. 1976, leg. Tshinyenye (RMCA: 77.012-M-0002); 3 ♀♀, 3 ?? [1 SA, 4 AL,
1 UN], Beno, unknown date, leg. the Royal Museum for Central Africa (RMCA), prior to 3 Jan. 1950,
leg. P. Bayet (BMNH: 1954.784, 1954.785, RMCA: 19477a to 19477c, 19478); 3 ?? [3 AL], Benza
Masola, between 1 May 1913 and 31 May 1913, leg. Mayné (RMCA: 2099a to 2099c); 1 ♂ [SS],
Benzale, 28 Dec. 1976, leg. Tshinyenye (RMCA: 77.031-M-0008); 1 ♀, 3 ♂♂ [4 SS], Biakabomba,
24 Apr. 1979, leg. Betu Nsankulu (RMCA: 79.032-M-0002 to 79.032-M-0005); 7 ♀♀, 16 ♂♂, 6 ??
147
European Journal of Taxonomy 382: 1–327 (2017)
[29 AL], Boende, prior to 31 Jan. 1969, between 1 Jan. 1967 and 31 Dec. 1967, leg. P. Lootens (RMCA:
38657 to 38670, 38677 to 38685, 35629 to 35634); 1 ♀ [SS], Bokalakala, between 1 Oct. 1949 and
30 Nov. 1949, leg. Ngwe (RMCA: 19544); 1 ? [AL], Bokoro, prior to 26 Aug. 1952, leg. E. Jans (RMCA:
21297); 9 ♀♀, 1 ♂, 30 ?? [11 SS, 1 SA, 25 AL, 1 SO, 2 UN], Boma, unknown date, leg. unknown
collector, 20 Nov. 1920, 21 Nov. 1920, 24 Nov. 1920, 26 Nov. 1920, leg. Henri Schouteden, between
1 Aug. 1928 and 31 Aug. 1928, leg. Gerard, prior to 24 May 1912, leg. Barthélemy, prior to 26 May
1933, leg. Colsoulle, prior to 4 Aug. 1937, leg. Dartevelle (RMCA: 13851, 7292, 7293, 6542, 7295,
7297, 6543, 6545, 6544, 16181, 16216, 16219, 16220, 1051, 12125e to 12125h, 12126a to 12126j,
12127a to 12127i, 13947, 13951, 13954); 1 ♂ [SS], Byonga, 6 Oct. 1966, leg. Alexandre Prigogine
(RMCA: 34800); 1 ♀ [SK], Catchpole’s Farm, 24 Oct. 1924, leg. Edmund Heller (FMNH: 35684);
2 ♀♀, 1 ? [1 AL, 2 UN], Cel II, 16 Nov. 1951, leg. H. de Saeger et al., 16 Nov. 1951, leg. H. de Saeger
(RBINS: 13521.1, 13521.2, RMCA: 36555); 1 ♀, 1 ♂ [1 SS, 1 SO], Cinputu, 14 Sep. 1983, leg. Betu
Nsankulu (RMCA: 84.035-M-0009, 84.035-M-0012); 2 ?? [2 AL], Congo da Lemba, between
13 Jan. 1900 and 22 Jan. 1900, leg. Paul Van Bellighen, prior to 13 Mar. 1913, leg. Mayné (RMCA:
408b, 1545); 1 ♀ [SS], Dedegwa River, between 1 Jan. 1950 and 17 May 1952, leg. H. de Saeger
(RMCA: 36554); 1 ? [AL], Faradje, 12 Sep. 2010, leg. Guy-Crispin Gembu Tungaluna
(RMCA: a1.097-M-0553); 1 ? [SS], Fwa, Lake, 25 May 1984, leg. Betu Nsankulu (RMCA: 84.035-M0008); 3 ?? [3 AL], Ganda-Sundi, 20 Apr. 1931, leg. James Paul Chapin, 31 Mar. 1964, leg. Alexandre
Fain (AMNH: 86860, 86861, RMCA: 33262); 4 ♀♀ [4 SO], Gangala [= Gangala-na-Bodio],
24 Mar. 1957, 25 Mar. 1957, 26 Mar. 1957, leg. Heinz Stephan (RMCA: 27426 to 27429); 1 ? [UN],
Gangala-na-Bodio, 25 May 1948, leg. unknown collector (RBINS: 11660); 1 ? [AL], Kabinda, between
1 Jan. 1945 and 31 Dec. 1946, leg. Hautmann (RMCA: 18148); 2 ?? [2 SS], Kaniki Kapangu,
12 Mar. 1966, 13 Mar. 1966, leg. Antoon Emeric Marcel De Roo (RMCA: 35278, 35280); 3 ♀♀, 2 ??
[3 SS, 2 SK], Kasai, 24 Dec. 1981, 17 Jan. 1982, 24 Feb. 1982, prior to 12 Aug. 1983, leg. Betu Nsankulu
(RMCA: 83.025-M-0009, 83.025-M-0010, 83.025-M-0008, 83.025-M-0020, 83.025-M-0022); 4 ♀♀,
2 ♂♂ [5 SS, 1 SO], Kaswabilenga, 6 Oct. 1947, 7 Oct. 1947, leg. unknown collector, 6 Oct. 1947,
7 Oct. 1947, leg. Gaston-François de Witte (RBINS: 10679 to 10684); 2 ?? [2 AL], Kavuaya, prior to
31 Dec. 2007, prior to 2 Feb. 2007, leg. A. Kimbembi-ma-Ibaka (RMCA: a7.006-M-0012,
a7.006-M-0013); 6 ♀♀, 1 ♂ [7 SA], Kikole-Pecheur, 17 May 1980, leg. M. Mutashia (SMF: 58058 to
58064); 7 ad ♀♀, 4 ad ♂♂ [5 SA, 6 UN], Kikwit ITPK [= Kikwit], 1 Aug. 1995, leg. Herwig Leirs
et al. (MSB: 274975 to 274979, 274985, RMCA: 97.021-M-0805 to 97.021-M-0809); 1 ad ♀ [UN],
Kimbinga Market, 20 Jul. 1995, leg. Herwig Leirs et al. (MSB: 274577); 1 ad ♀, 3 ad ♂♂, 2 juv ??,
7 ♀♀, 15 ♂♂ [20 SS, 8 SN], Kimpoko [= Kimpoko-Nsele], unknown date, leg. Tsh Mutashiya,
22 Aug. 1976, 27 Aug. 1976, 28 Aug. 1976, 30 Aug. 1976, 14 Oct. 1976, 15 Oct. 1976, 31 Oct. 1976,
1 Nov. 1976, 24 Dec. 1976, 25 Aug. 1979, 27 Apr. 1980, leg. Tshinyenye (MNHN: ZM-MO-1986-1059
to ZM-MO-1986-1061, ZM-MO-1986-299, RMCA: 77.012-M-0003, 77.031-M-0005, 77.031-M-0006,
77.031-M-0002, 77.031-M-0003, 76.046-M-0001, 77.031-M-0007, 80.026-M-0002 to 80.026-M-0009,
80.038-M-0003 to 80.038-M-0006, 76.046-M-0002 to 76.046-M-0004, 77.021-M-0011, 77.031-M0004); 1 ? [SS], Kingabwa, prior to 3 Mar. 1976, leg. Tshinyenye (RMCA: 76.010-M-0002); 3 ??
[3 UN], Kinkole, 15 Aug. 1980, leg. V. Wallach (ZMA: MAM.21146 to MAM.21148); 7 ♂♂, 31 ??
[7 SO, 31 UN], Kinshasa, between 15 Jan. 1975 and 22 Jan. 1975, leg. Michael D. Gallagher, between
5 Feb. 1993 and 31 Dec. 1993, between 3 Feb. 1993 and 31 Dec. 1993, between 24 Jan. 1993 and
31 Dec. 1993, between 22 Feb. 1993 and 31 Dec. 1993, between 2 Jan. 1993 and 31 Dec. 1993, between
19 Feb. 1993 and 31 Dec. 1993, leg. Tshinyenye (HZM: 10.7795 to 37.7822, 7.7792 to 9.7794, RMCA:
93.097-M-0020, 93.097-M-0021, 93.097-M-0019, 93.097-M-0017, 93.097-M-0018, 93.097-M-0016,
93.097-M-0015); 45 ?? [45 UN], Kinshasa City Province, 12 Feb. 2001, 28 Feb. 2001, between
1 Apr. 1999 and 30 Apr. 1999, leg. unknown collector (SMF: 92974 to 92978, 92972 to 92973, 93243 to
93280); 56 ad ♀♀, 3 ad ♂♂, 1 ad? ♀, 3 ♀♀ [31 SA, 1 AL, 1 SK, 30 UN], Kinzambi Mission, 25 Jul. 1995,
26 Jul. 1995, 27 Jul. 1995, 30 Jul. 1995, 31 Jul. 1995, leg. Herwig Leirs et al. (MSB: 274943 to 274945,
RMCA: 97.021-M-0773 to 97.021-M-0775, MSB: 274946 to 274949, RMCA: 97.021-M-0776 to
148
VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
97.021-M-0779, MSB: 274951 to 274962, 274964 to 274971, RMCA: 97.021-M-0780 to 97.021-M0801, 97.021-M-0849, MSB: 274972, RMCA: 97.021-M-0802, MSB: 274973, 274974, RMCA: 97.021M-0803, 97.021-M-0804); 1 ♀, 2 ♂♂ [3 SS], Kitimi, between 1 Dec. 1979 and 24 Dec. 1979, leg.
Tshinyenye (RMCA: 80.006-M-0001 to 80.006-M-0003); 3 ♀♀ [3 AL], Kitombongo [= Kintobongo],
25 Sep. 2009, 26 Sep. 2009, leg. Alexandre Hassanin, Nicolas Nesi, Célestin Pongombo, Peter Vallo and
Didier Tshikung (MNHN: ZM-2011-807 to ZM-2011-809); 1 ♀ [SS], Kitutu, 15 Feb. 1964, leg.
Alexandre Prigogine (RMCA: 32406); 2 ♂♂ [2 SS], Kuili, 4 Jan. 1977, leg. Tshinyenye (RMCA:
77.031-M-0010, 77.031-M-0011); 3 ♂♂ [3 SS], Kuili Ngongo [= Kuili], 5 Jan. 1977, 6 Jan. 1977, leg.
Tshinyenye (RMCA: 77.012-M-0005, 77.012-M-0006, 77.012-M-0001); 1 ♀ [SS], Kwamouth, 29 May
1921, leg. Henri Schouteden (RMCA: 6546); 3 ♂♂ [3 SS], Kwango, 29 Dec. 1976, leg. Tshinyenye
(RMCA: 77.012-M-0004, 77.021-M-0015, 77.031-M-0009); 1 ♀, 1 ♂ [1 SS, 1 SO], Léopoldville
[= Kinshasa], 5 Apr. 1912, leg. Cuthbert Christy, 8 Jun. 1943, leg. R. Fiasse (RMCA: 1431, 17656);
13 ?? [12 SS, 1 UN], Lovanium University [= Kinshasa], 1 Nov. 1962, 24 Nov. 1962, 26 Nov. 1962,
24 Dec. 1962, 28 Dec. 1962, 5 Jan. 1963, 17 Jan. 1963, 2 Apr. 1963, 19 Apr. 1963, 23 Nov. 1963, between
1 Apr. 1963 and 1 Apr. 1964, leg. De Bont and Jo Van Orshoven, between 6 Apr. 1964 and 7 Apr. 1964,
leg. Antoon Emeric Marcel De Roo (RMCA: 32618, 32617, 32622, 32619, 32613, 32623, 32614, 32615,
32620, 32621, 32612, 32616, 32783); 1 ? [AL], Lubondai-Tshimbulu [= Lubondaie-Tshimbulu],
12 Dec. 1955, leg. S. Marks (RMCA: 26274); 1 ♀, 1 ? [2 AL], Lukolela, 3 Nov. 1930, 28 Feb. 1931,
leg. James Paul Chapin (AMNH: 86862, 86905); 1 juv ? [SS], Lukonga, 6 Aug. 1979, leg. Betu Nsankulu
(RMCA: 80.013-M-0006); 2 ad ♀♀, 3 ad ♂♂, 2 juv ♂♂, 4 juv ??, 1 ad+foetus ♀, 9 ♀♀, 5 ♂♂, 155 ??
[111 SS, 58 AL, 9 SO, 3 UN], Luluabourg [= Kananga], unknown date, leg. unknown collector,
unknown date, 4 Nov. 1963, 5 Nov. 1963, 22 Nov. 1963, 25 Nov. 1963, 26 Nov. 1963, 29 Nov. 1963,
19 Dec. 1963, 11 Jan. 1964, 11 Feb. 1964, 8 Mar. 1964, 9 Mar. 1964, 10 Mar. 1964, 11 Mar. 1964,
12 Mar. 1964, 14 Mar. 1964, 16 Mar. 1964, 17 Mar. 1964, 19 Mar. 1964, 16 Apr. 1964, 17 Apr. 1964,
18 Apr. 1964, 19 Apr. 1964, 22 Apr. 1964, 23 Apr. 1964, 24 Apr. 1964, 25 Apr. 1964, 3 May 1964, 4 May
1964, 10 May 1964, 28 May 1964, 30 May 1964, 2 Jun. 1964, 3 Jun. 1964, 6 Jun. 1964, 10 Jun. 1964,
13 Jun. 1964, 21 Sep. 1964, 16 Oct. 1964, 19 Oct. 1964, 21 Oct. 1964, 22 Oct. 1964, 24 Oct. 1964,
31 Oct. 1964, 1 Nov. 1964, 4 Nov. 1964, 7 Nov. 1964, 11 Nov. 1964, 25 Nov. 1964, 1 Dec. 1964,
31 Dec. 1964, 2 Feb. 1965, 14 Feb. 1965, 15 Feb. 1965, 18 Feb. 1965, 20 Feb. 1965, 22 Feb. 1965,
27 Feb. 1965, 28 Feb. 1965, 3 Mar. 1965, 4 Mar. 1965, 6 Mar. 1965, 8 Mar. 1965, 12 Mar. 1965,
19 Mar. 1965, 20 Mar. 1965, 22 Mar. 1965, 25 Mar. 1965, 1 Apr. 1965, 12 Apr. 1965, 28 Apr. 1965,
2 May 1965, 8 May 1965, 16 Jun. 1965, 26 Jun. 1965, 3 Jul. 1965, 16 Nov. 1965, 3 Mar. 1966, 27 May
1966, 14 Aug. 1968, 20 Aug. 1968, prior to 18 Aug. 1965, between 1 Jan. 1965 and 31 Dec. 1965,
between 1 Jan. 1964 and 31 Dec. 1964, leg. Antoon Emeric Marcel De Roo (BMNH: 1926.7.6.60,
1926.7.6.61, RMCA: 32695, 33329 to 33331, 33431 to 33433, 33671, 33555, 33677, 33678, 34408,
34414, 32375, 32376, 32642 to 32649, 32768, 33579, 33580, AMNH: 206839 to 206842, KU: 125197,
RMCA: 32377, 33645, 32379, 33426, 33680, 33581, 33416, 32692, 32693, 35610 to 35612, 33323,
32381 to 32383, 32673, 32674, 33681, 33682, 33653 to 33655, 34411, 34415, 32384, 32385, 32675 to
32680, 33417, 32681, 32386, 33427, 33428, 32682, 32683, 33656, 33657, AMNH: 206836 to 206838,
RMCA: 33322, 33434 to 33438, 33575, 33321, 33327, 33328, 35613, 32689, 33658 to 33663, 33418,
33429, 32640, 32641, 33664, 33665, 32366, 32684 to 32687, 33666 to 33668, 33430, 32688, 32367 to
32369, 33675, 33676, 33683, 32370, 32371, 33419, 35281, 33420, 33556, 33572, 32372, 32373, 33421
to 33423, 34412, 34413, 32694, 33574, 33576, 33557, 33573, 33669, 33670, 33415, 33679, 33324,
32696, 32697, 32363, 32364, 33672 to 33674, 32365, 33325, 33326, 33424, 33558, 33577, 33578,
33320, 32691, 33425, 33439 to 33448, 32374, 34407, 34410, 33559 to 33561, 33563, 33565 to 33567,
33582, 33584); 2 ♀♀, 1 ♂, 1 ? [1 SS, 3 SO], Make, 1 Jul. 1989, 12 Aug. 1992, 1 Nov. 1992, between
18 Apr. 1993 and 31 Dec. 1993, leg. Tshinyenye (RMCA: 89.057-M-0004, 92.148-M-0003, 92.119-M0004, 93.097-M-0014); 1 ? [AL], Makengo, 11 Dec. 1946, leg. Dartevelle (RMCA: 18160); 1 ♂ [SS],
Masaba, 27 Sep. 1976, leg. Tshinyenye (RMCA: 77.012-M-0007); 1 ? [AL], Mayumbe, prior to
31 Aug. 1899, leg. Alphonse Cabra (RMCA: 331); 1 ad ♀ [SA], Mbalaka (Plantation), 23 Jul. 1995,
149
European Journal of Taxonomy 382: 1–327 (2017)
leg. Herwig Leirs et al. (RMCA: 97.021-M-0772); 3 ♂♂ [3 SS], Mbawu, 27 May 1979, leg. Betu
Nsankulu (RMCA: 79.032-M-0010 to 79.032-M-0012); 1 ♀ [SS], Mbelekelo, 24 Feb. 1965, leg.
Alexandre Prigogine (RMCA: 33287); 1 ♀, 1 ♂, 7 ?? [3 AL, 6 UN], Mboga [= Boga], unknown date,
leg. unknown collector, unknown date, leg. Howard de Walden (BMNH: 1930.11.11.70 to 1930.11.11.78);
1 ♀ [SS], Moanda [= Muanda], 18 Aug. 1920, leg. Henri Schouteden (RMCA: 6547); 1 emb ♂, 27 ♀♀,
16 ♂♂ [44 AL], Mukulakulu, 22 Sep. 2009, leg. Alexandre Hassanin, Nicolas Nesi, Célestin Pongombo,
Peter Vallo and Didier Tshikung (MNHN: ZM-2011-810 to ZM-2011-853); 2 ♀♀, 2 ♂♂ [4 SS],
Mukwaya, 20 May 1979, 22 May 1979, leg. Betu Nsankulu (RMCA: 79.032-M-0006 to 79.032-M0009); 2 ♀♀ [2 AL], 4 km from Muvule Cave, 8 Oct. 2009, leg. Alexandre Hassanin, Nicolas Nesi,
Célestin Pongombo, Peter Vallo and Didier Tshikung (MNHN: ZM-2011-854, ZM-2011-855); 4 ♀♀,
6 ♂♂ [10 SO], Ndjili, between 4 Apr. 1993 and 31 Dec. 1993, between 22 Mar. 1993 and 31 Dec. 1993,
between 19 Mar. 1993 and 31 Dec. 1993, between 15 Apr. 1993 and 31 Dec. 1993, leg. Tshinyenye
(RMCA: 93.097-M-0013, 93.097-M-0011, 93.097-M-0012, 93.097-M-0004 to 93.097-M-0010); 1 ♂,
1 ? [2 SS], Nganza, 19 Sep. 1983, leg. Betu Nsankulu (RMCA: 84.035-M-0007, 84.035-M-0013); 1 ♂
[SS], Ngula, 6 Mar. 1977, leg. Tshinyenye (RMCA: 77.031-M-0012); 1 sad ♂ [SS], Niangara,
18 Nov. 1910, leg. Herbert Lang, James Paul Chapin and The American Museum Congo Expedition
(AMNH: 48756); 1 ♀ [SS], Nkiene, 30 Sep. 1980, leg. Tshinyenye (RMCA: 80.053-M-0001); 1 ? [UN],
N’soro, unknown date, leg. unknown collector (BMNH: 1907.7.8.29); 2 ♀♀, 2 ♂♂ [4 AL], Nyankunde,
16 Aug. 1976, 18 Aug. 1976, leg. Frieder Meier (MHNG: 1872.055 to 1872.058); 1 ♂ [UN], Parc
National de Garamba [= Garamba National Park], 22 May 1952, leg. H. de Saeger et al. (RBINS:
13522); 1 ♂ [UN], Rethy, 23 Oct. 2015, leg. Anne Laudisoit et al. (UNIKIS: ITU708); 1 ♂ [AL],
Sacred Heart Mission, 2 Aug. 1995, leg. Herwig Leirs et al. (RMCA: 97.021-M-1174); 2 ♂♂, 1 ?
[1 SS, 1 SO, 1 UN], St. Jos. (Misson) [= Mikalaya], 23 Nov. 1922, 19 Oct. 1924, 29 Oct. 1924, leg.
Richard Callewaert (AMNH: 86147, 86257, 86151); 2 ♀♀, 2 ?? [1 SS, 2 SK, 1 SO], St. Joseph de
Luluabourg Mission [= Mikalaya], 5 May 1922, 23 Sep. 1922, 23 Feb. 1923, between 19 Feb. 1923
and 1 Jun. 1923, leg. Richard Callewaert (RMCA: 7115, 7547, 7548, 7109); 1 ? [SS], Sud-Kivu
Province, 3 Nov. 1966, leg. A. Elbl (USNM: 535399); 1 ♀, 2 ?? [1 SA, 2 AL], Thysville [= MbanzaNgungu], 13 Jul. 1949, leg. Narcisse Leleup (RMCA: 22658 to 22660); 1 ♀, 2 ♂♂ [3 SS], Topetope,
18 Jun. 1966, 4 Jul. 1966, 8 Jul. 1966, leg. Alexandre Prigogine (RMCA: 34386, 34384, 34385); 2 ♂♂
[2 SS], Tschinputu [= Cinputu], 19 Sep. 1983, 20 Sep. 1983, leg. Betu Nsankulu (RMCA: 84.035-M0011, 84.035-M-0010); 3 ♀♀, 1 ♂ [4 SS], Tshibasha, 18 Nov. 1970, leg. Betu Nsankulu (RMCA:
36251 to 36254); 3 ?? [3 SO], Tshimputu, 23 May 1975, leg. Betu Nsankulu (RMCA: 75.061-M-0008
to 75.061-M-0010); 1 ♀, 7 ?? [8 AL], Tshuapa District, between 1 Jan. 1960 and 31 Dec. 1964, leg.
P. Lootens (RMCA: 34166, 34167, 34169, 34174, 34175, 34179, 34182, 34183); 1 ? [AL], Vista
[= Nsiamfumu], between 1 Jan. 1933 and 31 Dec. 1933, leg. Dartevelle (RMCA: 12167); 1 ? [UN],
Zambi, prior to 31 Jan. 1899, leg. J.M. Derscheid (RMCA: 149); 1 ♀ [SK], Zongo, 2 Jul. 1948, leg.
unknown collector (RBINS: 7715).
Nanonycteris Matschie, 1899
Nanonycteris veldkampii (Jentink, 1888)
DEMOCRATIC REPUBLIC OF THE CONGO: 3 ?? [3 SK], unknown locality, between 1 May 2010
and 31 May 2010, leg. Nicolas Nesi, Blaise Kadjo and Christiane Denys (MNHN: ZM-2014-1169, ZM2014-1171, ZM-2014-1173).
150
VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
Myonycterini Lawrence & Novick, 1963
Megaloglossus Pagenstecher, 1885
Megaloglossus woermanni Pagenstecher, 1885
DEMOCRATIC REPUBLIC OF THE CONGO: 1 ♂, 1 ? [2 UN], unknown locality, unknown date,
12 Feb. 2001, leg. unknown collector (BMNH: 1990.224, SMF: 92984); 7 ♂♂ [7 AL], Aketi,
22 Jun. 2015, 23 Jun. 2015, leg. Vlir/Aketi (UNIKIS: AKETI608 to AKETI611, AKETI612 to
AKETI614); 1 ? [SS], Ba Balaye, 24 Apr. 1978, leg. Betu Nsankulu (RMCA: 78.020-M-0001); 4 ♀♀,
1 ♂ [2 AL, 3 SK], Baliko, 10 Jun. 2012, 11 Jun. 2012, 15 Jun. 2012, leg. William (“Bill”) T. Stanley,
prior to 31 Jan. 2007, leg. Guy-Crispin Gembu Tungaluna (FMNH: 219589 to 219591,
RMCA: a1.097-M-0410, a1.097-M-0411); 1 ♂ [SK], Banana, unknown date, leg. P. Hesse (SMF:
2513); 1 ♀ [SS], Bene Longo, 23 Aug. 1978, leg. Betu Nsankulu (RMCA: 79.022-M-0007); 1 ♀ [AL],
Beno, unknown date, leg. the Royal Museum for Central Africa (RMCA) (BMNH: 1954.787); 1 ♀ [SS],
Bikoro, 2 Mar. 1921, leg. Henri Schouteden (RMCA: 6548); 6 ♀♀, 12 ♂♂, 25 ?? [43 AL], Boende,
prior to 31 Jan. 1969, between 1 Dec. 1969 and 31 Dec. 1969, between 1 Nov. 1969 and 30 Nov. 1969,
between 1 Feb. 1970 and 28 Feb. 1970, between 1 Jan. 1974 and 31 Dec. 1974, between 1 Jan. 1973 and
31 Dec. 1973, between 1 Jan. 1970 and 31 Jan. 1970, between 1 Jan. 1967 and 31 Dec. 1967, leg.
P. Lootens (RMCA: 38686 to 38705, 38817, 38818, 75.035-M-0005 to 75.035-M-0019, 75.035-M0001, 75.035-M-0004, 35635 to 35638); 1 ? [AL], Bokuma, 14 Dec. 1952, leg. P. Lootens (RMCA:
22031); 5 ♀♀, 5 ♂♂ [4 SK, 6 UN], Bomane [= Bomaneh], 21 May 2010, 30 May 2010, leg. GuyCrispin Gembu Tungaluna, Prescott Musaba and José Akaibe (UNIKIS: CRT2159, CRT2171, CRT2174,
CRT2190, CRT2192, CRT2194, CRT2054 to CRT2056, CRT2076); 1 ♂ [SS], Byonga, 24 Jan. 1965,
leg. Alexandre Prigogine (RMCA: 33288); 7 ♀♀, 16 ♂♂, 1 ? [24 AL], Epulu, unknown date, leg. Luc
Bijnens, 25 Aug. 2015, 27 Aug. 2015, 29 Aug. 2015, 30 Aug. 2015, 31 Aug. 2015, leg. André Malekani
(RMCA: 90.031-M-0001, UNIKIS: EPLU630, EPLU632, EPLU654, EPLU658, EPLU659, EPLU662,
EPLU665 to EPLU668, EPLU671 to EPLU675, EPLU681, EPLU685, EPLU694, EPLU699, EPLU700,
EPLU703 to EPLU705); 10 ♂♂ [3 AL, 2 SK, 3 SB, 2 SN], Epulu, 2 km W, 2 Jul. 1991, 3 Jul. 1991,
4 Jul. 1991, leg. Julian C. Kerbis Peterhans (FMNH: 149392 to 149401); 3 ♂♂ [3 AL], Equateur
(former) Province, prior to 7 Oct. 1996, leg. unknown collector (SMF: 84407 to 84409); 1 ? [AL],
Ibembo, between 1 Nov. 1949 and 30 Nov. 1949, leg. Jozef Hutsebaut (RMCA: 19857); 1 ? [AL], Ikela,
prior to 4 Jan. 1958, leg. P. Lootens (RMCA: 27017); 1 ♀ [UN], Île Mai [= Mai Island], 15 Apr. 2012,
leg. Prescott Musaba, Tembele and Marc Boketshu (UNIKIS: CHIMA54); 9 ♀♀, 22 ♂♂, 7 ?? [26 AL,
12 UN], Île Mbiye [= Mbiye Island], 15 Jan. 2012, 24 Feb. 2012, 25 Feb. 2012, 26 Jul. 2012, leg.
Prescott Musaba, 15 Jan. 2012, 30 Mar. 2012, leg. Prescott Musaba, Tembele and Marc Boketshu,
19 Feb. 2012, 25 Feb. 2012, leg. Prescott Musaba et al., 19 Feb. 2012, 24 Feb. 2012, 25 Feb. 2012,
19 Apr. 2014, 22 Jun. 2014, 23 Jun. 2014, leg. Prescott Musaba, Moïse Bipoo, Richard Tamaru, Pauline
Isude and Joseph Kussa, 8 Aug. 2015, 9 Aug. 2015, 25 Dec. 2015, 26 Dec. 2015, leg. Prescott Musaba,
Moïse Bipoo, Charle Andabhati, Benjamin Kirongozi and Kambale Karupao (RMCA: a1.097-M-3206
to a1.097-M-3209, UNIKIS: CHIM01, RMCA: a1.097-M-3210, UNIKIS: CHIM09, CHIM04/14,
CHIM018/14, CHIM021/14, CHIM022/14, CHIM023/14, RMCA: a1.097-M-3199 to a1.097-M-3201,
UNIKIS: CHIM17, RMCA: a1.097-M-3202 to a1.097-M-3205, UNIKIS: CHIM24, CHIM25,
ABCHIM03 to ABCHIM05, RMCA: a1.097-M-3211, UNIKIS: ABCHIM06, CHIM36, CHIM37,
CHIMTB02 to CHIMTB10); 3 ♀♀, 3 ♂♂ [4 AL, 2 UN], Irangi, 26 Jan. 1984, 6 Feb. 1984, 13 Feb. 1984,
leg. Heinz Stephan, 17 Oct. 1990, leg. Wim Bergmans (SMF: 65004, ZMA: MAM.24195, MAM.24196,
SMF: 65001 to 65003); 1 ♀ [SS], Kakanda Cave, 29 Aug. 1964, leg. Alexandre Prigogine (RMCA:
33065); 1 juv ♀, 2 ♀♀, 2 ♂♂ [5 SS], Kamituga, 21 Dec. 1950, 1 Feb. 1964, 31 Dec. 1965, 30 Mar. 1967,
31 Mar. 1967, leg. Alexandre Prigogine (RMCA: 32394 [paratype Megaloglossus woermanni
prigoginei Hayman, 1966], 20429 [paratype Megaloglossus woermanni prigoginei Hayman, 1966],
35073, 35074, 33291); 1 ♂ [SS], Kamueshi [= Kamuesha], between 2 Apr. 1979 and 3 Apr. 1979, leg.
151
European Journal of Taxonomy 382: 1–327 (2017)
Betu Nsankulu (RMCA: 79.032-M-0014); 1 ♀ [AL], Kananga, 17 Feb. 1964, leg. Antoon Emeric
Marcel De Roo (AMNH: 212578); 1 ♀, 1 ♂ [2 UN], Karambi, 6 Apr. 1992, leg. Ngenge Masumbuko
Kamitongo (ZMA: MAM.24564, MAM.24565); 2 ♀♀ [2 SA], Kikole-Pecheur, 17 May 1980, leg.
M. Mutashia (SMF: 58067, 58068); 4 ♀♀, 1 ♂ [5 SS], Kiliza [River], 24 May 1964, 25 May 1964,
26 May 1964, leg. Alexandre Prigogine (RMCA: 32582 [paratype Megaloglossus woermanni
prigoginei Hayman, 1966], 32577 [holotype Megaloglossus woermanni prigoginei Hayman, 1966],
32579 [paratype Megaloglossus woermanni prigoginei Hayman, 1966], 32578, 32583 [paratype
Megaloglossus woermanni prigoginei Hayman, 1966]); 21 ?? [21 UN], Kinshasa City Province,
between 1 Apr. 1999 and 30 Apr. 1999, leg. unknown collector (SMF: 93297 to 93317); 4 ad ♀♀, 2 ad
♂♂, 2 sad ♀♀, 7 ♀♀, 1 ♂ [10 SA, 6 UN], Kinzambi Mission, 26 Jul. 1995, 27 Jul. 1995, 28 Jul. 1995,
31 Jul. 1995, leg. Herwig Leirs et al. (MSB: 274831, 274832, 274834, 274835, RMCA: 97.021-M-0762
to 97.021-M-0765, MSB: 274929, RMCA: 97.021-M-0766 to 97.021-M-0769, MSB: 274930, RMCA:
97.021-M-0770, 97.021-M-0771); 1 ? [AL], Kisangani, between 1 Jun. 1982 and 30 Jun. 1982, leg.
Marc Colyn (RMCA: 82.031-M-0011); 2 ♀♀ [2 SS], Kitongo, 3 Jun. 1964, 4 Jun. 1964, leg. Alexandre
Prigogine (RMCA: 32581 [paratype Megaloglossus woermanni prigoginei Hayman, 1966], 32580);
41 ♀♀, 8 ♂♂ [29 SA, 3 AL, 12 SK, 5 UN], Kona, 12 May 2010, 13 May 2010, 14 May 2010, 15 May
2010, 16 May 2010, 17 May 2010, 21 May 2010, between 13 May 2010 and 14 May 2010, leg. GuyCrispin Gembu Tungaluna, Prescott Musaba and José Akaibe (UNIKIS: CRT1908, CRT1912 to
CRT1915, CRT1919, CRT1922 to CRT1924, CRT1927, CRT1928, CRT1930 to CRT1936, CRT1938,
CRT1941, CRT1946, CRT1952, CRT1954 to CRT1957, CRT1961, CRT1962, CRT1964, CRT1965,
CRT1985 to CRT1990, CRT1998, CRT2002, CRT2004 to CRT2010, CRT2013, CRT2191, CRT2193,
CRT1951); 1 ? [AL], Konyola, prior to 5 Aug. 1988, leg. Luc De Vos, Kimbembi and Alain Vandelanoote
(RMCA: 88.025-M-0004); 4 ♀♀, 3 ♂♂ [2 SK, 5 UN], Lieki, 22 May 2010, 30 May 2010, 1 Jun. 2010,
3 Jun. 2010, leg. Guy-Crispin Gembu Tungaluna, Prescott Musaba and José Akaibe (UNIKIS: CRT2031,
CRT2026, CRT2024, CRT2025, CRT2027, CRT2049, CRT2053); 2 ♀♀ [2 SK], Luki Biosphere
Reserve, 13 Jul. 2013, leg. William (“Bill”) T. Stanley (FMNH: 222419, 222420); 1 ♀ [AL], Luki
Biosphere Reserve, Forest NE of, 17 Sep. 2005, leg. P. Lukamba (ZMA: MAM.26275); 2 ♂♂ [1 SS,
1 AL], Lukolela, 12 Aug. 1930, leg. Franklin Edson, 12 Aug. 1930, leg. James Paul Chapin (AMNH:
86771, 86908); 1 ♂, 11 ?? [2 SS, 10 AL], Luluabourg [= Kananga], 26 Apr. 1964, 15 Feb. 1965,
16 Feb. 1965, 20 Feb. 1965, 25 Feb. 1965, 25 Mar. 1965, 2 May 1965, 30 Jun. 1965, prior to 18 Aug. 1965,
leg. Antoon Emeric Marcel De Roo (RMCA: 33341 to 33343, 33371, 33344 to 33346, 33370, 33604,
33553, 33554, 33603); 1 ? [SS], Lundjulu, 14 Oct. 1952, leg. Schepens (RMCA: 21586 [paratype
Megaloglossus woermanni prigoginei Hayman, 1966]); 1 ♂ [UN], Malembe, 20 Apr. 1992, leg.
Ngenge Masumbuko Kamitongo (ZMA: MAM.24566); 1 ♀ [AL], Mandima, 7 Jul. 1976, leg. Frieder
Meier (MHNG: 1872.090); 1 ad ♂, 1 juv ♂, 20 ♀♀, 46 ♂♂ [9 AL, 59 SK], Masako, 12 Jun. 2015,
13 Jun. 2015, leg. Prescott Musaba, Moïse Bipoo, Charle Andabhati, Benjamin Kirongozi and Kambale
Karupao, 17 Dec. 1987, 18 Dec. 1987, 19 Dec. 1987, 20 Dec. 1987, 21 Dec. 1987, 22 Dec. 1987,
13 Jan. 1988, 15 Jan. 1988, 20 Jan. 1988, 23 May 1989, 18 Sep. 1989, 28 Nov. 1989, 22 Mar. 1990,
29 Apr. 1990, leg. Ndey B. Ifuta (UNIKIS: CHMS067, RMCA: 93.079-M-0080, 93.079-M-0081,
UNIKIS: CHMS069, CHMS073, CHMS074, RMCA: 93.079-M-0082, 93.079-M-0083, 93.079-M0269, 93.079-M-0229, 93.079-M-0257, 93.079-M-0060 to 93.079-M-0062, 93.079-M-0230 to 93.079M-0233, 93.079-M-0293 to 93.079-M-0297, 93.079-M-0063, 93.079-M-0064, 93.079-M-0298, 93.079M-0084 to 93.079-M-0091, 93.079-M-0065 to 93.079-M-0068, 93.079-M-0234 to 93.079-M-0239,
93.079-M-0069 to 93.079-M-0071, 93.079-M-0240, 93.079-M-0241, 93.079-M-0268, 93.079-M-0280,
93.079-M-0299, 93.079-M-0300, 93.079-M-0259, 93.079-M-0260, 93.079-M-0072 to 93.079-M-0079,
93.079-M-0242, 93.079-M-0301, 93.079-M-0369, 93.079-M-0258, 93.079-M-0261); 1 ad ♂, 1 sad ♀
[2 SA], Mbalaka (Plantation), 19 Jul. 1995, 23 Jul. 1995, leg. Herwig Leirs et al. (RMCA: 97.021-M0760, 97.021-M-0761); 13 ad ♀♀, 2 ad ♂♂, 1 juv ♀, 3 ♀♀ [10 SA, 9 UN], Mbwambala, 20 Jun. 1995,
21 Jun. 1995, 23 Jun. 1995, 1 Jul. 1995, 2 Jul. 1995, 3 Jul. 1995, 4 Jul. 1995, 6 Jul. 1995, 7 Jul. 1995,
11 Jul. 1995, leg. Herwig Leirs et al. (RMCA: 97.021-M-0756, MSB: 274637, 274648, RMCA: 97.021152
VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
M-0750, MSB: 274638, RMCA: 97.021-M-0751, MSB: 274643, 274644, 274646, 274647, RMCA:
97.021-M-0753 to 97.021-M-0755, 97.021-M-0752, 97.021-M-0757, MSB: 274635, RMCA: 97.021M-0758, MSB: 274636, RMCA: 97.021-M-0759); 1 ♂ [SS], Mukwaya, between 18 Apr. 1979 and
22 May 1979, leg. Betu Nsankulu (RMCA: 79.032-M-0015); 1 ♂ [SS], Munga, 22 Apr. 1978, leg.
Alexandre Prigogine (RMCA: 79.033-M-0001); 1 ♀ [SS], Mwenge, 4 Feb. 1967, leg. Alexandre
Prigogine (RMCA: 35075); 7 ♂♂ [6 SS, 1 SO], Ngula, 6 Mar. 1977, 8 Mar. 1977, 11 Mar. 1977, leg.
Tshinyenye (RMCA: 77.021-M-0009, 77.021-M-0012, 77.021-M-0013, 77.021-M-0010, 77.021-M0014, 77.021-M-0016, 77.031-M-0001); 4 ♀♀ [4 UN], Obenge, 7 Feb. 2013, leg. Prescott Musaba and
André Malekani (UNIKIS: TLL381 to TLL384); 4 ♀♀, 1 ♂ [2 SS, 2 AL, 1 SK], Quatorze, 25 Jun. 2013,
27 Jun. 2013, 1 Jul. 2013, 2 Jul. 2013, leg. William (“Bill”) T. Stanley (FMNH: 222417, 222418, 222620,
222416, 222621); 14 ♀♀, 6 ♂♂ [19 AL, 1 UN], Rubi-Tele, 17 Aug. 2013, 20 Aug. 2013, 23 Aug. 2013,
24 Aug. 2013, 25 Aug. 2013, 27 Aug. 2013, leg. Guy-Crispin Gembu Tungaluna, Prescott Musaba and
André Malekani (UNIKIS: RBTL486 to RBTL491, RBTL495, RBTL521, RBTL522, RBTL548 to
RBTL551, RBTL555, RBTL556, RBTL562 to RBTL564, RBTL576, RBTL577); 1 ♀ [SS], Tandala,
6 Jul. 1979, leg. C. Brian Robbins (USNM: 537700); 7 juv ??, 1 ♀, 1 ♂ [9 AL], Tshakala, between
1 Jan. 1982 and 31 Jan. 1982, leg. Marc Colyn (RMCA: 82.019-M-0013 to 82.019-M-0021); 1 ? [SO],
Tshimputu, 23 Mar. 1975, leg. Betu Nsankulu (RMCA: 75.061-M-0007); 1 ♀, 7 ?? [8 AL], Tshuapa
District, between 1 Jan. 1960 and 31 Dec. 1964, leg. P. Lootens (RMCA: 34168, 34171 to 34173, 34176
to 34178, 34180); 1 ♂ [UN], Uma, 10 Apr. 2014, leg. Guy-Crispin Gembu Tungaluna, Prescott Musaba
and André Malekani (UNIKIS: UMA544); 19 ?? [19 AL], Wafanya [= Wafania], unknown date,
17 Aug. 1991, 30 Aug. 1991, 7 Sep. 1991, 12 Sep. 1991, 20 Mar. 1992, 26 Apr. 1992, 27 Apr. 1992,
4 May 1992, 10 May 1992, leg. P. Lootens (RMCA: 92.079-M-0125, 92.079-M-0126, 92.079-M-0018,
92.079-M-0024, 91.076-M-0078, 91.076-M-0056, 92.079-M-0020, 92.079-M-0021, 92.079-M-0023,
92.079-M-0017, 92.079-M-0022, 92.079-M-0025, 92.079-M-0014, 92.079-M-0015, 92.079-M-0019,
91.076-M-0063, 92.079-M-0016, 91.076-M-0072, 91.076-M-0073); 5 ♀♀ [2 SK, 3 UN], Yaekela,
3 May 2010, 4 May 2010, 5 May 2010, leg. Guy-Crispin Gembu Tungaluna, Prescott Musaba and José
Akaibe (UNIKIS: CRT2361, CRT1809, CRT1812, CRT1814, CRT1846); 2 ♀♀, 1 ♂ [2 AL, 1 UN],
Yoko, 4 Sep. 2012, 7 Sep. 2012, leg. Prescott Musaba, 4 Sep. 2012, leg. Prescott Musaba, Tembele and
Marc Boketshu (RMCA: a1.097-M-3197, UNIKIS: YK002CH, RMCA: a1.097-M-3198); 3 ♀♀, 9 ♂♂
[1 SK, 11 UN], Zookis [= Kisangani], 16 Feb. 2013, leg. Prescott Musaba and André Malekani,
24 Dec. 2015, leg. Prescott Musaba, Moïse Bipoo, Charle Andabhati, Benjamin Kirongozi and Kambale
Karupao (UNIKIS: CSB2, CSB5, CSB6, ZCH01 to ZCH04, ZCH07 to ZCH11).
Myonycteris Matschie, 1899
Myonycteris angolensis (Bocage, 1898)
BURUNDI: 4 ♀♀ [4 SB], Giserama, Colline, 30 Jul. 1991, 31 Jul. 1991, 10 Aug. 1991, leg. Julian
C. Kerbis Peterhans, 31 Mar. 1992, leg. L. Davenport (FMNH: 148975, 148974, 148350, 148973); 1
juv ♂, 3 ♀♀ [1 SA, 3 SK], Kayanza, 26 Mar. 1979, leg. Heinz Stephan (SMF: 57579 to 57582); 1 ♀,
1 ♂ [1 AL, 1 SO], Kigwena Hill, 12 Oct. 1990, leg. Matthew Criblez, 24 Apr. 1982, leg. Danny Meirte
(FMNH: 148349, RMCA: 82.016-M-0069); 2 ♀♀, 10 ♂♂, 1 ? [2 AL, 6 SB, 5 SN], Kwogofe, Colline,
1 Apr. 1992, 3 Apr. 1992, 4 Apr. 1992, 5 Apr. 1992, 8 Apr. 1992, 9 Apr. 1992, 10 Apr. 1992, 11 Apr. 1992,
12 Apr. 1992, leg. L. Davenport (FMNH: 148351, 148363, 148360, 148361, 148352, 148358, 148353 to
148357, 148359, 148362); 2 ♀♀, 2 ♂♂ [1 AL, 1 SK, 1 SB, 1 SN], Mumushwizi Valley, 18 Aug. 1993,
23 Aug. 1993, 24 Aug. 1993, leg. Jay L. Udelhoven (FMNH: 155943 to 155946); 7 ♂♂, 1 ? [2 AL, 3 SB,
3 SN], Nyagatarugwa River Valley, 9 Aug. 1992, 10 Aug. 1992, leg. L. Davenport (FMNH: 148343,
148341, 148342, 148344 to 148348); 1 ♂ [SN], Nyamugari Hill, 11 Sep. 1990, leg. L. Davenport
(FMNH: 148364); 1 ♀, 18 ♂♂, 3 ?? [6 AL, 5 SK, 7 SB, 4 SN], Ruhinga Hill, 9 Mar. 1993, 10 Mar. 1993,
11 Mar. 1993, 12 Mar. 1993, 13 Mar. 1993, leg. Jay L. Udelhoven (FMNH: 155951 to 155968, 155947
to 155950); 5 ♀♀, 2 ♂♂ [1 SS, 1 AL, 5 SB], Teza, 2.3 km N, 0.7 km W [= Kibira National Park],
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European Journal of Taxonomy 382: 1–327 (2017)
14 Mar. 1990, 16 Mar. 1990, 19 Mar. 1990, 26 Mar. 1990, 28 Mar. 1990, leg. Julian C. Kerbis Peterhans
(FMNH: 137620 to 137624, 137619, 137476).
DEMOCRATIC REPUBLIC OF THE CONGO: 6 ♀♀ [6 AL], Alambi, 5 Aug. 1976, 6 Aug. 1976,
7 Aug. 1976, leg. Frieder Meier (MHNG: 1872.072 to 1872.077); 1 ? [SS], Banjan Caves, prior to
3 Nov. 1936, leg. Schwetz (RMCA: 13689); 3 ♀♀, 4 ?? [7 AL], Barainda Cave B 13, 15 Jun. 1949,
leg. Narcisse Leleup (RMCA: 22504 to 22510); 4 ♀♀, 1 ♂ [5 AL], Bossainda, unknown date, leg.
the Royal Museum for Central Africa (RMCA) (BMNH: 1954.846 to 1954.850); 8 ♀♀, 5 ♂♂ [1 SS,
12 AL], Bushokwe Caves, 1 Aug. 1956, leg. Alvin Novick, 3 Jun. 1957, leg. Jean Dubois (MCZ:
49032, 48288 to 48299); 2 ♀♀, 1 ♂, 4 ?? [6 AL, 1 UN], Butembo, 1 Jan. 1970, leg. M.J. Celis and
M. Lejeune, 21 Feb. 1956, prior to 29 Jul. 1957, prior to 25 Jan. 1956, leg. P. Dyleff (RMCA: 36194,
36204, 36207, 36208, 23675, 23449, 26435); 1 ? [UN], Djelube River, lower, unknown date, leg.
unknown collector (RBINS: 13228); 1 ? [SS], Djugu Forest, between 1 Jan. 1952 and 31 Dec. 1952,
leg. Alexandre Fain (RMCA: 23151); 1 ♂ [SS], Drodro, 22 Sep. 1952, leg. Alexandre Fain (MHNG:
921.028); 1 ♀, 2 ♂♂ [3 AL], Eidolon Land, 4 Oct. 2009, leg. Alexandre Hassanin, Nicolas Nesi,
Célestin Pongombo, Peter Vallo and Didier Tshikung (MNHN: ZM-2011-804 to ZM-2011-806); 2 ♀♀,
4 ♂♂ [6 UN], Goroumbwa Mine, 21 Oct. 1999, 22 Oct. 1999, 24 Oct. 1999, 25 Oct. 1999, leg. Robert
(Bob) Swanepoel (TM: DBA069, DBA076, DBA126, DBA227, DBA230, DBA287); 1 ♂ [AL], Grand
Ebouli de Lemera [= Itombwe], 10 Sep. 1957, leg. Narcisse Leleup (RMCA: 26706); 1 ♀ [SK],
Irangi, 22 Jan. 1984, leg. Heinz Stephan (SMF: 65027); 1 imm ♀ [SS], Jaima, 26 Aug. 1906, leg.
The Alexander-Gosling Expedition (BMNH: 1907.7.8.24); 1 ♀, 1 ? [1 SS, 1 UN], Kabambi, unknown
date, leg. unknown collector, 17 Aug. 1947, leg. Jean Hiernaux (INPBC: W1461, RBINS: 7055); 1 sad
♀ [SB], Kabobo Forest, 16 Feb. 2007, leg. B.D. Marks (FMNH: 195081); 1 juv ♀, 2 ♀♀ [3 SA],
Kahuzi-Biega National Park, 16 Oct. 1965, 20 Apr. 1966, 21 Apr. 1966, leg. P. Kunkel (SMF: 31820,
31818, 31819); 2 ♀♀ [2 AL], Kalumbu Cave, 1 Dec. 1957, leg. Félix / Michel Anciaux de Faveaux
(RMCA: 27592, 27593); 1 ♂ [SO], Kasai, 22 Jan. 1982, leg. Betu Nsankulu (RMCA: 83.025-M-0023);
1 ? [UN], Katanda, unknown date, leg. unknown collector (INPBC: W975.2); 3 ?? [3 AL], Kibwe Lya
Mikako Cave, 28 Dec. 1966, leg. M.J. Celis and M. Lejeune (RMCA: 35168, 35169, 35204); 1 ♂ [SS],
Kiliza [River], 17 Jan. 1966, leg. Alexandre Prigogine (RMCA: 34210); 1 ♀, 4 ♂♂ [5 SS], Kilo mines
[= Kilomines], 20 Apr. 1949, leg. Jean Hiernaux (RMCA: 19090 to 19094); 1 ? [AL], Kimbemba
Cataracts, between 1 Sep. 1936 and 30 Sep. 1936, leg. Schwetz (RMCA: 13701); 1 ♀ [UN], Kinshasa,
unknown date, leg. Henri Schouteden (RMCA: 15760); 1 ? [AL], Kisangani, 20 Mar. 1988, leg.
Luc Bijnens (RMCA: 88.017-M-0001); 3 ♂♂ [3 AL], Komanda, 21 Aug. 1976, leg. Frieder Meier
(MHNG: 1872.078 to 1872.080); 1 ? [SS], Léopoldville [= Kinshasa], prior to 31 Dec. 1926, leg.
Henri Schouteden (RMCA: 15760a); 1 ♀ [SO], Lubango [= Lobango], 25 May 1950, leg. Alexandre
Prigogine (RMCA: 20691); 1 ♂ [SS], Lusilubi Valley, 12 Aug. 1946, leg. J. de Wilde (RBINS: 13328);
1 ♀, 4 ♂♂ [5 SS], Lutunguru, 30 Mar. 1949, 5 Apr. 1949, 6 Jul. 1949, 4 Aug. 1949, 16 Feb. 1952,
leg. Alexandre Prigogine (RMCA: 20820, 18863, 19053, 18862, 19052); 1 ♂ [AL], Lwana River,
19 Apr. 1992, leg. Ngenge Masumbuko Kamitongo (ZMA: MAM.24560); 2 ♀♀, 2 ♂♂ [2 SS, 2 SA],
Lwiro [= Luiro], 1 Apr. 1967, 4 Apr. 1967, leg. Alexandre Prigogine, 28 May 1964, between 1 Jul. 1965
and 31 Jul. 1965, leg. P. Kunkel (RMCA: 35064, SMF: 31816, RMCA: 35065, SMF: 31817); 1 ♂ [SS],
Make, 23 Jul. 1993, leg. Tshinyenye (RMCA: 93.097-M-0022); 2 foetus ??, 1 ♀ [2 AL, 1 SK], Masako,
16 Dec. 1987, leg. Ndey B. Ifuta, between 23 May 1988 and 28 May 1988, leg. Luc Bijnens (RMCA:
93.079-M-0227, 88.017-M-0002, 88.017-M-0003); 2 ♀♀, 2 ♂♂ [4 SS], Matadi, 15 Mar. 1987,
16 Mar. 1987, leg. Ntotoji (RMCA: 87.019-M-0001 to 87.019-M-0004); 2 ?? [1 AL, 1 UN], Matupi Cave,
unknown date, leg. unknown collector, 4 Aug. 1955, leg. Gaston-François de Witte (INPBC: W1459,
RMCA: 37118); 1 ♀ [SS], Miki, 20 Aug. 1954, leg. Alexandre Prigogine (RMCA: 23423); 1 ? [SS],
Mongbwalu, 14 Aug. 1939, leg. Janssens (RMCA: 15759); 3 ad ♀♀ [3 AL], Montembessa Cave,
between 1 Nov. 1969 and 30 Nov. 1969, between 1 Sep. 1969 and 30 Sep. 1969, leg. unknown collector
(MNHN: ZM-MO-1985-1165 to ZM-MO-1985-1167); 2 ♀♀, 6 ♂♂, 7 ?? [8 SS, 1 SA, 6 AL], Mount
154
VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
Hoyo Caves, 17 Aug. 1947, leg. Jean Hiernaux, 19 Sep. 1955, leg. James Paul Chapin, 4 May 1955,
leg. W. Smith and E. Baker, 4 May 1955, leg. Waldo LaSalle Schmitt and E. Baker (RBINS: 7065,
AMNH: 180902 to 180905, USNM: 301703 to 301712); 1 ♂, 3 ?? [4 SS], Mount Wago [= Wago,
Mount], unknown date, prior to 28 Mar. 1952, leg. Alexandre Fain, 1 Jan. 1952, leg. unknown collector
(BMNH: 1953.477, 1953.478, RBINS: 21665, RMCA: 20804); 2 ♀♀ [1 SS, 1 SO], Munoi [= Munoi
conluence], 14 Apr. 1948, leg. Gaston-François de Witte, 14 Jun. 1947, leg. unknown collector (RBINS:
10663, 10662); 1 ? [UN], Muvo, Mount, unknown date, leg. unknown collector (INPBC: V1235); 1 ♀
[SS], Mwenge, 8 Feb. 1967, leg. Alexandre Prigogine (RMCA: 35077); 1 ♀, 1 ? [1 SS, 1 SO], Pelenge,
Gorges, 4 Jun. 1947, leg. unknown collector, between 4 Apr. 1947 and 4 Jun. 1947, leg. Gaston-François
de Witte (RBINS: 10660, 10661); 1 ♀ [SS], Rutshuru, 26 Feb. 1938, leg. J. Ghesquière (RMCA:
14597); 1 ? [SO], Ruwenzori, Foothills, between 1 Jan. 1925 and 31 Dec. 1925, leg. Henri Martin
Hackars (RMCA: 9904a); 1 ♂, 3 ?? [4 SS], Saliboko, 22 Sep. 1952, leg. unknown collector, between
22 Sep. 1952 and 31 Dec. 1952, leg. Alexandre Fain (RBINS: 21666, RMCA: 81.012-M-0044 to 81.012M-0046); 1 juv ?, 1 ? [2 SS], Shabunda, 16 Aug. 1952, leg. P.G. Vercammen (RMCA: 23177, 23178);
2 ad ♀♀, 3 juv ??, 1 ♀, 3 ?? [5 SS, 4 AL], Thysville [= Mbanza-Ngungu], 29 Mar. 1964, 29 Apr. 1964,
prior to 18 Aug. 1965, leg. Antoon Emeric Marcel De Roo (RMCA: 32655, 32656, 32767, 32777 to
32779, 32388, 32389, 33568); 3 ?? [1 SA, 2 AL], Thysville Cave b 13 a [= Banjan Caves], 23 May
1949, 17 Aug. 1949, leg. Narcisse Leleup (RMCA: 22671, 22672, 22503); 2 ♂♂ [2 SS], Wago, Mount,
prior to 16 Oct. 1952, leg. Alexandre Fain (RMCA: 21436, 21437); 2 ?? [2 SK], Yoko, unknown date,
leg. Prescott Musaba (RMCA: a1.097-M-3162, a1.097-M-3163).
RWANDA: 1 ♀, 1 ? [2 AL], Gisovu, 2 Oct. 1980, 31 Oct. 1980, leg. Danny Meirte (RMCA: 80.055-M0096, 80.055-M-0097); 1 ? [SA], Kibuye, 2 Dec. 1971, leg. Alena Vanderborght-Elbl, G. Mathys and
A. Elbl (RMCA: 73.058-M-0002); 32 ad ♀♀, 10 ad ♂♂, 15 sad ♀♀, 4 sad ♂♂, 1 juv ♂ [8 SA, 54 AL],
Ntango, 24 May 1982, 25 May 1982, 26 May 1982, leg. Benny Baeten (RMCA: 87.046-M-0063 to
87.046-M-0124); 2 ♀♀ [1 SA, 1 AL], Nyungwe Forest Reserve, 18 Oct. 1976, leg. Frieder Meier
(MHNG: 1872.081, 1872.082); 1 ad ♀, 3 ad ♂♂, 4 sad ♀♀, 5 sad ♂♂ [13 AL], Routabansougera,
14 Jul. 1982, 15 Jul. 1982, 17 Jul. 1982, leg. Benny Baeten, 19 Oct. 1981, 20 Oct. 1981, leg. Frits
De Vree, Benny Baeten and Victor Van Cakenberghe (RMCA: 87.046-M-0125 to 87.046-M-0134,
82.006-M-0040, 82.006-M-0042, 82.006-M-0041); 1 sad ♀, 2 sad ♂♂ [3 AL], Rukira, 22 Nov. 1981,
23 Nov. 1981, 26 Nov. 1981, leg. Frits De Vree, Benny Baeten and Victor Van Cakenberghe (RMCA:
82.006-M-0043 to 82.006-M-0045); 1 ad ♀ [SA], Shangi, 5 Jan. 1985, leg. Benny Baeten and Luc
Janssens (RMCA: 85.006-M-0464); 1 ad ♀, 1 ad ♂, 1 sad ♀, 1 sad ♂ [2 SA, 2 AL], Uwinka, 16 Oct. 1981,
17 Oct. 1981, 18 Oct. 1981, leg. Frits De Vree, Benny Baeten and Victor Van Cakenberghe (RMCA:
82.006-M-0036 to 82.006-M-0039).
Myonycteris relicta Bergmans, 1980
DEMOCRATIC REPUBLIC OF THE CONGO: 1 ♂ [SS], Beyi - Kalonji, 22 Feb. 1980, leg. Betu
Nsankulu (RMCA: 80.020-M-0001).
Myonycteris torquata (Dobson, 1878)
DEMOCRATIC REPUBLIC OF THE CONGO: 4 ?? [3 AL, 1 SK], unknown locality, unknown date,
leg. Prescott Musaba, between 1 May 2010 and 31 May 2010, leg. Nicolas Nesi, Blaise Kadjo and
Christiane Denys (RMCA: a1.097-M-3194 to a1.097-M-3196, MNHN: ZM-2014-1168); 1 ? [UN],
Albert National Park [= Virunga National Park], unknown date, leg. unknown collector
(INPBC: V1694.2); 1 ♂ [SS], Bakwakenge [= Bakwa-Kenge], 3 Feb. 1980, leg. Betu Nsankulu
(RMCA: 80.020-M-0005); 4 ♀♀, 2 ♂♂ [1 SS, 5 SK], Baliko, 11 Jun. 2012, 12 Jun. 2012, 13 Jun. 2012,
14 Jun. 2012, leg. William (“Bill”) T. Stanley (FMNH: 219592, 219593, 219680, 219594 to 219596);
2 ♂♂ [2 SS], Bena Mbala, 21 Feb. 1984, leg. Betu Nsankulu (RMCA: 84.035-M-0004, 84.035-M155
European Journal of Taxonomy 382: 1–327 (2017)
0006); 6 ♂♂, 4 ?? [10 AL], Boende, between 1 Jan. 1974 and 31 Dec. 1974, between 1 Jan. 1967 and
31 Dec. 1967, leg. P. Lootens (RMCA: 38650, 38651, 38671 to 38676, 75.035-M-0002, 75.035-M0003); 4 ♀♀, 5 ♂♂ [9 UN], Bomane [= Bomaneh], 21 May 2010, 30 May 2010, leg. Guy-Crispin
Gembu Tungaluna, Prescott Musaba and José Akaibe (UNIKIS: CRT2127, CRT2130, CRT2144,
CRT2156, CRT2157, CRT2195, CRT2196, CRT2200, CRT2078); 1 ♂ [SS], Cinputu, 13 Sep. 1983,
leg. Betu Nsankulu (RMCA: 84.035-M-0005); 2 ♂♂ [2 UN], Congo-Nil-Aka, 20 May 1952, leg. H. de
Saeger et al., 20 May 1952, leg. H. de Saeger (RBINS: 13525, RMCA: 13525); 14 ad ♀♀, 17 ad ♂♂,
15 sad ♀♀, 14 sad ♂♂, 38 ♀♀, 40 ♂♂ [1 AL, 60 SK, 77 UN], Djabir, unknown date, 8 May 2011,
9 May 2011, 10 May 2011, 12 May 2011, leg. Prescott Musaba, 8 May 2011, 9 May 2011, 10 May 2011,
11 May 2011, 12 May 2011, 13 May 2011, leg. Prescott Musaba and Jonathan Kosele
(RMCA: a1.097-M-3248, a1.097-M-3249, a1.097-M-3265 to a1.097-M-3268, a1.097-M-3282,
a1.097-M-3250 to a1.097-M-3264, UNIKIS: DJB083, DJB084, DJB087, DJB088, DJB091, DJB092,
DJB094, DJB097, DJB098, DJB102, DJB104, DJB106, DJB109, DJB111, DJB122,
RMCA: a1.097-M-3269 to a1.097-M-3281, UNIKIS: DJB127, DJB128, DJB130, DJB133, DJB139,
DJB142, DJB147, DJB154 to DJB167, DJB170, RMCA: a1.097-M-3223 to a1.097-M-3230,
UNIKIS: DJB001, DJB003, DJB006, DJB008 to DJB011, DJB014, DJB016, DJB018 to DJB020,
DJB022 to DJB024, DJB026, DJB027, DJB029, DJB030, DJB032, DJB034, DJB036, DJB037,
RMCA: a1.097-M-3231 to a1.097-M-3247, UNIKIS: DJB040, DJB044, DJB046, DJB048 to DJB050,
DJB052, DJB053, DJB055, DJB056, DJB058, DJB060, DJB062, DJB067, DJB070, DJB071, DJB169,
DJB172); 5 ♀♀, 2 ♂♂ [7 UN], Epulu, 29 Aug. 2015, 30 Aug. 2015, leg. André Malekani
(UNIKIS: EPLU660, EPLU669, EPLU670, EPLU683, EPLU684, EPLU695, EPLU696); 1 imm ♂, 1 ♀
[2 UN], Gangala-na-Bodio, 11 May 1948, leg. unknown collector (RBINS: 11657, RMCA: 11657);
4 ♀♀, 16 ♂♂ [20 UN], Île Mbiye [= Mbiye Island], 16 Jan. 2012, 25 Feb. 2012, 30 Mar. 2012,
31 Mar. 2012, leg. Prescott Musaba, Tembele and Marc Boketshu, 19 Apr. 2014, 21 Apr. 2014, leg.
Prescott Musaba, Moïse Bipoo, Richard Tamaru, Pauline Isude and Joseph Kussa, 7 Aug. 2015,
28 Dec. 2015, 8 Jan. 2016, leg. Prescott Musaba, Moïse Bipoo, Charle Andabhati, Benjamin Kirongozi
and Kambale Karupao (UNIKIS: CHIM07, CHIM07/14, CHIM011/14, CHIM012/14, CHIM29,
ABCHIM07, ABCHIM08, CHIM34, CHIM35, CHIM40 to CHIM45, CHIM47, CHIMA38, CHIMTB01,
ABCHIM09, ABCHIM11); 1 ♀ [SA], Irangi, 11 Feb. 1984, leg. Heinz Stephan (SMF: 65030); 2 ♀♀
[2 UN], Isai River, between 16 Dec. 1974 and 23 Dec. 1974, leg. Michael D. Gallagher (HZM: 1.7798,
2.7799); 1 ♀ [UN], Kamikoni, 17 May 1960, leg. unknown collector (RBINS: 1694); 1 imm ♀ [UN],
Karambi, 4 Apr. 1992, leg. Ngenge Masumbuko Kamitongo (ZMA: MAM.24567); 1 ♀ [SO], Kimpoko
[= Kimpoko-Nsele], 7 Nov. 1978, leg. Tshinyenye (RMCA: 79.003-M-0001); 1 ♂ [UN], Kingabwa,
31 Dec. 1975, leg. Tshinyenye (RMCA: 76.010-M-0004); 1 imm ? [UN], Kinshasa, 2 May 1964, leg.
Jo Van Orshoven (ZMA: MAM.11163); 21 ?? [21 UN], Kinshasa City Province, 12 Feb. 2001, between
1 Apr. 1999 and 30 Apr. 1999, leg. unknown collector (SMF 92979 to 92983, 93281 to 93296); 1 ♀, 1 ♂
[2 AL], Kisangani, between 1 Jun. 1982 and 30 Jun. 1982, leg. Marc Colyn (RMCA: 82.031-M-0012,
82.031-M-0013); 2 ♂♂ [1 SK, 1 SO], Kitangua, 24 Dec. 1981, leg. Betu Nsankulu (RMCA: 83.025-M0006, 83.025-M-0019); 20 ♀♀, 8 ♂♂, 1 ? [13 SK, 16 UN], Kona, unknown date, leg. Prescott Musaba,
12 May 2010, 13 May 2010, 14 May 2010, 15 May 2010, between 12 May 2010 and 13 May 2010, leg.
Guy-Crispin Gembu Tungaluna, Prescott Musaba and José Akaibe (RMCA: a1.097-M-3160,
UNIKIS: CRT1910, CRT1911, CRT1920, CRT1926, CRT1937, CRT1942 to CRT1945, CRT1947 to
CRT1950, CRT1953, CRT1960, CRT1963, CRT1966 to CRT1972, CRT1979 to CRT1982, CRT1918);
3 ?? [3 AL], Konyola, prior to 5 Aug. 1988, leg. Luc De Vos, Kimbembi and Alain Vandelanoote
(RMCA: 88.025-M-0005 to 88.025-M-0007); 2 ♂♂ [2 UN], Kumbi, 7 Dec. 1993, leg. Ngenge
Masumbuko Kamitongo (ZMA: MAM.24896, MAM.24897); 13 ♀♀, 14 ♂♂, 1 ? [1 SK, 27 UN], Lieki,
21 May 2010, 30 May 2010, 1 Jun. 2010, 2 Jun. 2010, 3 Jun. 2010, leg. Guy-Crispin Gembu Tungaluna,
Prescott Musaba and José Akaibe (UNIKIS: CRT2029, CRT2030, CRT2032 to CRT2035, CRT2037,
CRT2282 to CRT2288, CRT2290 to CRT2296, CRT2057, CRT2021 to CRT2023, CRT2028, CRT2050,
CRT2051); 1 ad ♂ [UN], Likandi River [= Litaki River], 18 Apr. 1906, leg. The Alexander-Gosling
156
VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
Expedition (BMNH: 1907.7.8.25 [holotype Myonycteris wroughtoni K. Andersen, 1908]); 1 ♂ [UN],
Likati River, 18 Apr. 1906, leg. unknown collector (BMNH: 1907.7.8.26); 14 ?? [14 SS], Lovanium
University [= Kinshasa], 10 Jan. 1962, 13 Jan. 1962, 2 May 1962, leg. De Bont, 22 Nov. 1962,
3 Dec. 1962, 20 Feb. 1963, 17 Mar. 1963, 23 Mar. 1963, 30 Mar. 1963, 2 Apr. 1963, 19 Apr. 1963,
26 Dec. 1966, between 30 Mar. 1963 and 1 Apr. 1963, between 1 Nov. 1962 and 1 Dec. 1962, leg.
De Bont and Jo Van Orshoven (RMCA: 31197, 31198, 32606, 32608, 32611, 31199, 32609, 32602,
32607, 32601, 32604, 32605, 32610, 32603); 1 ♂ [AL], Lubumbashi, 12 Oct. 2009, leg. Alexandre
Hassanin, Nicolas Nesi, Célestin Pongombo, Peter Vallo and Didier Tshikung (MNHN: ZM-2011-859);
1 ♂ [SS], Lukonga, 7 Oct. 1978, leg. Betu Nsankulu (RMCA: 79.022-M-0006); 12 ?? [3 SS, 9 AL],
Luluabourg [= Kananga], 1 May 1964, 7 Nov. 1964, 13 Feb. 1965, 20 Feb. 1965, 4 Mar. 1965,
1 Apr. 1965, 2 Apr. 1965, prior to 1 Jun. 1965, leg. Antoon Emeric Marcel De Roo (RMCA: 33336,
33338, 33413, 33414, 32660, 33333, 33339, 33340, 33334, 33335, 33332, 33337); 1 ♂ [UN], Lwana
River, unknown date, leg. Ngenge Masumbuko Kamitongo (ZMA: MAM.24568); 2 sad ♂♂, 1 juv ♀,
27 ♀♀, 45 ♂♂ [4 AL, 62 SK, 9 UN], Masako, unknown date, 27 Nov. 1987, 28 Nov. 1987, 29 Nov. 1987,
30 Nov. 1987, 16 Dec. 1987, 17 Dec. 1987, 22 Apr. 1988, 25 Aug. 1989, 29 Aug. 1989, 30 Aug. 1989,
24 Nov. 1989, 22 Mar. 1990, 25 Mar. 1990, leg. Ndey B. Ifuta, 10 Jun. 2015, 11 Jun. 2015, 13 Jun. 2015,
leg. Prescott Musaba, Moïse Bipoo, Charle Andabhati, Benjamin Kirongozi and Kambale Karupao
(RMCA: 93.079-M-0278, UNIKIS: CHMS062, CHMS063, CHMS065, CHMS066, CHMS070 to
CHMS072, CHMS075, CHMS077, RMCA: 93.079-M-0043 to 93.079-M-0051, 93.079-M-0287 to
93.079-M-0291, 93.079-M-0052 to 93.079-M-0059, 93.079-M-0228, 93.079-M-0292, 93.079-M-0276,
93.079-M-0272, 93.079-M-0275, 93.079-M-0277, 93.079-M-0270, 93.079-M-0026, 93.079-M-0027,
93.079-M-0214 to 93.079-M-0216, 93.079-M-0028 to 93.079-M-0033, 93.079-M-0217 to 93.079-M0219, 93.079-M-0267, 93.079-M-0273, 93.079-M-0034, 93.079-M-0220, 93.079-M-0284, 93.079-M0274, 93.079-M-0035 to 93.079-M-0042, 93.079-M-0221 to 93.079-M-0226, 93.079-M-0285, 93.079M-0286); 2 ad ♀♀, 1 ad ♂, 1 imm ♀ [4 UN], Medje, 16 Apr. 1910, 6 Sep. 1910, 4 Apr. 1914, leg.
Herbert Lang, James Paul Chapin and The American Museum Congo Expedition (AMNH: 48752,
48755, 48753, 48754); 3 ♂♂ [3 AL], Pandé, Forest N of, 11 Oct. 2009, leg. Alexandre Hassanin,
Nicolas Nesi, Célestin Pongombo, Peter Vallo and Didier Tshikung (MNHN: ZM-2011-856 to ZM2011-858); 6 ♀♀ [1 SS, 1 AL, 4 SK], Quatorze, 15 Jun. 2012, 25 Jun. 2013, 29 Jun. 2013, 30 Jun. 2013,
leg. William (“Bill”) T. Stanley (FMNH: 219597, 222421, 222622, 222422 to 222424); 6 ♀♀, 3 ♂♂
[9 UN], Rubi-Tele, 18 Aug. 2013, 19 Aug. 2013, 20 Aug. 2013, 22 Aug. 2013, 27 Aug. 2013, leg. GuyCrispin Gembu Tungaluna, Prescott Musaba and André Malekani (UNIKIS: RBTL502, RBTL503,
RBTL516, RBTL520, RBTL529, RBTL530, RBTL541, RBTL542, RBTL575); 2 ♀♀, 1 ? [1 SK, 2 UN],
Sciere Forest, between 24 Nov. 1974 and 25 Nov. 1974, leg. Michael D. Gallagher (HZM: 3.7866 to
5.7868); 1 ♀, 2 ♂♂ [3 SS], Topetope, 10 Jun. 1966, 5 Jul. 1966, 10 Jul. 1966, leg. Alexandre Prigogine
(RMCA: 34381, 34383, 34382); 3 ♂♂ [3 UN], Uma, 10 Apr. 2014, 12 Apr. 2014, 13 Apr. 2014, leg.
Guy-Crispin Gembu Tungaluna, Prescott Musaba and André Malekani (UNIKIS: UMA501, UMA624,
UMA635); 14 ?? [14 AL], Wafanya [= Wafania], 3 Dec. 1988, 2 Sep. 1991, 14 Sep. 1991, 3 Oct. 1991,
18 Apr. 1992, 12 May 1992, 29 May 1992, between 1 Nov. 1991 and 30 Nov. 1991, between 1 Apr. 1991
and 30 Apr. 1991, between 1 Jan. 1991 and 31 Dec. 1991, leg. P. Lootens (RMCA: 92.079-M-0068,
91.076-M-0082, 92.079-M-0065, 92.079-M-0070, 91.076-M-0065, 92.079-M-0066, 92.079-M-0069,
91.076-M-0098, 91.076-M-0100, 89.020-M-0039, 92.079-M-0063, 91.076-M-0020, 92.079-M-0064,
92.079-M-0067); 41 ♀♀, 73 ♂♂ [37 SK, 77 UN], Yaekela, 2 May 2010, 3 May 2010, 4 May 2010,
5 May 2010, 6 May 2010, 7 May 2010, between 5 May 2010 and 13 May 2010, between 5 May 2010
and 12 May 2010, leg. Guy-Crispin Gembu Tungaluna, Prescott Musaba and José Akaibe
(UNIKIS: CRT2305, CRT2307, CRT2316 to CRT2318, CRT2320, CRT2351 to CRT2360, CRT2362,
CRT2365, CRT2366, CRT2369 to CRT2373, CRT2375 to CRT2383, CRT2385, CRT1801, CRT1802,
CRT1804 to CRT1808, CRT1810, CRT1811, CRT1815, CRT1819, CRT1820, CRT2391, CRT2393,
CRT2394, CRT2397 to CRT2400, CRT1824 to CRT1830, CRT1833 to CRT1840, CRT1842 to CRT1845,
CRT1847, CRT1848, CRT1850 to CRT1853, CRT1855, CRT1857, CRT1859 to CRT1866, CRT1869 to
157
European Journal of Taxonomy 382: 1–327 (2017)
CRT1878, CRT1883 to CRT1887, CRT1889 to CRT1896, CRT1831, CRT1832, CRT1841); 1 ♀, 2 ♂♂
[2 SS, 1 AL], Yalosemba, 18 Jun. 1979, 19 Jun. 1979, leg. C. Brian Robbins, 23 Jun. 1979, leg. Frieder
Meier (USNM: 537694, 537695, MHNG: 1871.014); 3 ♀♀, 11 ♂♂, 1 ? [3 AL, 1 SK, 11 UN], Yoko,
unknown date, 4 Sep. 2012, 7 Sep. 2012, leg. Prescott Musaba, 18 May 2009, 23 Sep. 2009, 24 Sep. 2009,
28 Sep. 2009, leg. unknown collector, 4 Sep. 2012, 5 Sep. 2012, 6 Sep. 2012, leg. Prescott Musaba,
Tembele and Marc Boketshu (RMCA: a1.097-M-3161, UNIKIS: GE246, GE281, GE284, GE293,
RMCA: a1.097-M-3192, a1.097-M-3193, UNIKIS: YK005CH, YK010CH, YK013CH, YK015CH,
YK018CH, YK019CH, YK025CH, RMCA: a1.097-M-3191); 2 ♂♂ [2 UN], Zookis [= Kisangani],
24 Dec. 2015, leg. Prescott Musaba, Moïse Bipoo, Charle Andabhati, Benjamin Kirongozi and Kambale
Karupao (UNIKIS: ZCH05, ZCH12).
Plerotini Bergmans, 1997
Plerotes K. Andersen, 1910
Plerotes anchietae (Seabra, 1900)
DEMOCRATIC REPUBLIC OF THE CONGO: 1 ♂ [AL], Kaloba, 5 km from, 3 Oct. 2009, leg.
Alexandre Hassanin, Nicolas Nesi, Célestin Pongombo, Peter Vallo and Didier Tshikung (MNHN: ZM2011-860); 1 ad ♀ [SO], Kasai, 17 Jan. 1982, leg. Betu Nsankulu (RMCA: 83.025-M-0021); 2 ♂♂
[1 SS, 1 SO], Kipangaribwe River, 10 Dec. 1947, 14 Jan. 1948, leg. Gaston-François de Witte et al.
(RBINS: 10685, 10686); 1 ♂ [UN], Kisantu, 11 Dec. 1920, leg. Henri Schouteden (RMCA: 7300b).
Rousettini K. Andersen, 1912
Rousettus Gray, 1821
DEMOCRATIC REPUBLIC OF THE CONGO: 1 juv ? [UN], Djomba [= Djemba], unknown date, leg.
unknown collector (INPBC: V93.1); 1 juv ? [UN], Kabambi, unknown date, leg. unknown collector
(INPBC: W1467); 1 ? [AL], Mount Hoyo [= Hoyo, Mount], unknown date, leg. The Cambridge East
Africa Expedition 1959 (BMNH: 1960.113); 1 juv ? [UN], Rutshuru, unknown date, leg. unknown
collector (INPBC: V1447).
Rousettus aegyptiacus (E. Geoffroy St.-Hilaire, 1810)
BURUNDI: 1 ♂ [UN], Bujumbura, 1 Aug. 1976, leg. Jacques Verschuren (RBINS: 19903); 1 juv ♀
[SA], Gihanga, 14 Mar. 1979, leg. Heinz Stephan (SMF: 57530); 1 ♀ [SK], Kayanza, 27 Mar. 1979,
leg. Heinz Stephan (SMF: 57531); 1 ♀, 1 ♂ [2 SO], Kigwena Hill, 13 Oct. 1990, leg. Matthew Criblez
(FMNH: 148339, 148340); 1 ♀ [SB], Nyagatarugwa River Valley, 9 Aug. 1992, leg. L. Davenport
(FMNH: 148338); 2 ♀♀ [2 SK], Nyanza-Lac, 3 Mar. 1979, leg. Heinz Stephan (SMF: 57528, 57529);
2 ♀♀, 1 ♂ [2 SK, 1 CO], Resha, 2 Mar. 1979, 3 Apr. 1979, 4 Apr. 1979, leg. Heinz Stephan (SMF:
57526, 57525, 57527); 3 ?? [3 SS], Rumonge, 22 Jul. 1944, leg. J.M. Vrijdagh (RMCA: 17759 to
17761).
DEMOCRATIC REPUBLIC OF THE CONGO: 1 ♀, 4 ♂♂, 4 ?? [5 SS, 3 AL, 1 SK], unknown locality,
unknown date, leg. Guy-Crispin Gembu Tungaluna, between 1 May 2010 and 31 May 2010, leg. Nicolas
Nesi, Blaise Kadjo and Christiane Denys, prior to 12 Jun. 1958, leg. Alexandre Fain, prior to 14 Dec. 1957,
leg. Félix / Michel Anciaux de Faveaux (RMCA: a1.097-M-0548, a1.097-M-0549, MNHN: ZM-20141156, RMCA: 27182 to 27186, 26823); 4 ?? [4 UN], Albert National Park [= Virunga National
Park], unknown date, leg. unknown collector (INPBC: V1765, V1839.3, V5006, V705); 1 ♀, 2 ♂♂
[3 AL], Baliko, prior to 31 Jan. 2007, leg. Guy-Crispin Gembu Tungaluna (RMCA: a1.097-M-0414 to
a1.097-M-0416); 8 ?? [8 AL], Banza Ninda Caves, 15 Jun. 1949, leg. Narcisse Leleup (RMCA: 22511
to 22514, 22566 to 22569); 3 ?? [1 SS, 1 SK, 1 SO], Beni, prior to 2 Jun. 1914, leg. Bonnevie (RMCA:
158
VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
3238a, 3238b, 3239); 1 ♀ [SS], Bushushu, 20 Jun. 1959, leg. Uwe Rahm (RMCA: 28893); 1 ad ♂,
1 imm ♂ [2 SS], Buta, 24 Dec. 1946, leg. F.L. Hendricks (RBINS: 12932, 12933); 2 ♀♀, 1 ♂ [2 SS,
1 AL], Butembo, 1 Jan. 1970, leg. M.J. Celis and M. Lejeune, 22 Jan. 1958, 26 Jul. 1960, leg. P. Dyleff
(RMCA: 36206, 27384, 29242); 1 ♀ [SS], Butembo Region, 31 Jul. 1947, leg. Jean Hiernaux (RBINS:
7054); 3 ♀♀, 3 ♂♂, 24 ?? [1 SS, 27 AL, 1 UN, 1 AB], Defrenne Cave [= Mwanga Cave], 19 Jun. 1957,
leg. Félix / Michel Anciaux de Faveaux, 25 Sep. 1948, leg. Narcisse Leleup, 3 Apr. 1957, leg.
G. Verhoustraete (RMCA: 26711 to 26715, 26798, 22615 to 22637, 26214); 18 ?? [18 UN], Djemba,
unknown date, leg. unknown collector (INPBC: V417, V93.1.01 to V93.1.17); 1 ? [UN], Djomba [=
Djemba], unknown date, leg. unknown collector (INPBC: V1535); 1 ? [AL], Elisabethville [=
Lubumbashi], prior to 15 Oct. 1963, leg. Parent (RMCA: 31509); 97 ♀♀, 108 ♂♂, 1 ? [206 UN],
Goroumbwa Mine, 14 May 1999, 17 May 1999, 18 May 1999, 19 May 1999, 19 Oct. 1999, 20 Oct. 1999,
21 Oct. 1999, 22 Oct. 1999, 23 Oct. 1999, 24 Oct. 1999, 25 Oct. 1999, 27 Oct. 1999, leg. Robert (Bob)
Swanepoel (TM: DRC0001 to DRC0004, DRC0028, DRC0030, DRC0031, DRC0035 to DRC0052,
DRC0054, DRC0057 to DRC0068, DRC0070, DRC0072, DRC0073, DRC0087, DRC0088, DRC0090
to DRC0101, DRC0108 to DRC0119, DRC0123, DRC0125, DRC0126, DRC0132, DRC0133,
DRC0136 to DRC0140, DRC0143 to DRC0145, DRC0152, DRC0153, DRC0157, DBA002 to DBA007,
DBA009, DBA029, DBA070, DBA074, DBA077 to DBA084, DBA110, DBA111, DBA114, DBA116
to DBA123, DBA125, DBA127, DBA128, DBA131, DBA142, DBA157, DBA187 to DBA193,
DBA197, DBA226, DBA229, DBA231 to DBA238, DBA254, DBA281, DBA284 to DBA286, DBA335
to DBA353, DBA355 to DBA359, DBA361 to DBA400, DBA407); 3 ♀♀, 6 ♂♂ [3 AL, 6 UN],
Jadotville [= Likasi], 3 Apr. 1957, 6 Dec. 1959, 18 Dec. 1959, leg. Félix / Michel Anciaux de Faveaux
(RBINS: 14600, RMCA: 26258 to 26264, RBINS: 14601); 5 ♂♂ [5 AL], Jomba, 4 Sep. 1976, leg.
Frieder Meier (MHNG: 1872.062 to 1872.066); 1 ad ♂ [AL], Kadjudju [= Kajuju], 16 Apr. 1905, leg.
unknown collector (MNHN: ZM-MO-1983-1267); 2 ?? [2 SS], Kakindo Cave [= Kakondo Cave],
27 Jul. 1962, 28 Jul. 1962, leg. A. Elbl (USNM: 375234, 375235); 1 ♂ [SS], Kakonda, 8 Oct. 1942, leg.
F.L. Hendricks (AMNH: 118865); 6 ♀♀, 4 ♂♂ [6 SS, 2 SK, 2 UN], Kakondo, 1 Dec. 1946, 24 Dec. 1946,
leg. F.L. Hendricks, 1 Dec. 1946, leg. unknown collector, 14 Jan. 1959, leg. Uwe Rahm (RBINS:
13114.1, 13114.2, 13283, 13284, RMCA: 28031, 28890 to 28892, 19652, 19653); 1 ♀, 1 ♂ [2 SS],
Kakondo Cave, between 1 Dec. 1946 and 31 Dec. 1946, leg. F.L. Hendricks (RMCA: 19650, 19655);
3 ♀♀, 1 ♂ [4 AL], Kakontwe, unknown date, leg. the Royal Museum for Central Africa (RMCA)
(BMNH: 1954.778 to 1954.781); 1 ad ♀, 2 ?? [3 SS], Kakontwe Cave, between 1 Sep. 1948 and
30 Sep. 1948, leg. Narcisse Leleup, between 1 Dec. 1958 and 31 Dec. 1958, leg. Laurent (RMCA:
22483, 22484, 29219); 1 ? [AL], Kalimabenge River, 12 Jul. 1958, leg. Georges Marlier (RMCA:
27988); 3 ?? [3 AL], Kasuo Cave Kabwe ka, 26 Jan. 1955, leg. Wim Bergmans and Collard (RMCA:
22955a to 22955c); 1 sad ♀, 12 ♀♀, 11 ♂♂ [3 SS, 1 SA, 10 AL, 10 UN], Katana, unknown date, leg.
J.J. Laarman, 11 Jul. 1956, leg. Alvin Novick, 22 Jul. 1954, leg. James Paul Chapin, 30 May 1963, leg.
P. Kunkel (RMNH: MAM.16347 to MAM.16355, MAM.16401, MCZ: 48186 to 48194, 48536, AMNH:
180899 to 180901, SMF: 31823); 1 ♀ [SS], Katana Cave [= Mahyusha Cave], 23 Jan. 1957, leg.
Heinz Stephan (RMCA: 27419); 2 ♂♂ [2 SS], Kibanda, 15 May 1975, leg. Alexandre Prigogine
(RMCA: 76.022-M-0021, 76.022-M-0022); 1 ♀, 1 ♂ [2 SS], Kilo Mines [= Kilomines], 16 Apr. 1949,
17 Apr. 1949, leg. Jean Hiernaux (RMCA: 19100, 19101); 1 ♀, 1 ♂ [2 AL], Kinyamahura, 11 Apr. 1938,
leg. Serge Frechkop (RMCA: 17083, 17084); 6 ?? [6 AL], Kisangani, between 1 Jul. 1985 and
31 Jul. 1985, leg. Marc Colyn, between 1 Oct. 1985 and 31 Mar. 1986, leg. Ndey B. Ifuta (RMCA:
86.024-M-0001, 86.024-M-0002, 87.010-M-0007 to 87.010-M-0010); 1 ad ♀, 1 ad ♂, 1 juv ♀, 1 juv ♂,
1 ♀ [1 SA, 4 AL], Kivu [= Kivu region], 10 Apr. 1905, leg. unknown collector, 19 Jul. 1964, leg.
P. Kunkel (MNHN: ZM-MO-1983-1076 to ZM-MO-1983-1079, SMF: 31824); 2 ?? [2 AL], Kivu
region, between 1 Jan. 1934 and 31 Dec. 1934, leg. Guy Babault (MCZ: 46063, 46064); 1 ♀ [SO],
Kunungu, unknown date, leg. Henri Schouteden (RMCA: 7291a); 1 ♀, 1 ♂ [1 SO, 1 UN], Kyniamahura,
7 May 1938, leg. Serge Frechkop (RBINS: 4784, 4785); 2 ?? [2 AL], Lake Kivu, near, between
1 Jan. 1935 and 31 Dec. 1935, leg. Guy Babault (MCZ: 46154, 46155); 2 sad ♀♀, 1 juv ♀, 6 ♀♀, 4 ♂♂,
159
European Journal of Taxonomy 382: 1–327 (2017)
9 ?? [6 SS, 5 SA, 5 AL, 4 SO, 2 SB], Lwiro [= Luiro], unknown date, 2 Jun. 1964, 22 Jun. 1965,
between 1 Jan. 1965 and 31 Dec. 1965, leg. P. Kunkel, 16 Jan. 1962, leg. Kiss, 6 May 1955, 16 May
1955, leg. Waldo LaSalle Schmitt and E. Baker, 16 May 1955, leg. Waldo LaSalle Schmitt, 31 Jul. 2002,
leg. Michael H. Huhndorf, 7 Apr. 1959, 8 Apr. 1959, leg. Uwe Rahm (SMF: 25859, RMCA: 31004 to
31007, USNM: 301694, 301696 to 301702, SMF: 31822, 31821, FMNH: 173295, 173296, USNM:
301695, RMCA: 28894 to 28896, SMF: 25860); 1 ♂ [SS], Mahysha Cave [= Mahyusha Cave],
unknown date, leg. Alexandre Fain (RMCA: 27717); 2 ♀♀ [2 SS], Mahyusa Cave [= Mahyusha
Cave], unknown date, leg. unknown collector (BMNH: 1958.647, 1958.648); 1 ♀, 1 ? [2 SS], Mai-yaMoto [= May ya Moto], 16 May 1955, leg. James Paul Chapin (AMNH: 180897, 180898); 2 ad ♂♂,
1 sad ♂, 10 ♀♀, 6 ♂♂, 2 ?? [2 AL, 19 SK], Masako, 22 May 1986, 24 Nov. 1987, 26 Nov. 1987, 22 May
1989, 23 May 1989, 24 May 1989, 25 May 1989, 21 Jan. 1990, between 1 May 1989 and 31 May 1989,
leg. Ndey B. Ifuta (RMCA: 93.079-M-0317, 87.010-M-0002, 87.010-M-0006, 93.079-M-0318, 93.079M-0305 to 93.079-M-0309, 93.079-M-0315, 93.079-M-0316, 93.079-M-0281, 93.079-M-0310 to
93.079-M-0312, 93.079-M-0319, 93.079-M-0282, 93.079-M-0283, 93.079-M-0313, 93.079-M-0320,
93.079-M-0321); 1 ♂ [SS], May ya Moto, sources chaudes [= May ya Moto], 22 Aug. 1949, leg. J. de
Wilde (RBINS: 12930); 1 juv ♀, 1 juv ♂, 2 ♀♀, 3 ♂♂ [5 SS, 1 SO, 1 UN], Mount Hoyo [= Hoyo,
Mount], 12 Aug. 1947, leg. Jean Hiernaux, 12 Aug. 1947, leg. unknown collector (RBINS: 7057 to
7061, 7064, 7474); 1 imm ♀ [AL], Mufunga, 23 Jul. 1958, leg. Félix / Michel Anciaux de Faveaux
(RMCA: 27915); 1 imm ? [AL], Mulungu, prior to 1 Feb. 1940, leg. F.L. Hendricks (RMCA: 15879);
1 ♂ [SS], Mumba, 7 Jun. 1967, leg. Alexandre Prigogine (RMCA: 35112); 1 ? [UN], Murambi,
unknown date, leg. unknown collector (INPBC: W831); 1 ♂ [SS], Mutsora, 27 Sep. 1949, leg. J. de
Wilde (RBINS: 12931); 2 ♀♀, 1 ♂ [3 SS], Nyarusambo, 7 Jan. 1947, leg. G. Van Cools (RBINS: 12871
to 12873); 15 ad ♀♀, 1 ad ♂, 5 juv ♀♀, 2 juv ♂♂, 2 imm ♀♀, 1 imm ♂, 4 ♀♀, 2 ♂♂, 1 ? [2 SA, 31 AL],
Pempere Cave, 19 Jun. 1957, 20 Oct. 1957, 21 Apr. 1958, leg. Félix / Michel Anciaux de Faveaux
(MHNG: 1046.032, RMCA: 26777 to 26780, MHNG: 1046.033, 1046.034, RMCA: 27596 to 27606,
27608 to 27621, 27595); 1 ♀ [UN], Rethy, 23 Oct. 2015, leg. Anne Laudisoit et al. (UNIKIS: ITU702);
1 ? [UN], Rumoka, unknown date, leg. unknown collector (INPBC: V1636.1); 2 ?? [2 UN], Rutshuru,
unknown date, leg. unknown collector (INPBC: V1247, V1625); 1 ♂ [SS], Ruvara, 2 Apr. 1950, leg.
J. de Wilde (RBINS: 12929); 2 ♀♀, 13 ?? [11 SA, 4 AL], Tamba Tamba Cave b 12, unknown date, leg.
the Royal Museum for Central Africa (RMCA), 27 Sep. 1948, leg. Narcisse Leleup (BMNH: 1954.782,
1954.783, RMCA: 22673 to 22685); 15 ♀♀, 9 ♂♂ [6 AL, 3 SK, 15 UN], Uma, 18 May 2012,
16 Aug. 2012, 17 Aug. 2012, 18 Aug. 2012, 10 Apr. 2014, 11 Apr. 2014, 12 Apr. 2014, leg. Guy-Crispin
Gembu Tungaluna, Prescott Musaba and André Malekani, 16 Aug. 2012, 17 Aug. 2012, 18 Aug. 2012,
leg. Prescott Musaba (UNIKIS: UMA540, UMA548, UMA552, UMA549, UMA562, UMA579,
UMA580, UMA582, UMA585, UMA634, UMA610, UMA611, UMA620, UMA633,
RMCA: a1.097-M-3166, a1.097-M-3167, a1.097-M-3169, a1.097-M-3171, UNIKIS: UM013,
RMCA: a1.097-M-3170, UNIKIS: UM014, UM025, RMCA: a1.097-M-3168, UNIKIS: UM021); 3 ♀♀
[3 AL], Upemba National Park, 25 Sep. 2009, leg. Alexandre Hassanin, Nicolas Nesi, Célestin
Pongombo, Peter Vallo and Didier Tshikung (MNHN: ZM-2011-861 to ZM-2011-863); 9 ♀♀, 5 ♂♂
[14 UN], Yoko, 11 Feb. 2009, 12 Feb. 2009, 12 May 2009, 13 May 2009, 18 May 2009, 19 May 2009,
20 May 2009, 19 Jul. 2009, 25 Sep. 2009, leg. unknown collector (UNIKIS: GE125, GE126, GE234/
G15, GE236/G22, GE235, GE237, GE238/G21, GE242, GE243, GE245/G13, GE248, GE267/G14,
GE250, GE286); 1 ♀, 1 ? [2 SK], Yolokairi, 31 Aug. 1947, leg. unknown collector, between 30 Jul. 1947
and 30 Aug. 1947, leg. Jean Hiernaux (RBINS: 7056, 7063); 1 ♀ [UN], Zookis [= Kisangani],
24 Dec. 2015, leg. Prescott Musaba, Moïse Bipoo, Charle Andabhati, Benjamin Kirongozi and Kambale
Karupao (UNIKIS: ZCH06).
RWANDA: 5 ♀♀, 4 ♂♂ [8 AL, 1 UN], Gisenyi, 13 Oct. 1980, 15 Oct. 1980, leg. Hans H. de Iongh,
21 Oct. 1980, leg. Danny Meirte (ZMA: MAM.21243 to MAM.21247, MAM.21364, RMCA: 80.055M-0065 to 80.055-M-0067); 2 ♂♂, 1 ? [3 AL], Kibuye, 7 Oct. 1980, 9 Oct. 1980, leg. Danny Meirte
160
VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
(RMCA: 80.055-M-0059 to 80.055-M-0061); 4 sad ♀♀, 3 sad ♂♂, 2 juv ♀♀ [9 AL], Kidaho,
8 Jun. 1982, 10 Jun. 1982, 11 Jun. 1982, 12 Jun. 1982, 13 Jun. 1982, 17 Jun. 1982, 19 Jun. 1982, leg.
Benny Baeten (RMCA: 87.046-M-0002 to 87.046-M-0005, 87.046-M-0038, 87.046-M-0039, 87.046M-0061, 87.046-M-0062, 87.046-M-0001); 1 sad ♂ [AL], Kitabi, 2 Nov. 1981, leg. Frits De Vree,
Benny Baeten and Victor Van Cakenberghe (RMCA: 82.006-M-0033); 28 ad ♀♀, 19 ad ♂♂, 15 sad ♀♀,
18 sad ♂♂, 3 juv ♂♂, 2 ♀♀, 5 ♂♂, 5 ?? [11 SA, 82 AL, 2 UN], Musanze, 16 Jun. 1982, 17 Jun. 1982,
leg. Benny Baeten, 24 Oct. 1981, 25 Oct. 1981, 27 Oct. 1981, leg. Frits De Vree, Benny Baeten and
Victor Van Cakenberghe, 25 Oct. 1980, leg. Danny Meirte, 8 Jun. 1966, leg. Félix / Michel Anciaux
de Faveaux (RMCA: 87.046-M-0006 to 87.046-M-0037, 87.046-M-0040 to 87.046-M-0060, 82.006M-0003, 80.055-M-0053 to 80.055-M-0058, 80.055-M-0068 to 80.055-M-0072, 82.006-M-0004 to
82.006-M-0030, 82.006-M-0032, RBINS: 15485.1, 15485.2); 1 juv ♂ [AL], Nyamata, 16 Nov. 1981,
leg. Frits De Vree, Benny Baeten and Victor Van Cakenberghe (RMCA: 82.006-M-0034); 1 ♀, 3 ??
[4 AL], Nyundo, 20 Oct. 1980, leg. Danny Meirte (RMCA: 80.055-M-0052, 80.055-M-0062 to 80.055M-0064); 1 juv ♀, 1 imm ♂, 7 ♀♀, 5 ♂♂ [13 AL, 1 UN], Ruhengeri, 15 Oct. 1980, leg. Hans H. de
Iongh, 20 Sep. 1976, leg. Frieder Meier, 30 Nov. 1985, leg. unknown collector (ZMA: MAM.21248
to MAM.21255, MHNG: 1872.067 to 1872.069, SMF: 66863 to 66865); 1 sad ♂ [AL], Rukira,
22 Nov. 1981, leg. Frits De Vree, Benny Baeten and Victor Van Cakenberghe (RMCA: 82.006-M-0035);
1 sad ♀ [AL], Tamira, 26 Oct. 1981, leg. Frits De Vree, Benny Baeten and Victor Van Cakenberghe
(RMCA: 82.006-M-0031); 1 sad ♀ [AL], Uwinka, 17 Oct. 1981, leg. Frits De Vree, Benny Baeten and
Victor Van Cakenberghe (RMCA: 82.006-M-0002).
Scotonycterini Bergmans, 1997
Casinycteris Thomas, 1910
Casinycteris argynnis Thomas, 1910
DEMOCRATIC REPUBLIC OF THE CONGO: 1 ♂, 2 ?? [1 SS, 1 SA, 1 UN], unknown locality,
unknown date, between 1 Jun. 1911 and 30 Jun. 1911, leg. unknown collector, between 1 Jan. 1984 and
31 Dec. 1984, leg. Betu Nsankulu (RMNH: MAM.24558, RMCA: 84.035-M-0003, SMF: 6361); 1 ♂
[AL], Baliko, prior to 31 Jan. 2007, leg. Guy-Crispin Gembu Tungaluna (RMCA: a1.097-M-0412); 2 ♀♀
[2 SS], Bena Mbala, 15 Jan. 1984, leg. Betu Nsankulu (RMCA: 84.035-M-0001, 84.035-M-0002); 1 ♀,
1 ♂ [2 SA], Boende, between 1 Jan. 1967 and 31 Dec. 1967, leg. P. Lootens (RMCA: 38652, 38653);
1 ♀ [AL], Bomane [= Bomaneh], 31 May 2010, leg. Guy-Crispin Gembu Tungaluna, Prescott Musaba
and José Akaibe (UNIKIS: CRT2093); 1 ? [AL], Boteka, between 16 Feb. 1985 and 30 Jun. 1985, leg.
P. Lootens (RMCA: 85.030-M-0001); 2 ♀♀ [2 AL], Djabir, 8 May 2011, 10 May 2011, leg. Prescott
Musaba and Jonathan Kosele (UNIKIS: DJB096, DJB035); 1 ♀ [AL], Epulu, 29 Aug. 2015, leg.
André Malekani (UNIKIS: EPLU664); 3 ♀♀, 4 ♂♂, 1 ? [4 AL, 4 UN], Île Mbiye [= Mbiye Island],
25 Feb. 2012, 13 Apr. 2012, 21 Apr. 2014, leg. Prescott Musaba, 19 Feb. 2012, 24 Feb. 2012, 25 Feb. 2012,
leg. Prescott Musaba, Tembele and Marc Boketshu (RMCA: a1.097-M-3183, UNIKIS: CHIM11,
CHIM013/14, CHIM15, RMCA: a1.097-M-3180 to a1.097-M-3182, UNIKIS: CHIM28); 1 ad ♀, 1 ad
♂, 5 ♀♀, 9 ♂♂, 2 ?? [1 SS, 9 AL, 7 SK, 1 UN], Irangi, unknown date, 30 Dec. 1956, 11 Jan. 1984,
18 Jan. 1984, 5 Feb. 1984, 17 Feb. 1984, 18 Feb. 1984, leg. Heinz Stephan, 16 Oct. 1990, 19 Oct. 1990,
leg. Wim Bergmans (SMF: 65046, 64988 to 64991, ZMA: MAM.24193, SMF: 64994 to 64998, 65052,
64992, 64999, 65000, ZMA: MAM.24194, RMCA: 27430, SMF: 64993); 1 ♀ [SK], Kivu region,
12 Jan. 1984, leg. Heinz Stephan (SMF: 69400); 1 ♂ [UN], Kona, 15 May 2010, leg. Guy-Crispin
Gembu Tungaluna, Prescott Musaba and José Akaibe (UNIKIS: CRT1974); 2 ♀♀ [2 AL], Kumbi,
9 Dec. 1993, leg. Ngenge Masumbuko Kamitongo (ZMA: MAM.24894, MAM.24895); 1 ? [UN],
Kumi, unknown date, leg. unknown collector (RMNH: MAM.24894); 4 ♀♀, 1 ♂ [2 AL, 1 SK, 2 UN],
Lieki, 30 May 2010, 31 May 2010, 2 Jun. 2010, leg. Guy-Crispin Gembu Tungaluna, Prescott Musaba
and José Akaibe (UNIKIS: CRT2281, CRT2052, CRT2061, CRT2079, CRT2092); 1 ♀ [SS], Lukonga,
161
European Journal of Taxonomy 382: 1–327 (2017)
6 Aug. 1979, leg. Betu Nsankulu (RMCA: 80.013-M-0003); 1 ♀, 1 ♂, 2 ?? [2 SS, 2 SA], Luluabourg
[= Kananga], 25 Feb. 1965, 18 Mar. 1965, leg. Antoon Emeric Marcel De Roo (RMCA: 33412, 33347,
33348, 33411); 1 ♀, 1 ♂ [2 AL], Lwana River, 12 Apr. 1992, leg. Wim Bergmans, 16 Oct. 1990,
leg. Ngenge Masumbuko Kamitongo (ZMA: MAM.24558, MAM.24559); 4 ♀♀, 2 ♂♂ [5 SA, 1 SB],
Mandima, 7 Jul. 1976, 8 Jul. 1976, 15 Jul. 1976, 31 Jul. 1976, 1 Aug. 1976, leg. Frieder Meier (MHNG:
1871.096, 1871.094, 1871.095, 1871.091 to 1871.093); 5 ad ♂♂, 22 ♀♀, 28 ♂♂ [4 AL, 49 SK, 2 UN],
Masako, 13 Jun. 2015, leg. Prescott Musaba, Moïse Bipoo, Charle Andabhati, Benjamin Kirongozi and
Kambale Karupao, 30 Nov. 1987, 20 Jan. 1988, 21 Jan. 1988, 22 Jan. 1988, 23 Jan. 1988, 24 Jan. 1988,
25 Jan. 1988, 26 Jan. 1988, 27 Jan. 1988, 30 Jan. 1988, 21 Feb. 1988, 24 Jan. 1990, 16 Feb. 1990,
17 Feb. 1990, leg. Ndey B. Ifuta (UNIKIS: CHMS076, RMCA: 93.079-M-0324 to 93.079-M-0326,
93.079-M-0092, 93.079-M-0243, 93.079-M-0244, 93.079-M-0246, 93.079-M-0247, 93.079-M-0093
to 93.079-M-0095, 93.079-M-0248, 93.079-M-0119 to 93.079-M-0125, 93.079-M-0249 to 93.079-M0256, 93.079-M-0096 to 93.079-M-0101, 93.079-M-0103 to 93.079-M-0106, 93.079-M-0302, 93.079M-0323, 93.079-M-0107 to 93.079-M-0118, 93.079-M-0303, 93.079-M-0304, 93.079-M-0322); 2 ad
♀♀ [1 SA, 1 UN], Mbwambala, between 9 Jul. 1995 and 11 Jul. 1995, leg. Herwig Leirs et al. (MSB:
274606, RMCA: 97.021-M-0579); 1 ad ♀ [UN], Medje, 22 Apr. 1910, leg. Herbert Lang, James Paul
Chapin and The American Museum Congo Expedition (AMNH: 48751); 6 ♀♀, 4 ♂♂ [10 UN], Obenge,
5 Feb. 2013, 7 Feb. 2013, leg. Prescott Musaba and André Malekani (UNIKIS: TLL354, TLL356 to
TLL359, TLL363, TLL375 to TLL377, TLL385); 1 ♀ [AL], Rubi-Tele, 23 Aug. 2013, leg. Guy-Crispin
Gembu Tungaluna, Prescott Musaba and André Malekani (UNIKIS: RBTL547); 1 ♂ [SA], Stanleyville
[= Kisangani], between 1 May 1926 and 31 May 1926, leg. Henri Schouteden (RMCA: 16211); 6 ??
[6 AL], Wafanya [= Wafania], 12 Sep. 1991, 28 Apr. 1992, 4 May 1992, 8 May 1992, between 1 Jan. 1991
and 31 Dec. 1991, leg. P. Lootens (RMCA: 91.076-M-0079, 91.076-M-0080, 92.079-M-0073, 92.079M-0075, 92.079-M-0074, 92.079-M-0072); 1 ♀, 2 ♂♂ [3 UN], Yaekela, 3 May 2010, 5 May 2010,
6 May 2010, leg. Guy-Crispin Gembu Tungaluna, Prescott Musaba and José Akaibe (UNIKIS: CRT2364,
CRT1849, CRT1854); 2 ♂♂, 3 ?? [4 AL, 1 UN], Yoko, unknown date, 4 Sep. 2012, leg. Prescott
Musaba, 7 Sep. 2012, leg. Prescott Musaba, Tembele and Marc Boketshu (RMCA: a1.097-M-3184 to
a1.097-M-3187, UNIKIS: YK038CH); 1 ♀, 2 ♂♂ [3 AL], Zookis [= Kisangani], 16 Feb. 2013, leg.
Prescott Musaba and André Malekani (UNIKIS: CSB1, CSB3, CSB4).
Scotonycteris Matschie, 1894
Scotonycteris bergmansi
Hassanin, Khouider, Gembu, Goodman, Kadjo, Nesi, Pourrut, Nakouné & Bonillo, 2015
DEMOCRATIC REPUBLIC OF THE CONGO: 3 ♀♀ [1 SS, 2 SK], Baliko, 11 Jun. 2012,
12 Jun. 2012, leg. William (“Bill”) T. Stanley (FMNH: 219598, 219681, 219599); 1 ? [SS], Beni,
prior to 2 Jun. 1914, leg. Bonnevie (RMCA: 3244); 1 ♀ [SS], Bitale, 14 Jun. 1970, leg. Alexandre
Prigogine (RMCA: 36228); 2 ♂♂, 2 ?? [2 AL, 2 UN], Epulu, unknown date, leg. Luc Bijnens,
29 Aug. 2015, leg. André Malekani (RMCA: 90.031-M-0007, 90.031-M-0008, UNIKIS: EPLU676,
EPLU677); 2 ♀♀, 1 ♂ [1 SK, 2 SB], Epulu, 2 km W, 29 Jun. 1991, 3 Jul. 1991, 6 Jul. 1991, leg.
Julian C. Kerbis Peterhans (FMNH: 149402 to 149404); 2 ♀♀ [2 UN], Île Mbiye [= Mbiye Island],
25 Dec. 2015, 8 Jan. 2016, leg. Prescott Musaba, Moïse Bipoo, Charle Andabhati, Benjamin Kirongozi
and Kambale Karupao (UNIKIS: ABCHIM02, ABCHIM10); 2 ♀♀, 5 ♂♂ [3 AL, 3 SK, 1 UN],
Irangi, 11 Jan. 1984, 21 Jan. 1984, 28 Jan. 1984, 15 Feb. 1984, leg. Heinz Stephan, 14 Nov. 1955,
leg. J.J. Laarman, 18 Oct. 1990, leg. Wim Bergmans (SMF: 65006, RMNH: MAM.26325, SMF:
65009, 65010, ZMA: MAM.24197, SMF: 65007, 65008); 1 ♀ [SS], Kiliza [River], 25 May 1964, leg.
Alexandre Prigogine (RMCA: 32584); 1 ♂ [SS], Kiloboze [= Kilobono], 2 Feb. 1963, leg. Alexandre
Prigogine (RMCA: 31345); 2 ♂♂ [2 UN], Lieki, 29 May 2010, leg. Guy-Crispin Gembu Tungaluna,
Prescott Musaba and José Akaibe (UNIKIS: CRT2041, CRT2042); 1 ? [AL], Maganga, unknown
date, leg. Luc Bijnens (RMCA: 90.031-M-0009); 3 ♂♂ [1 SA, 2 AL], Mandima, 7 Jul. 1976,
162
VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
8 Jul. 1976, leg. Frieder Meier (MHNG: 1871.088 to 1871.090); 1 ad ♀, 3 ad ♂♂, 1 sad ♂, 2 juv
♂♂, 5 ♀♀, 2 ♂♂ [4 AL, 10 SK], Masako, 20 Jan. 1988, 23 Jan. 1988, 23 Mar. 1988, 28 Jun. 1988,
26 Aug. 1988, 30 Apr. 1989, 14 Sep. 1989, 16 Sep. 1989, 22 Dec. 1989, 13 Feb. 1990, 14 Feb. 1990,
leg. Ndey B. Ifuta (RMCA: 93.079-M-0364 to 93.079-M-0366, 93.079-M-0359 to 93.079-M-0361,
93.079-M-0245, 93.079-M-0363, 93.079-M-0102, 93.079-M-0170, 93.079-M-0357, 93.079-M-0358,
93.079-M-0356, 93.079-M-0362); 1 ♀ [SS], Mwenge, 5 Feb. 1967, leg. Alexandre Prigogine (RMCA:
35076); 1 ♀, 1 ♂ [1 SS, 1 SK], Quatorze, 23 Jun. 2013, 29 Jun. 2013, leg. William (“Bill”) T. Stanley
(FMNH: 222426, 222623); 9 ♂♂ [9 UN], Rubi-Tele, 17 Aug. 2013, 18 Aug. 2013, 19 Aug. 2013,
20 Aug. 2013, 22 Aug. 2013, leg. Guy-Crispin Gembu Tungaluna, Prescott Musaba and André
Malekani (UNIKIS: RBTL481, RBTL482, RBTL492, RBTL494, RBTL504, RBTL515, RBTL505,
RBTL534, RBTL540); 1 ad ♀ [AL], Sukisa, unknown date, leg. unknown collector (MNHN: ZM2014-564 [holotype Scotonycteris bergmansi congoensis Hassanin, Khouider, Gembu, Goodman,
Kadjo, Nesi, Pourrut, Nakouné, and Bonillo, 2015]); 3 ♀♀, 1 ♂ [4 UN], Uma, 10 Apr. 2014,
11 Apr. 2014, 12 Apr. 2014, leg. Guy-Crispin Gembu Tungaluna, Prescott Musaba and André
Malekani (UNIKIS: UMA525, UMA583, UMA609, UMA615); 2 ♀♀, 1 ? [3 AL], Yoko, 4 Sep. 2012,
5 Sep. 2012, leg. Prescott Musaba (RMCA: a1.097-M-3188 to a1.097-M-3190).
Stenonycterini Nesi, Kadjo, Pourrut, Shongo, Cruaud & Hassanin, 2012
Stenonycteris Gray, 1870
Stenonycteris lanosus (Thomas, 1906)
BURUNDI: 1 ♀, 3 ♂♂ [2 SK, 2 SB], Giserama, Colline, 30 Jul. 1991, 31 Jul. 1991, 2 Aug. 1991, leg.
Julian C. Kerbis Peterhans (FMNH: 148979, 148976 to 148978); 2 ♀♀, 1 ♂ [3 SB], Ruhinga Hill,
9 Mar. 1993, 10 Mar. 1993, 13 Mar. 1993, leg. Jay L. Udelhoven (FMNH: 155970, 155971, 155969).
DEMOCRATIC REPUBLIC OF THE CONGO: 1 juv ♀, 2 ♀♀ [2 SS, 1 AL], Butagu Valley [= Butahu
Valley Cave], 24 Dec. 1926, leg. James Paul Chapin, Dewitt L. Sage, Frank P. Mathews and The
Ruwenzori-Kivu Expedition, 24 Dec. 1926, leg. James Paul Chapin (AMNH: 82380, 82527, 82528);
1 ♀, 4 ♂♂, 1 ? [3 SS, 3 SA], Butembo, 1 Jan. 1970, leg. M.J. Celis and M. Lejeune, 31 Jan. 1956,
21 Feb. 1956, leg. P. Dyleff (RMCA: 36200, 36202, 36205, 23674, 23676, 23553); 1 ♂ [SS], Kabira,
19 Mar. 1959, leg. Uwe Rahm (RMCA: 28025); 3 ♀♀, 2 ♂♂, 2 ?? [5 SS, 2 UN], Kalonge, unknown
date, 23 Jan. 1953, 28 Jan. 1953, 22 Feb. 1953, leg. Gaston-François de Witte, 21 Jan. 1925, leg. Edmund
Heller, 22 Dec. 1926, leg. James Paul Chapin (RBINS: 37116, 37117, FMNH: 26237, RMCA: 37117,
AMNH: 82526, RBINS: 17299, RMCA: 37116); 1 ? [UN], Kikura, unknown date, leg. unknown
collector (INPBC: V1178); 1 imm ♀ [SS], Lamya Valley, 16 Nov. 1945, leg. Gaston-François de Witte
(RBINS: 12928); 1 ♀ [AL], Lemera, SW of, 16 Jul. 1927, leg. James Paul Chapin (AMNH: 82379); 1 ♂
[SS], Lushala, 4 Nov. 1966, leg. A. Elbl (USNM: 535400); 2 ♀♀ [2 SA], Lwiro [= Luiro], 8 Sep. 1963,
23 Oct. 1965, leg. P. Kunkel (SMF: 31829, 31830); 1 ? [UN], Marion, Lake, unknown date, leg. unknown
collector (INPBC: V1159); 1 ♂ [SK], Marumbi, 6 Dec. 1954, leg. Gaston-François de Witte (RMCA:
37119); 1 ♂ [SO], Mount Hoyo [= Hoyo, Mount], 12 Aug. 1947, leg. Jean Hiernaux (RMCA: 35806);
3 ♂♂ [2 AL, 1 SK], Mount Tshiabirimu [= Tshiabirimu, Mount], 5 Jun. 2004, leg. Prince Kiswele
Kaleme (FMNH: 188790 to 188792); 2 ♀♀ [1 SS, 1 SK], Mugaba Cave, 19 Jul. 1962, leg. A. Elbl
(USNM: 375232, 375233); 1 ♀ [SO], Mutsora, 13 Aug. 1949, leg. Gaston-François de Witte (RBINS:
12927); 1 juv ♂, 2 ♀♀, 1 ♂ [2 AL, 2 UN], Nyabiondo, 27 Dec. 1990, 28 Dec. 1990, 1 Jan. 1991,
leg. J.P. Lubula Bulambo and Ngenge Masumbuko Kamitongo (ZMA: MAM.24346, MAM.24347,
MAM.24344, MAM.24345); 8 ♀♀, 1 ♂ [3 SS, 6 AL], Old Kalonge [= Ugongo], 1 Feb. 1925,
2 Feb. 1925, 3 Feb. 1925, 4 Feb. 1925, 5 Feb. 1925, 8 Feb. 1925, leg. Edmund Heller (FMNH: 26234 to
26236, 30572 to 30574, USNM: 260141, FMNH: 30576, 30577); 1 ♂ [SS], Tshibasha, 18 Nov. 1970,
leg. Betu Nsankulu (RMCA: 36250); 1 ♀ [SS], Tshibati, 28 Dec. 1963, leg. Antoon Emeric Marcel
De Roo (AMNH: 207735).
163
European Journal of Taxonomy 382: 1–327 (2017)
RWANDA: 1 ♂ [SS], Bugundi, 19 Nov. 1955, leg. Alexandre Fain (RMCA: 81.012-M-0043); 1 ♀,
1 ♂ [2 SS], Gasiza, 23 Feb. 1978, leg. unknown collector (RBINS: 20215, 20273); 1 ad ♂ [SA],
Kanyamiketo, 4 Jun. 1982, leg. Benny Baeten (RMCA: 87.046-M-0518); 3 ad ♂♂, 1 sad ♀ [4 SA],
Kayove, 10 Jul. 1982, 11 Jul. 1982, leg. Benny Baeten (RMCA: 87.046-M-0515, 87.046-M-0516,
87.046-M-0519, 87.046-M-0520); 4 ?? [4 SK], Kibumbu Crater, 9 Dec. 1991, leg. J.R. Wilson and
Anthony Michael Hutson (BMNH: 1992.385 to 1992.388); 1 ad ♂ [SA], Kinigi, 1 Jun. 1982, leg. Benny
Baeten (RMCA: 87.046-M-0514); 5 ♀♀, 2 ♂♂, 16 ?? [23 SA], Murunda, 12 Oct. 1980, leg. Danny
Meirte (RMCA: 80.055-M-0073 to 80.055-M-0095); 1 ad ♂ [SA], Routabansougera, 17 Jul. 1982, leg.
Benny Baeten (RMCA: 87.046-M-0517); 1 ? [UN], Ruhengeri, 22 Jun. 2003, leg. unknown collector
(SMF: 92441); 2 ad ♀♀, 1 ad ♂ [3 SA], Uwinka, 17 Oct. 1981, leg. Frits De Vree, Benny Baeten and
Victor Van Cakenberghe (RMCA: 82.006-M-0932 to 82.006-M-0934).
Hipposideridae Lydekker, 1891
DEMOCRATIC REPUBLIC OF THE CONGO: 1 ? [SK], Aka, between 15 May 1952 and 28 Dec. 1972,
leg. H. de Saeger (RMCA: 36837); 1 ♀, 1 ♂, 7 ?? [9 AL], Boende, between 1 Jan. 1967 and 31 Dec. 1967,
leg. P. Lootens (RMCA: 38706 to 38714); 1 ? [SK], Keroma, between 21 Apr. 1952 and 28 Dec. 1972,
leg. H. de Saeger (RMCA: 36836); 1 ♂ [SS], Kilibozi, 12 Apr. 1975, leg. Alexandre Prigogine (RMCA:
76.022-M-0023); 2 ?? [2 SS], Kiligi, 11 Feb. 1975, leg. Alexandre Prigogine (RMCA: 75.017-M-0004,
75.017-M-0005); 1 ? [SS], Kinda, 17 Aug. 1978, leg. Betu Nsankulu (RMCA: 79.022-M-0008); 1 ♀
[SK], Luluabourg [= Kananga], unknown date, leg. Richard Callewaert (RMCA: 7543 b); 1 ♂, 24 ??
[25 AL], Masako, unknown date, 19 Dec. 1989, leg. Ndey B. Ifuta (RMCA: 93.079-M-0371 to 93.079M-0394, 93.079-M-0329); 3 ?? [2 SK, 1 SN], PPK 72, between 18 Mar. 1952 and 28 Dec. 1972, leg.
H. de Saeger (RMCA: 36833 to 36835).
Asellia Gray, 1838
Asellia tridens (E. Geoffroy St.-Hilaire, 1813)
DEMOCRATIC REPUBLIC OF THE CONGO: 2 ♀♀, 15 ♂♂ [17 AL], Kivu Region, unknown date,
leg. Guy Babault (MNHN: ZM-MO-1985-846 to ZM-MO-1985-854, ZM-MO-1985-856 to ZMMO-1985-863); 1 ♂ [AL], Kivu, near, unknown date, leg. Guy Babault (MNHN: ZM-MO-1985-855).
Doryrhina Peters, 1871
Doryrhina cyclops (Temminck, 1853)
BURUNDI: 1 ♀, 2 ♂♂ [1 AL, 2 SB], Giserama, Colline, 3 Aug. 1991, 11 Aug. 1991, 23 Aug. 1991,
leg. Julian C. Kerbis Peterhans (FMNH: 148982, 148983, 148981).
DEMOCRATIC REPUBLIC OF THE CONGO: 1 ♂ [SS], Aba, 17 Apr. 1945, leg. unknown collector
(RBINS: 13332); 9 ?? [9 UN], Abatupi River, unknown date, leg. unknown collector (INPBC: V1285.5,
V1290.1 to V1290.8); 1 ♂ [SK], Abimva, 22 Jun. 1925, leg. Henri Schouteden (RMCA: 9028b); 1 ?
[UN], Albert National Park [= Virunga National Park], unknown date, leg. unknown collector
(INPBC: V1852.2); 1 ad ♂, 4 ♀♀, 1 ? [4 SS, 1 SA, 1 SK], Avakubi, 1 Oct. 1913, 24 Jan. 1914, leg.
Herbert Lang, James Paul Chapin and The American Museum Congo Expedition, 26 Jan. 1914, leg.
Herbert Lang, between 24 Jan. 1914 and 24 Dec. 1914, leg. unknown collector, between 7 Nov. 1912 and
30 Apr. 1913, leg. Cuthbert Christy (AMNH: 49404, 49098 [holotype Hipposideros langi J.A. Allen,
1917], 49099, FMNH: 43878, MCZ: 17370, RMCA: 1606b); 1 ♂ [SK], Baliko, 12 Jun. 2012, leg.
William (“Bill”) T. Stanley (FMNH: 219601); 1 ? [SS], Beni, prior to 2 Jun. 1914, leg. Bonnevie (RMCA:
3243); 5 ♀♀, 1 ♂ [6 AL], Bikoro, 25 Jun. 1956, leg. Alvin Novick (MCZ: 48219 to 48224); 1 ♀ [UN],
Bomane [= Bomaneh], 30 May 2010, leg. Guy-Crispin Gembu Tungaluna, Prescott Musaba and José
164
VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
Akaibe (UNIKIS: CRT2067); 1 ♀ [AL], Bozade, 20 Jul. 1979, leg. Frieder Meier (MHNG: 1870.045);
2 ♂♂ [2 UN], Buluku River, 22 May 1952, leg. H. de Saeger et al. (RBINS: 13678.1, 13678.2); 1 ♀,
1 ♂ [2 SO], Buta, 1 Feb. 1950, 6 Dec. 1950, leg. unknown collector (RBINS: 13335, 13334); 4 ??
[4 UN], Bwirina, unknown date, leg. unknown collector (INPBC: V1661.4, V1662.1 to V1662.3); 1 ?
[SA], Congo River, 75 mi [= 121 km] up, between 1 Aug. 1897 and 31 Aug. 1897, leg. unknown
collector (USNM: 102512); 5 ♀♀, 2 ♂♂, 3 ?? [3 AL, 7 UN], Dedegwa River, 19 May 1952, leg. H. de
Saeger et al., 19 May 1952, leg. H. de Saeger (RBINS: 13676, 13677.1 to 13677.6, RMCA: 36820 to
36822); 1 ? [UN], Djelube River, lower, unknown date, leg. unknown collector (INPBC: V1093.1);
2 ?? [2 UN], Djelube River, middle, unknown date, leg. unknown collector (INPBC: V1078.3, V1100);
1 ♀, 1 ? [1 SS, 1 AL], Epulu, unknown date, leg. Luc Bijnens, 2 May 1951, leg. A. Schwartz and
R.N. Porter (RMCA: 90.031-M-0004, KU: 151340); 2 ♀♀, 1 ♂ [3 UN], Fataki, between 1 Jan. 1948
and 6 Apr. 1964, leg. unknown collector (RBINS: 18529.1 to 18529.3); 2 ♀♀, 5 ?? [2 AL, 1 SO, 4 UN],
Inimvua, 15 May 1952, leg. H. de Saeger et al., 15 May 1952, leg. H. de Saeger (RBINS: 13505, 13674.1
to 13674.4, RMCA: 36818, 36819); 1 ♀, 5 ♂♂ [6 SS], Irumu, 40 mi [= 64 km] SW [= Ituri Forest],
unknown date, leg. Howard de Walden (BMNH: 1930.11.11.146 to 1930.11.11.150, 1930.11.11.45);
1 ? [UN], Kambemba, unknown date, leg. unknown collector (INPBC: V1657); 1 ♀ [SO], Liki,
6 Mar. 1952, leg. unknown collector (RBINS: 13333); 1 ? [SS], Lovanium University [= Kinshasa],
30 Mar. 1962, leg. De Bont (RMCA: 31201); 1 ♀, 1 ♂, 2 ?? [1 SS, 3 AL], Lukolela, 28 Jul. 1930,
17 Mar. 1931, leg. James Paul Chapin (AMNH: 86858, 86863, 86866, 86909); 1 ? [UN], Makayova
River, unknown date, leg. unknown collector (INPBC: V1257.2); 1 ? [UN], Malulu, unknown date,
leg. unknown collector (INPBC: V185); 1 ? [UN], Mamudioma, unknown date, leg. unknown collector
(INPBC: W1152); 2 ♀♀ [2 AL], Masako, 15 Sep. 1988, 11 Aug. 1989, leg. Ndey B. Ifuta (RMCA:
93.079-M-0328, 93.079-M-0327); 1 juv ♂, 4 ♀♀ [1 SS, 4 UN], Medje, 22 Apr. 1910, 27 Apr. 1910,
3 Sep. 1910, leg. Herbert Lang, James Paul Chapin and The American Museum Congo Expedition
(RMCA: 12379, AMNH: 49093 to 49096); 2 ad ♀♀, 1 juv ?, 1 foetus ?, 7 ♀♀, 4 ?? [1 SS, 7 AL, 1 SO,
6 UN], Mogbwamu River, 8 Mar. 1952, leg. H. de Saeger et al., 8 Mar. 1952, between 8 Mar. 1952
and 28 Dec. 1972, leg. H. de Saeger (RBINS: 13502, 13503, 13669.1 to 13669.4, RMCA: 36808 to
36812, 36813a, 36813b, 36815, 36814); 1 ♀, 2 ♂♂ [1 SS, 2 AL], Niangara, 3 Dec. 1910, 6 Dec. 1910,
leg. Herbert Lang, James Paul Chapin and The American Museum Congo Expedition, 6 Dec. 1910,
leg. Herbert Lang (AMNH: 49097, 49415, FMNH: 43879); 1 ad ♂ [SA], Niapu, 28 Jan. 1914, leg.
Herbert Lang, James Paul Chapin and The American Museum Congo Expedition (AMNH: 49391);
3 ♀♀, 1 ♂ [4 AL], Obenge, 5 Feb. 2013, 6 Feb. 2013, leg. Prescott Musaba and André Malekani
(UNIKIS: TLL361, TLL364 to TLL366); 1 ♀ [SS], Parc National de Garamba [= Garamba National
Park], 7 Feb. 1952, leg. H. de Saeger et al. (RBINS: 13675); 2 ♀♀ [2 UN], PFNK.16, 7 Feb. 1952,
leg. H. de Saeger et al. (RBINS: 13668.1, 13668.2); 2 ♂♂ [2 UN], Pidigala River, 23 Apr. 1952, leg.
H. de Saeger et al. (RBINS: 13672, 13673); 2 ?? [2 UN], Ruanoli-Mavea conluence, unknown date,
leg. unknown collector (INPBC: V1668.1, V1668.3.3); 2 ?? [2 UN], Rwere Rwindi, unknown date, leg.
unknown collector (INPBC: V1325.1, V1325.2); 2 ♂♂ [2 UN], Taere River, 10 Mar. 1952, leg. H. de
Saeger et al. (RBINS: 13670, 13671); 1 ♂ [SS], Tandala, 20 Jul. 1979, leg. C. Brian Robbins (USNM:
537712); 2 ♀♀ [2 UN], Yaekela, 3 May 2010, leg. Guy-Crispin Gembu Tungaluna, Prescott Musaba
and José Akaibe (UNIKIS: CRT2367, CRT2368); 1 ♀ [SS], Yalosemba, 19 Jun. 1979, leg. C. Brian
Robbins (USNM: 537711).
Hipposideros Gray, 1831
DEMOCRATIC REPUBLIC OF THE CONGO: 1 juv ? [UN], Abatupi River, unknown date,
leg. unknown collector (INPBC: V1057.2b); 1 ♀ [AL], Île Mai [= Mai Island], 8 Jan. 2016, leg.
Prescott Musaba, Moïse Bipoo, Charle Andabhati, Benjamin Kirongozi and Kambale Karupao
(UNIKIS: ABCHIMAF12); 3 ♂♂ [3 AL], Irangi, slightly E of [= Irangi], 15 Oct. 1990, leg. Wim
Bergmans (ZMA: MAM.24182 to MAM.24184); 2 ♀♀ [2 AL], Luhohov (River), 18 Oct. 1990, leg.
165
European Journal of Taxonomy 382: 1–327 (2017)
Wim Bergmans (ZMA: MAM.24187, MAM.24188); 1 ♀ [AL], Musenge, 3 May 1992, leg. Ngenge
Masumbuko Kamitongo (ZMA: MAM.24553); 1 ♀, 1 ♂ [2 UN], Obenge, 7 Feb. 2013, 8 Feb. 2013, leg.
Prescott Musaba and André Malekani (UNIKIS: TLL390, TLL393); 3 ♂♂ [3 AL], Uma, 12 Apr. 2014, leg.
Guy-Crispin Gembu Tungaluna, Prescott Musaba and André Malekani (UNIKIS: UMA504, UMA618,
UMA619); 2 ♂♂ [2 UN], Yoko, 12 Feb. 2009, 4 Oct. 2009, leg. unknown collector (UNIKIS: GE128,
GE318).
Hipposideros abae J.A. Allen, 1917
DEMOCRATIC REPUBLIC OF THE CONGO: 1 ad ♂, 20 ♀♀, 17 ♂♂ [37 SS, 1 UN], Aba, 3 Dec. 1911,
13 Dec. 1911, leg. Herbert Lang, James Paul Chapin and The American Museum Congo Expedition,
13 Dec. 1911, leg. unknown collector (AMNH: 49120 to 49122, 49123 [holotype Hipposideros abæ
J.A. Allen, 1917], 49124, 49125, 49128 to 49130, 49146 to 49167, 49169, 49170, MCZ: 17371, 17372,
RMCA: 12366, TM: 4564, RMCA: 12367); 2 ?? [1 SO, 1 UN], Aka, 15 May 1952, leg. H. de Saeger
et al., 15 May 1952, leg. H. de Saeger (RBINS: 13684.1, RMCA: 36694); 1 ♂, 2 ?? [3 UN], Cel II,
21 May 1951, 25 Jun. 1951, 15 May 1952, leg. H. de Saeger et al. (RBINS: 13684.2, 13679, 13681);
1 ♂ [AL], Digba, between 1 Jul. 1981 and 31 Jul. 1981, leg. Marc Colyn (RMCA: 82.031-M-0005);
20 juv ♀♀, 24 juv ♂♂, 1 imm ♀, 21 ♀♀, 163 ?? [1 SS, 80 AL, 148 UN], Keroma, 21 Apr. 1952, leg.
H. de Saeger et al., 21 Apr. 1952, between 21 Apr. 1952 and 28 Dec. 1972, leg. H. de Saeger (RBINS:
13683, 13790.01 to 13790.25, 13791.01 to 13791.35, 13792.01 to 13792.30, 13793.01 to 13793.30,
13794.01 to 13794.27, RMCA: 36610, 36611, 36612a, 36613 to 36689, 36693); 1 ♀, 1 ♂ [2 UN],
Maleli, 13 Jun. 1951, leg. H. de Saeger et al. (RBINS: 13680.1, 13680.2); 1 ♀ [SS], Oriental (former)
Province, 13 Dec. 1911, leg. unknown collector (SMF: 37915); 4 ♀♀, 4 ♂♂ [8 UN], Pidigala River,
23 Apr. 1952, leg. H. de Saeger et al. (RBINS: 13682.1 to 13682.8).
Hipposideros beatus (K. Andersen, 1906)
DEMOCRATIC REPUBLIC OF THE CONGO: 1 juv ?, 4 ♀♀, 3 ♂♂ [2 SA, 6 UN], unknown locality,
unknown date, between 11 Aug. 1989 and 14 Aug. 1989, leg. unknown collector (BMNH: 1990.225
to 1990.230, SMF: 77085, 77086); 3 ♀♀, 3 ♂♂ [1 AL, 5 UN], Aka, 23 Apr. 1952, 14 May 1952, leg.
H. de Saeger et al., 14 May 1952, leg. H. de Saeger (RBINS: 13770, 4041, RMCA: 36690, RBINS:
13771, 4037 [holotype Hipposideros beatus maximus Verschuren, 1957], 4039); 1 ♂ [AL], Aketi,
21 Jun. 2015, leg. Vlir/Aketi (UNIKIS: AKETI603); 1 ♂ [UN], Bomane [= Bomaneh], 21 May 2010,
leg. Guy-Crispin Gembu Tungaluna, Prescott Musaba and José Akaibe (UNIKIS: CRT2197); 2 ??
[2 AL], Buluku, 22 May 1952, leg. H. de Saeger (RMCA: 36691, 36692); 2 ?? [2 UN], Buluku River,
22 May 1952, leg. H. de Saeger et al. (RBINS: 13774, 4044); 1 ? [AL], Buta, between 1 Jan. 1925
and 31 Dec. 1925, leg. Jozef Hutsebaut (RMCA: 16227); 2 ♂♂ [2 SA], Bwandza, 30 Jun. 1979, leg.
Frieder Meier (MHNG: 1870.095, 1870.096); 2 ♂♂ [2 UN], Dedegwa River, 19 May 1952, leg. H. de
Saeger et al. (RBINS: 13772, 4042); 1 ♀ [SA], Equateur (former) Province, prior to 7 Oct. 1996, leg.
unknown collector (SMF: 84412); 2 ♂♂, 1 ? [3 SS], Faradje, 6 Jul. 1925, 7 Jul. 1925, 11 Jul. 1925, leg.
Henri Schouteden (RMCA: 9056, 9044, 9048); 1 ? [UN], Ikengo, unknown date, leg. unknown collector
(RMCA: 5646); 2 ♀♀, 2 ♂♂ [4 UN], Kona, 12 May 2010, 15 May 2010, leg. Guy-Crispin Gembu
Tungaluna, Prescott Musaba and José Akaibe (UNIKIS: CRT1909, CRT1973, CRT1975, CRT1983);
2 ♂♂ [2 UN], Lieki, 29 May 2010, leg. Guy-Crispin Gembu Tungaluna, Prescott Musaba and José
Akaibe (UNIKIS: CRT2046, CRT2047); 2 ♂♂ [2 AL], Mabali, 18 Jun. 1956, leg. Alvin Novick (MCZ:
48217, 48218); 1 ♀, 1 ♂ [2 AL], Mandima, 21 Jul. 1976, leg. Frieder Meier (MHNG: 1873.057,
1873.058); 1 ♀ [SS], Medje, 3 Aug. 1910, leg. unknown collector (MCZ: 17374); 1 ♀, 1 ? [2 AL],
Moenge, 15 May 2010, 16 May 2010, leg. Guy-Crispin Gembu Tungaluna (RMCA: a1.097-M-3218,
a1.097-M-3217); 2 ♂♂ [2 UN], Mogbwamu River, 8 Mar. 1952, leg. H. de Saeger et al. (RBINS:
13767, 4038); 2 ♂♂ [2 UN], Nakpanga, 18 Apr. 1952, leg. H. de Saeger et al. (RBINS: 13769, 4040);
1 ? [AL], Panga, 1 Dec. 1914, leg. J. Bequaert (RMCA: 5446); 1 ♂ [UN], Pidigala River, 23 Apr. 1952,
166
VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
leg. H. de Saeger et al. (RBINS: 13768; 2 ♂♂ [2 UN], Soudan River, 20 May 1952, leg. H. de Saeger
et al. (RBINS: 13773, 4043); 1 ? [UN], Thysville [= Mbanza-Ngungu], 11 Feb. 1987, leg. unknown
collector (RBINS: 22426); 1 ♂ [UN], Yaekela, 6 May 2010, leg. Guy-Crispin Gembu Tungaluna,
Prescott Musaba and José Akaibe (UNIKIS: CRT1856); 1 ♀, 3 ♂♂ [2 SS, 1 SA, 1 AL], Yalosemba,
16 Jun. 1979, leg. Lynn W. Robbins, 16 Jun. 1979, leg. C. Brian Robbins, 18 Jun. 1979, 30 Jun. 1979,
leg. Frieder Meier (USNM: 537707, 537708, MHNG: 1870.093, 1870.094).
Hipposideros caffer (Sundevall, 1846)
BURUNDI: 1 ad ♀, 1 juv ?, 2 ♀♀, 1 ♂ [3 SA, 2 AL], unknown locality, 3 Mar. 1965, leg. P.L. Niort,
3 Mar. 1979, leg. Heinz Stephan (MHNG: 1896.013, 1896.014, SMF: 57956 to 57958); 1 ? [AL],
Bujumbura, between 1 Jul. 1968 and 31 Jul. 1968, leg. Lewall (RMCA: 35154); 1 ♂ [AL], Mabaye
[= Mabayi], 12 Jul. 1973, leg. P.L. Niort (MHNG: 1896.012); 4 ?? [3 AL, 1 UN], Rubindi, between
1 Sep. 1958 and 30 Sep. 1958, leg. Narcisse Leleup (RMCA: 28455 to 28458).
DEMOCRATIC REPUBLIC OF THE CONGO: 1 juv ♂, 18 ?? [1 SA, 2 AL, 9 SK, 1 SO, 6 UN],
unknown locality, unknown date, 12 Feb. 2001, 28 Feb. 2001, prior to 31 Jan. 1897, between 1 Jun. 1911
and 30 Jun. 1911, between 1 Apr. 1999 and 30 Apr. 1999, leg. unknown collector, unknown date, leg.
Alexandre Prigogine, prior to 1 Aug. 1957, leg. Heinz Stephan, prior to 18 Aug. 1965, leg. Antoon
Emeric Marcel De Roo (RBINS: 208C, RMCA: 32866, SMF: 92971, 92970, 93318 to 93320, 6363,
RMCA: 27451 to 27459, 33599, RBINS: 208e); 3 ♂♂ [3 AL], Aketi, 25 Jun. 2015, 27 Jun. 2015, leg.
Vlir/Aketi (UNIKIS: AKETI618, AKETI621, AKETI626); 21 ?? [21 UN], Albert National Park
[= Virunga National Park], unknown date, leg. unknown collector (INPBC: V1711, V1723.01 to
V1723.20); 1 ♀ [SS], Alima, between 5 Mar. 1959 and 5 Oct. 1959, leg. P. Dyleff (RMCA: 28154);
1 juv ♂ [SS], Ango, between 14 Apr. 1941 and 23 Jul. 1946, leg. J.M. Vrijdagh (RMCA: 17756); 3 ??
[3 AL], Ango Cave, 13 Apr. 1941, leg. J.M. Vrijdagh (RMCA: 18103, 18105, 18106); 1 ♀ [SS],
Avakubi, between 18 Sep. 1912 and 24 Feb. 1913, leg. Cuthbert Christy (RMCA: 1429); 2 ?? [2 AL],
Banana, between 1 Aug. 1952 and 31 Aug. 1952, leg. Is. Mesmaekers (RMCA: 21478, 21479); 3 ??
[3 AL], Baya Carrière [= Baya Quarry], 17 May 1965, leg. Walter N. Verheyen (RMCA: 35023,
35024, 35026); 1 ? [SS], Baye via Bondo, 13 Dec. 1956, leg. Rooyakkers (RMCA: 25863); 1 ♂ [SS],
Bela, 1 Aug. 1947, leg. unknown collector (RBINS: 7066); 14 ♀♀, 3 ♂♂ [3 SS, 1 SA, 13 AL], Beni,
8 Mar. 1925, 9 Mar. 1925, 11 Mar. 1925, 18 Mar. 1925, 22 Mar. 1925, 24 Mar. 1925, 28 Mar. 1925, leg.
Edmund Heller (FMNH: 30625, 26242, 26243, 30626 to 30634, 30622 to 30624, MCZ: 33500, FMNH:
26240); 21 ?? [21 AL], Boende, between 1 Nov. 1969 and 31 Dec. 1974, leg. P. Lootens (RMCA:
75.035-M-0036 to 75.035-M-0056); 2 ♀♀ [2 SS], Bolobo, prior to 27 Sep. 1951, leg. Ngwe (RMCA:
20535, 20536); 3 ?? [3 SO], Bombo Lumene, 20 Aug. 1992, leg. Tshinyenye (RMCA: 92.119-M-0001
to 92.119-M-0003); 7 ?? [7 AL], Bunia, between 1 Jan. 1951 and 31 Dec. 1951, leg. Alexandre Fain,
prior to 27 Dec. 1938, leg. Putnam (RMCA: 21468, 14950 to 14955); 1 ? [UN], Busholinka River,
unknown date, leg. unknown collector (INPBC: W1006); 2 ?? [2 AL], Buta, unknown date, leg. Jozef
Hutsebaut (RMCA: 14582a, 14582b); 2 ♀♀, 2 ♂♂, 9 ?? [3 SS, 6 AL, 4 SO], Butembo, 10 Aug. 1947,
14 Aug. 1947, leg. unknown collector, 4 Feb. 1950, leg. P. Dyleff, between 1 Nov. 1971 and 30 Nov. 1971,
leg. M. Lejeune, prior to 5 Jul. 1950, leg. Alexandre Prigogine (RBINS: 7078 to 7080, RMCA: 20574,
81.049-M-0003 to 81.049-M-0005, 19916 to 19921); 1 ♀, 1 ♂, 2 ?? [2 SA, 2 AL], Congo River, 75 mi
[= 121 km] up, unknown date, leg. George Latimer Bates, between 1 Aug. 1897 and 31 Aug. 1897, leg.
unknown collector (USNM: 102513, 102515, 102514, 102516); 1 ? [SS], Defrenne Cave [= Mwanga
Cave], between 19 Sep. 1948 and 22 Jun. 1949, leg. Narcisse Leleup (RMCA: 22490); 1 ♀, 4 ♂♂
[5 AL], Dethioux Farm, Cave I [= Shinkolobwe], 15 Jun. 1958, leg. Félix / Michel Anciaux de Faveaux
(RMCA: 27909 to 27913); 12 ?? [12 AL], Dikuluwe Cave [= Dikulwe Cave], 2 Jan. 1954, leg. R.P. Th.
de Caters (RMCA: 22090 to 22101); 1 ? [UN], Djelube River, lower, unknown date, leg. unknown
collector (INPBC: V1093.2); 1 ? [UN], Djuma River, upper, unknown date, leg. unknown collector
167
European Journal of Taxonomy 382: 1–327 (2017)
(INPBC: V1278); 16 ?? [16 UN], Dungu, 1 May 1948, leg. unknown collector (RBINS: 8128.01 to
8128.16); 2 ♀♀ [2 UN], Edouard, Lac [= Edward, Lake], 26 Jan. 1954, leg. unknown collector
(RBINS: 15450, 15451); 2 ?? [1 AL, 1 UN], Elisabethville [= Lubumbashi], between 1 Apr. 1926 and
30 Apr. 1926, leg. Henri Schouteden (RMCA: 16183, 16243); 1 ♂, 1 ? [2 AL], Epulu, 26 Aug. 2015,
leg. André Malekani, between 1 Oct. 1956 and 31 Oct. 1956, leg. Max Poll (UNIKIS: EPLU651, RMCA:
25653); 1 ♂ [UN], Epulu, 2 km W, 14 Jul. 1991, leg. unknown collector (FMNH: 149406); 1 ? [SO],
Etembo, 31 Mar. 1950, leg. P. Dyleff (RMCA: 19903); 2 ad ♀♀, 2 ♀♀, 2 ?? [1 SA, 2 AL, 3 UN],
Faradje, 1 Apr. 1952, leg. H. de Saeger et al., 1 Apr. 1952, leg. H. de Saeger, 24 Oct. 1912, leg. Herbert
Lang, James Paul Chapin and The American Museum Congo Expedition (RBINS: 13796.1 to 13796.3,
RMCA: 36823, 36824, AMNH: 49426 [holotype Hipposideros nanus J.A. Allen, 1917]); 1 ? [AL],
Funda Biabo, 26 Mar. 1914, leg. Charliers (RMCA: 5382); 2 ?? [2 AL], Garamba National Park,
prior to 28 Dec. 1972, leg. H. de Saeger (RMCA: 36826, 36827); 1 ? [AL], Gemena, 13 Feb. 1939, leg.
Leontovitch (RMCA: 15841); 7 ♀♀, 6 ♂♂ [13 UN], Goroumbwa Mine, 17 May 1999, 18 May 1999,
19 May 1999, 23 Oct. 1999, 24 Oct. 1999, 25 Oct. 1999, 27 Oct. 1999, leg. Robert (Bob) Swanepoel
(TM: DRC0015, DRC0034, DRC0069, DRC0105, DRC0106, DRC0121, DBA207, DBA261, DBA266,
DBA268, DBA274, DBA320, DBA410); 4 ♀♀, 7 ♂♂, 1 ? [10 SS, 1 AL, 1 UN], Hoyo, Mount, unknown
date, 11 Aug. 1947, 14 Aug. 1947, 16 Aug. 1947, leg. unknown collector, 5 Oct. 1957, leg. Robin
E. Leech and Edward S. Ross, between 17 Apr. 1949 and 16 Dec. 1949, between 14 Apr. 1949 and
16 Dec. 1949, leg. Jean Hiernaux (INPBC: V1549, RBINS: 7068 to 7075, CAS: 12497, RMCA: 19098,
19097); 2 ?? [2 AL], Ibambi, 10 Nov. 1938, prior to 23 Jan. 1939, leg. Degotte (RMCA: 14987, 14988);
1 ? [AL], Ibembo, between 1 Dec. 1950 and 31 Dec. 1950, leg. Jozef Hutsebaut (RMCA: 20280); 3 ??
[3 AL], Ikela, prior to 4 Jan. 1958, prior to 3 Nov. 1956, leg. P. Lootens (RMCA: 25829, 25831, 27021);
18 ?? [8 AL, 10 UN], Ilimba, unknown date, leg. unknown collector, 3 Jul. 1954, leg. Gaston-François
de Witte (INPBC: W1191.01 to W1191.10, RMCA: 37147 to 37154); 1 ♀, 6 ♂♂, 3 ?? [9 SS, 1 AL],
Irangi, 24 Dec. 1958, leg. Narcisse Leleup, between 24 Aug. 1959 and 16 Nov. 1959, leg. Uwe Rahm,
between 4 Jan. 1957 and 1 Aug. 1957, between 3 Jan. 1957 and 1 Aug. 1957, between 27 Dec. 1956 and
1 Aug. 1957, leg. Heinz Stephan (RMCA: 28037, 28381 to 28384, 27440, 27441, 27444, 27445, 27443);
4 ?? [1 SS, 3 AL], Irangi Forest, between 1 Jan. 1957 and 31 Jan. 1957, leg. Narcisse Leleup, between
1 Nov. 1955 and 10 Aug. 1956, leg. Alexandre Fain (RMCA: 25872 to 25874, 23749); 1 ? [AL], Irumu,
25 km W [= Ituri Forest], prior to 17 Jan. 1955, leg. Pierrard (RMCA: 22800); 1 ? [AL], Ishango
River, 11 Dec. 1954, leg. Gaston-François de Witte (RMCA: 37155); 1 ♂ [UN], Ituri, south, 1 Jan. 1947,
leg. unknown collector (RBINS: 7473); 5 ♀♀, 5 ♂♂ [10 UN], Jadotville [= Likasi], 23 Feb. 1966, leg.
unknown collector (RBINS: 17206.01 to 17206.10); 1 ? [SS], Kabalo, unknown date, leg. Henri
Schouteden (RMCA: 9166); 1 ♂, 4 ?? [2 AL, 1 SK, 2 UN], Kabambi, unknown date, 14 Aug. 1947, leg.
unknown collector, 27 Jul. 1955, leg. Gaston-François de Witte (INPBC: W1464.1, W1464.2, RBINS:
7087, RMCA: 37160, 37161); 2 ♀♀, 6 ♂♂ [8 SN], Kafwe, 14 Jul. 1947, leg. l’ Institut des Parcs
nationaux du Congo belge, 14 Jul. 1947, leg. unknown collector (RBINS: 10727 to 10734); 1 ♂ [AL],
Kailo, 6 Mar. 1964, leg. Alexandre Prigogine (RMCA: 32863); 3 ♀♀, 13 ♂♂, 41 ?? [1 SS, 54 AL,
2 UN], Kakanda Cave, 22 Apr. 1959, leg. Félix / Michel Anciaux de Faveaux, between 12 Aug. 1964
and 11 Jan. 1965, leg. Alexandre Prigogine, between 21 Apr. 1954 and 22 Apr. 1954, leg. R.P. Th. de
Caters (RMCA: 28167, 28168, 28170, 28175, 28177, 28178, 28180, 28182, 28183, 28185 to 28190,
33066, 22851 to 22885, 22891 to 22896); 64 ?? [64 AL], Kakontwe, 15 Aug. 1948, leg. Narcisse Leleup,
23 Feb. 1966, leg. Xavier Misonne (RMCA: 22638 to 22651, 34714 to 34763); 1 ♂ [AL], Kakontwe
Camp [= Kakontwe], 7 Jun. 1956, leg. Félix / Michel Anciaux de Faveaux (RMCA: 23859); 2 ♀♀,
11 ♂♂, 4 ?? [1 SS, 15 AL, 1 SK], Kakontwe Cave, 26 Jun. 1956, 13 Feb. 1957, 8 Jul. 1957, 12 Jul. 1957,
leg. Félix / Michel Anciaux de Faveaux, 6 Feb. 1958, leg. Robin E. Leech and Edward S. Ross, between
19 Sep. 1948 and 22 Jun. 1949, leg. Narcisse Leleup, prior to 24 Sep. 1962, leg. L. Poelman (RMCA:
26744, 26746, 26247, 26251 to 26253, 23814, 23861, 23862, 26232, CAS: 12496, 12527, RMCA:
26725, 26726, 22489, 31215, 31216); 2 ♂♂ [2 SS], Kakyelo, between 13 Nov. 1930 and 21 Feb. 1931,
leg. Gaston-François de Witte and de Crawhez (RMCA: 10850, 10851); 1 ? [UN], Kalenda [Kasai],
168
VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
8 Sep. 1952, leg. unknown collector (RBINS: 7743); 1 ? [UN], Kamande, unknown date, leg. unknown
collector (INPBC: V1057.2a); 2 ♀♀, 4 ♂♂, 1 ? [4 SS, 3 AL], Kamituga, 11 Dec. 1964, between
11 Dec. 1963 and 11 May 1964, between 1 Nov. 1961 and 30 Nov. 1961, leg. Alexandre Prigogine
(RMCA: 32864, 32865, 31151, 32395 to 32397, 32399); 1 ? [AL], Kapolowe, prior to 24 May 1954,
leg. De Smet (RMCA: 22191); 1 ♀, 2 ♂♂ [2 SS, 1 SO], Karibumba, 30 Jan. 1949, between 30 Jan. 1949
and 26 Sep. 1949, leg. Alexandre Prigogine (RMCA: 18864 to 18866); 1 ? [AL], Kasaji Malonga, prior
to 17 Jul. 1952, leg. Fisher (RMCA: 21278); 1 ♂ [SS], Kasenga Road, km 106, 10 May 1956, leg.
M. Lips (RMCA: 23803); 1 ♀ [SS], Kasenye, 10 km N, 22 Sep. 1955, leg. James Paul Chapin (AMNH:
180907); 1 ♀, 2 ?? [2 AL, 1 UN], Kasongo, 1 Dec. 1938, leg. unknown collector, 23 Aug. 1959, leg.
Segers and Benoit Mission (RBINS: 6281.01, RMCA: 29201, 29205); 12 ♀♀, 7 ♂♂ [19 UN], Kasongo
Mwano, 1 Dec. 1938, leg. unknown collector (RBINS: 6281.02 to 6281.20); 1 ♂ [AL], Katanga,
unknown date, leg. Félix / Michel Anciaux de Faveaux (MNHN: ZM-MO-1957-129); 9 ♀♀, 10 ♂♂
[19 AL], Katanga (former) Province, 19 Apr. 1962, 22 Apr. 1962, 24 Apr. 1962, 31 May 1962,
21 Jul. 1962, 13 Aug. 1962, leg. Félix / Michel Anciaux de Faveaux (SMF: 21279 to 21281, 21296,
21297, 21282, 21285, 21278, 21283, 21284, 21286, 21288 to 21295); 2 ?? [1 AL, 1 UN], Katanga s,
unknown date, leg. André Pilette (RMCA: 16223, 16224); 9 ?? [9 SS], Kele Cave 1, 18 Aug. 1938,
between 18 Aug. 1938 and 10 Nov. 1938, leg. J. Bequaert (RMCA: 14916, 14918, 14920, 14921, 14923,
14924, 14917, 14919, 14922); 2 ?? [2 AL], Kiambi, unknown date, leg. Gerard (RMCA: 16225, 16226);
2 ♀♀, 6 ♂♂ [6 SS, 2 SO], Kiamokoto, 7 Jul. 1948, 8 Jul. 1948, 7 Aug. 1948, 8 Aug. 1948, leg. unknown
collector, 8 Jul. 1948, leg. l’ Institut des Parcs nationaux du Congo belge (RBINS: 10763, 10768, 10761,
10762, 10764 to 10767); 1 ♀, 43 ?? [44 AL], Kibwe Lya Mikako Cave, 26 Jan. 1955, leg. Wim
Bergmans and Collard, 28 Dec. 1966, leg. M.J. Celis and M. Lejeune (RMCA: 22949 to 22954, 35171,
35175 to 35203, 35205, 35207, 35209, 35210, 35212 to 35215); 1 ? [AL], Kimuenza Cave, unknown
date, leg. Alexandre Fain (RMCA: 35917); 2 ?? [2 AL], Kisangani, between 1 Jun. 1982 and 30 Jun. 1982,
leg. Marc Colyn (RMCA: 82.031-M-0014, 82.031-M-0015); 1 ♂ [SS], Kisantu, 12 Dec. 1920, leg.
Henri Schouteden (RMCA: 6563); 1 ♂ [SS], Koteli [= Kotili], 15 Jan. 1925, leg. Henri Schouteden
(RMCA: 8918); 2 ?? [2 AL], Kwama Kwama Cave, 13 Jun. 1955, leg. Gaston-François de Witte
(RMCA: 37156, 37157); 6 ♀♀, 5 ♂♂ [1 SA, 10 AL], Kyasala Cave [= Kiasala Cave], 31 Mar. 1956,
15 Apr. 1956, 28 Jul. 1957, between 31 Mar. 1956 and 12 Sep. 1956, leg. Félix / Michel Anciaux de
Faveaux (RMCA: 23811, 23812, 23857, 23858, 26865, 23809, 23852, 23854 to 23856, 23851); 1 ?
[AL], Kyavinionge, 21 May 1954, leg. Gaston-François de Witte (RMCA: 37146); 1 ? [AL], Lake
Kivu, near, between 1 Jan. 1934 and 31 Dec. 1934, leg. Guy Babault (MCZ: 46090); 1 juv ?, 1 ♀, 3 ??
[1 SS, 1 SA, 3 AL], Léopoldville [= Kinshasa], unknown date, leg. J. Camp, 1 Feb. 1942, 12 Nov. 1942,
leg. R. Fiasse, 15 Mar. 1943, leg. Colback (USNM: 21663, 21664, RMCA: 18127, 18128, 31128); 14 ??
[14 UN], Lonzo [= Loso], unknown date, leg. unknown collector (INPBC: V1672.1, V1672.2.01 to
V1672.2.13); 4 ?? [4 AL], Lubudi, large Cave [= Lubudi], 1 Nov. 1948, leg. Narcisse Leleup (RMCA:
22654 to 22657); 5 ?? [5 AL], Lubumbashi, 15 Feb. 1921, between 15 Feb. 1921 and 1 Jan. 1926, leg.
J. Bequaert (RMCA: 16185, 16221, 16222, 16244, 16207); 1 ? [UN], Lufu, prior to 1 Oct. 1898, leg.
unknown collector (RBINS: 208f); 2 ?? [2 AL], Lukatu Cave [= Ntoto Cave], prior to 2 Feb. 2007, leg.
A. Kimbembi-ma-Ibaka (RMCA: a7.006-M-0005, a7.006-M-0006); 17 ?? [16 AL, 1 UN], Lula, between
1 Aug. 1956 and 31 Aug. 1956, leg. Max Poll, prior to 4 Jul. 1958, leg. A.J. Jobaert (RMCA: 25612 to
25627, 27199); 2 ♀♀, 3 ?? [4 SS, 1 AL], Luluabourg [= Kananga], unknown date, 9 May 1922,
10 May 1922, leg. Richard Callewaert, prior to 6 Mar. 1937, leg. Puissant (BMNH: 1927.12.21.6,
1927.12.21.7, RMCA: 7567, 7566, 13708); 1 ♂ [SS], Lundjulu, between 22 Sep. 1952 and 29 Dec. 1952,
leg. Schepens (RMCA: 21585); 4 ♀♀ [3 SS, 1 SO], Mabwe, 28 Jul. 1947, leg. l’ Institut des Parcs
nationaux du Congo belge, 7 Aug. 1947, 8 Aug. 1947, leg. unknown collector (RBINS: 10753, 10771,
10754, 10755); 1 ? [UN], Maginda, unknown date, leg. unknown collector (INPBC: W1077); 1 ? [SO],
Make, 1 Jul. 1989, leg. Tshinyenye (RMCA: 89.057-M-0002); 1 ♀ [SS], Mambabwanga Hill [=
Batala’s Village], 9 Mar. 1925, leg. Edmund Heller (FMNH: 26241); 2 ?? [2 AL], Mani Grot of Nkela
Grot van Mbumba-Nsi [= Mani Cave or Nkela Cave or Mbumba-Nsi], between 1 Jan. 2002 and
169
European Journal of Taxonomy 382: 1–327 (2017)
13 Jul. 2002, leg. A. Kimbembi-ma-Ibaka (RMCA: a2.040-M-0003, a2.040-M-0004); 5 ?? [5 AL],
Masako, unknown date, leg. Ndey B. Ifuta (RMCA: 93.079-M-0330 to 93.079-M-0334); 2 ?? [2 UN],
Masimengo, unknown date, leg. unknown collector (INPBC: V1094.1, V1094.2); 1 ♀, 7 ♂♂ [8 SS],
Masombwe, 10 Oct. 1948, 19 Oct. 1948, leg. unknown collector, 12 Oct. 1948, leg. l’ Institut des Parcs
nationaux du Congo belge (RBINS: 10778, 10780, 10781, 10784, 10785, 10788, 10774, 10795); 1 ♂
[SK], Matupi Cave, 16 Aug. 1947, leg. unknown collector (RBINS: 7088); 1 ♂, 1 ? [1 SS, 1 UN],
Medje, 24 Jan. 1910, leg. Herbert Lang, James Paul Chapin and The American Museum Congo
Expedition, 25 Aug. 1925, leg. Henri Schouteden (AMNH: 49530, RMCA: 9180); 1 ♀, 3 ♂♂, 2 ??
[6 SS], Mongbwalu, 7 Jun. 1939, between 30 May 1939 and 8 Aug. 1939, leg. Janssens, between
21 Jul. 1939 and 27 Nov. 1939, leg. Lepersonne (RMCA: 15630, 15747 to 15750, 15561); 1 ♂ [SK],
Mount Hoyo Caves, 5 Oct. 1957, leg. Robin E. Leech and Edward S. Ross (CAS: 12528); 1 ♂ [SS],
Mount Sombwe, 22 May 1949, leg. unknown collector (RBINS: 10799); 5 ?? [4 SS, 1 AL], Mount
Wago [= Wago, Mount], unknown date, between 1 Jul. 1951 and 31 Jul. 1951, between 1 Jan. 1951 and
31 Dec. 1951, leg. Alexandre Fain, 1 Jul. 1951, leg. unknown collector (BMNH: 1953.496, RBINS:
21660, RMCA: 21460, 23294, 23295); 4 ♀♀, 6 ♂♂ [9 AL, 1 UN], Mpopola Cave, 2 May 1959, leg.
Félix / Michel Anciaux de Faveaux (RMCA: 28193 to 28202); 2 ♀♀, 3 ♂♂ [4 SS, 1 SO], Muhulu,
15 Jan. 1957, between 15 Jan. 1957 and 1 Aug. 1957, leg. Heinz Stephan (RMCA: 27450, 27446 to
27449); 11 ♀♀, 17 ♂♂ [26 SS, 2 SO], Mulungwishi, prior to 15 Dec. 1931, between 10 Jan. 1930 and
14 Jan. 1930, leg. Gaston-François de Witte (RMCA: 11669 to 11695, 11522); 1 ♀, 1 ♂ [2 AL],
Mulungwishi, Cave, 28 Jun. 1956, leg. Félix / Michel Anciaux de Faveaux (RMCA: 23863, 23864); 1
ad ?, 3 imm ?? [4 AL], Musenge, 27 Oct. 1954, leg. Narcisse Leleup (RMCA: 22736 to 22739); 1 ♀
[SS], Mutsora, 4 Nov. 1949, leg. unknown collector (RBINS: 12946); 1 ? [SS], Mutwanga, between
1 Sep. 1937 and 30 Sep. 1937, leg. Henri Martin Hackars (RMCA: 16607); 35 ?? [35 UN], Muyirimbo,
unknown date, leg. unknown collector (INPBC: V1139, V1144.01 to V1144.34); 4 ?? [4 AL],
Mwanakusu [= Mwana-Kusu], 17 Sep. 1959, leg. Segers and Benoit Mission (RMCA: 29200, 29202
to 29204); 2 ad ??, 4 imm ??, 1 ? [6 AL, 1 UN], Mwanakusu Cave, 24 Dec. 1954, leg. Narcisse Leleup,
28 Aug. 1959, leg. Segers and Benoit Mission (RMCA: 22730 to 22735, 29199); 1 ? [UN], Nagero,
prior to 25 Sep. 1951, leg. unknown collector (RBINS: 14637); 6 ?? [6 UN], Nambiliki / Dungu,
unknown date, leg. unknown collector, 18 Aug. 1950, leg. H. de Saeger et al. (INPBC: 2128bis.1,
2128bis.2, RBINS: 13795.1 to 13795.4); 1 ? [AL], Ngamaye, 17 Jun. 1957, leg. Gaston-François de
Witte (RMCA: 37145); 2 ♀♀, 2 ♂♂ [4 UN], Obenge, 7 Feb. 2013, leg. Guy-Crispin Gembu Tungaluna,
Prescott Musaba and André Malekani, 7 Feb. 2013, leg. Prescott Musaba and André Malekani
(UNIKIS: TLL386 to TLL389); 2 ♀♀, 1 ♂, 1 ? [4 SS], Panga, between 1 Sep. 1925 and 30 Sep. 1925,
leg. Henri Schouteden (RMCA: 9128 to 9131); 1 ? [SS], Paulis [= Isiro], between 1 May 1949 and
5 Nov. 1949, leg. Abbeloos (RMCA: 18944); 7 ♂♂ [6 SS, 1 UN], Pelenge, Gorges, 1 Jun. 1947, leg. l’
Institut des Parcs nationaux du Congo belge, 1 Jun. 1947, 4 Jun. 1947, leg. unknown collector (RBINS:
10735, 10737, 10738, 10801, 10739, 10741, 10742); 11 ?? [9 SS, 1 SO, 1 UN], Poko, unknown date,
17 Jun. 1914, 24 Jun. 1914, 12 Jul. 1914, 17 Jul. 1914, leg. Cuthbert Christy, unknown date, leg. unknown
collector (BMNH: 1919.5.8.10, 1919.5.8.11, RMCA: 4688, 4332, 4334, 4333, 4339, 4336a, 4337, 4338,
4335); 1 ♀ [AL], Rambira, 16 Dec. 1959, leg. Uwe Rahm (RMCA: 28904); 1 ♀, 3 ♂♂ [4 AL], RubiTele, 17 Aug. 2013, 20 Aug. 2013, leg. Guy-Crispin Gembu Tungaluna, Prescott Musaba and André
Malekani (UNIKIS: RBTL497 to RBTL499, RBTL528); 17 ?? [17 UN], Rutshuru, unknown date, leg.
unknown collector (INPBC: V124, V1344, V1351, V1637.01 to V1637.12, V414, W26); 1 ? [UN],
Rwere Rwindi, unknown date, leg. unknown collector (INPBC: V1336); 1 ♂, 2 ?? [2 AL, 1 SK], Saga
Saga Cave, 10 Aug. 1947, leg. unknown collector, 16 Jul. 1955, leg. Gaston-François de Witte (RBINS:
7067, RMCA: 37158, 37159); 1 ? [SS], Sakania, 7 Mar. 1948, leg. Jozef Hutsebaut (RMCA: 18244);
1 ? [SS], Saliboko, 22 Sep. 1952, leg. Alexandre Fain (RMCA: 81.012-M-0076); 1 ♀, 2 ♂♂ [3 AL],
St Therese Mission [= Likasi], 8 Feb. 1957, 25 Mar. 1957, 20 Jan. 1958, leg. Félix / Michel Anciaux
de Faveaux (RMCA: 27537, 26255, 26246); 2 ?? [2 AL], Stanleyville [= Kisangani], unknown date,
leg. Weyns, 5 Apr. 1928, leg. Collart (RMCA: 16214, 16217); 1 ♂ [SS], Tandala, 11 Jun. 1979, leg.
170
VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
C. Brian Robbins (USNM: 537709); 1 ♀, 1 ♂, 1 ? [3 SS], Tembwe, unknown date, leg. Henri Schouteden
(RMCA: 8854, 8855, 9178); 1 ad ♀, 1 foetus ♀, 2 ♀♀ [2 SS, 2 UN], Teturi, 5 Aug. 1947, 21 Aug. 1947,
15 Jun. 1948, leg. unknown collector (RBINS: 7077, 7076, 7472.1, 7472.2); 17 ?? [16 AL, 1 UN],
Thysville [= Mbanza-Ngungu], 11 Feb. 1987, leg. unknown collector, between 1 Aug. 1949 and
31 Aug. 1949, leg. Narcisse Leleup (RBINS: 22427, RMCA: 22586 to 22601); 2 ?? [2 AL], Thysville,
Cave b 13 a [= Banjan Caves], between 1 Jul. 1949 and 31 Jul. 1949, leg. Narcisse Leleup (RMCA:
22584, 22585); 1 ♀ [SS], Tschambutscha [= Tshambutschu], between 31 Dec. 1956 and 1 Aug. 1957,
leg. Heinz Stephan (RMCA: 27442); 2 ?? [2 AL], Tshakala, between 1 Jan. 1982 and 31 Jan. 1982, leg.
Marc Colyn (RMCA: 82.019-M-0003, 82.019-M-0004); 1 ♂ [UN], Tshapona [= Tshiapona],
5 Dec. 1937, leg. unknown collector (RBINS: 6119); 1 ♀ [AL], Tshuapa District, between 1 Jan. 1960
and 31 Dec. 1964, leg. P. Lootens (RMCA: 34181); 4 ?? [4 UN], Uduku, 23 Jul. 1952, leg. H. de Saeger
et al. (RBINS: 13797.1 to 13797.3, 13799); 2 ♀♀, 1 ♂, 2 ?? [1 SS, 1 AL, 1 SO, 2 UN], Uduku Moke
[= Uduku], 24 Jul. 1952, leg. H. de Saeger et al., 24 Jul. 1952, between 24 Jul. 1952 and 28 Dec. 1972,
leg. H. de Saeger (RBINS: 13798.1, 13798.2, RMCA: 36816, 36825, 36817); 1 ♂ [AL],
Uma, 10 Apr. 2014, leg. Prescott Musaba and André Malekani (UNIKIS: UMA503); 1 ♀ [SS], Utu
[River], between 6 Feb. 1959 and 28 Jun. 1960, leg. Uwe Rahm (RMCA: 28903); 9 ?? [8 AL, 1 UN],
Wafanya [= Wafania], 30 Sep. 1988, 12 Oct. 1991, 19 May 1992, between 12 Oct. 1991 and 18 Oct. 1991,
between 1 Dec. 1991 and 31 Dec. 1991, between 1 May 1991 and 31 May 1991, between 1 Apr. 1992
and 30 Apr. 1992, between 1 Jan. 1992 and 31 Jan. 1992, leg. P. Lootens (RMCA: 91.076-M-0110,
92.079-M-0077, 89.020-M-0038, 92.079-M-0078, 92.079-M-0079, 91.076-M-0048, 92.079-M-0076,
92.079-M-0149, 91.076-M-0101); 1 ? [SS], Wago, Mount, between 1 Jul. 1951 and 31 Jul. 1951, leg.
Alexandre Fain (RMCA: 23293); 3 ?? [3 AL], Zambi, prior to 31 Jan. 1899, leg. J.M. Derscheid (RMCA:
150a to 150c); 1 ? [SO], Zobia, prior to 18 Apr. 1946, leg. unknown collector (RBINS: 6974).
RWANDA: 1 ? [UN], Kissenji [= Gisenyi], unknown date, leg. Stegmann-Stein (ZMB: 67685).
Hipposideros camerunensis Eisentraut, 1956
DEMOCRATIC REPUBLIC OF THE CONGO: 1 ♂ [SN], Shabunda, unknown date, leg. Uwe Rahm
(BMNH: 1963.1166).
Hipposideros fuliginosus (Temminck, 1853)
DEMOCRATIC REPUBLIC OF THE CONGO: 3 ♂♂ [3 SS], Epulu, 2 May 1951, 3 May 1951, leg.
A. Schwartz and R.N. Porter (KU: 151336 to 151338); 1 ♂ [SA], Equateur (former) Province, prior to
7 Oct. 1996, leg. unknown collector (SMF: 84411); 2 ♂♂ [1 SK, 1 SN], Irumu, 40 mi [= 64 km] SW
[= Ituri Forest], unknown date, leg. Howard de Walden (BMNH: 1930.11.11.143, 1930.11.11.144); 1 ♂
[SS], Yalosemba, 26 Jun. 1979, leg. Lynn W. Robbins (USNM: 537713).
Hipposideros ruber (Noack, 1893)
BURUNDI: 20 ♂♂ [20 AL], Grotte de Rubindi [= Rubindi], 9 Apr. 1969, 8 Mar. 1970, leg. P.L. Niort
(MHNG: 1895.091 to 1895.100, 1896.001 to 1896.010); 1 ♂ [AL], Mpinga Cave [= Mpinga Mission],
1 Jan. 1969, leg. P.L. Niort (MHNG: 1896.011); 1 ♀ [SB], Mumushwizi Valley, 17 Aug. 1993, leg. Jay
L. Udelhoven (FMNH: 155973); 3 ?? [3 SK], Nyanza-Lac, 10 km N, 3 Mar. 1979, leg. Heinz Stephan
(SMF: 57576 to 57578); 1 ♀, 1 ♂ [1 SB, 1 SN], Teza, 2.3 km N, 0.7 km W [= Kibira National Park],
10 Mar. 1990, 14 Mar. 1990, leg. Julian C. Kerbis Peterhans (FMNH: 137627, 137628).
DEMOCRATIC REPUBLIC OF THE CONGO: 9 ♀♀, 8 ♂♂, 21 ?? [14 SS, 11 AL, 2 SK, 11 UN],
unknown locality, unknown date, 1 Jan. 1948, leg. unknown collector, unknown date, leg. Guy-Crispin
Gembu Tungaluna (BMNH: 1926.7.6.69, 1930.11.11.105 to 1930.11.11.108, 1930.11.11.110 to
1930.11.11.112, 1930.11.11.114 to 1930.11.11.119, 1930.11.11.128, 1990.231, 1990.232,
171
European Journal of Taxonomy 382: 1–327 (2017)
RMCA: a1.097-M-0554 to a1.097-M-0564, RBINS: 8204.01 to 8204.10); 1 ♀, 2 ♂♂ [3 SS],
Aba, 13 Jan. 1911, 13 Dec. 1911, leg. Herbert Lang, James Paul Chapin and The American Museum
Congo Expedition (AMNH: 49172, 49127, RMCA: 12370); 23 ?? [1 SS, 6 AL, 16 UN], Aka, 15 May
1950, between 15 May 1952 and 28 Dec. 1972, leg. H. de Saeger, 23 Apr. 1952, 15 May 1952, leg. H. de
Saeger et al. (RMCA: 36802 to 36807, RBINS: 13765.01 to 13765.11, 13749.1 to 13749.5, RMCA:
36711); 1 ♀ [SS], Angumu, 1 Jul. 1937, leg. James Paul Chapin (AMNH: 119614); 7 ♀♀, 6 ♂♂, 3 ??
[7 SS, 9 AL], Avakubi, 29 Sep. 1909, 1 Oct. 1909, 15 Nov. 1909, 17 Nov. 1909, 22 Nov. 1909,
24 Nov. 1909, 27 Nov. 1909, 28 Nov. 1909, 1 Dec. 1909, 2 Dec. 1909, between 1 Oct. 1909 and
1 Oct. 1915, leg. Herbert Lang, James Paul Chapin and The American Museum Congo Expedition
(AMNH: 49372, 49369, 49373, 49178, 49179, 49375 to 49377, 49180, MCZ: 17373, AMNH: 49370,
49181, 49182, 49171, 49374, MCZ: 17223); 2 ?? [2 SS], Bolobo, unknown date, leg. the Royal Museum
for Central Africa (RMCA) (BMNH: 1954.818, 1954.819); 1 ? [AL], Buta, prior to 31 May 1939, leg.
Jozef Hutsebaut (RMCA: 15329); 1 ad ♀, 1 ♂ [2 UN], Cel II, 8 Feb. 1951, 9 Feb. 1951, leg. H. de
Saeger et al. (RBINS: 13737, 13738); 1 ♀, 2 ♂♂ [2 SA, 1 SK], Dethiox Caves [= Kambove], 4 Jun. 1960,
leg. Félix / Michel Anciaux de Faveaux (SMF: 18922 to 18924); 1 ♀ [SA], Detioux Cave [= Kambove],
4 Jun. 1960, leg. Félix / Michel Anciaux de Faveaux (SMF: 18925); 18 ?? [18 UN], Dungu, unknown
date, leg. unknown collector (RBINS: 8203.01 to 8203.18); 1 ♂ [SS], Eala, between 1 Aug. 1936 and
31 Aug. 1936, leg. J. Ghesquière (RMCA: 13411); 8 ♀♀, 11 ♂♂ [1 SS, 12 AL, 1 SO, 1 UN, 2 SB, 2 SN],
Epulu, 2 May 1951, 3 May 1951, leg. A. Schwartz and R.N. Porter, 21 Apr. 1981, leg. John A. Hart,
26 Aug. 2015, leg. André Malekani, 6 Jul. 1991, leg. Julian C. Kerbis Peterhans (KU: 151344, FMNH:
138139, 138140, UNIKIS: EPLU637, EPLU639, EPLU641 to EPLU643, KU: 151343, FMNH: 149407
to 149416); 3 ♀♀, 1 ♂ [1 SS, 3 AL], Faradje, 20 Nov. 1911, 22 Oct. 1912, 25 Oct. 1912, leg. Herbert
Lang, James Paul Chapin and The American Museum Congo Expedition (AMNH: 49168, 49381, 49385,
49416); 1 ♀ [UN], Gangala-na-Bodio, 4 May 1951, leg. H. de Saeger et al. (RBINS: 13739); 1 ♀ [UN],
Grottes Kiwakishi, 18 Oct. 1974, leg. Michael D. Gallagher (HZM: 55.8027); 1 ? [SS], Irangi, unknown
date, leg. Alexandre Fain (BMNH: 1956.465); 3 ♂♂ [3 AL], Irangi, slightly E of [= Irangi],
15 Oct. 1990, leg. Wim Bergmans (ZMA: MAM.24180, MAM.24181, MAM.24185); 9 ♀♀ [9 AL],
Ituri Forest, 19 Jul. 1956, leg. Alvin Novick (MCZ: 48226 to 48232, 48234, 48235); 6 ♂♂ [2 SA,
4 AL], Jomba, 4 Sep. 1976, leg. Frieder Meier (MHNG: 1873.062 to 1873.067); 1 ? [UN], Kabare,
unknown date, leg. unknown collector (RMCA: 15839); 2 ?? [2 SS], Kabatibi Cave [= Mount Hoyo
Caves], unknown date, leg. The Cambridge East Africa Expedition 1959 (BMNH: 1960.83, 1960.84);
1 ad ♀ [SB], Kabobo Forest, 16 Feb. 2007, leg. Michael H. Huhndorf (FMNH: 195085); 1 ♂ [SS],
Kabwe, 12 Nov. 1973, leg. Western New Mexico University (FMNH: 152682); 1 ♂ [AL], Kadjidji
[= Kajuju], unknown date, leg. Guy Babault (MNHN: ZM-MO-1984-1344); 4 ♀♀, 17 ♂♂, 1 ? [8 AL,
13 UN, 1 SS], Kakanda Cave, 31 May 1962, leg. Félix / Michel Anciaux de Faveaux, 31 May 1962,
leg. unknown collector, between 13 Aug. 1964 and 11 Jan. 1965, leg. Alexandre Prigogine (MHNG:
1047.067 to 1047.074, RBINS: 14563 to 14575, RMCA: 33067); 6 ♀♀ [1 SA, 5 AL], Kakontwe Cave,
26 Jun. 1956, 8 Jul. 1957, 12 Jul. 1957, leg. Félix / Michel Anciaux de Faveaux (MHNG: 1046.039 to
1046.041, 922.096, 1046.037, 1046.038); 3 ♀♀, 3 ♂♂ [6 UN], Kasoma, 13 Aug. 1962, 18 Aug. 1962,
leg. unknown collector (RBINS: 14587 to 14589, 14584 to 14586); 2 ♀♀, 4 ♂♂ [1 SA, 4 AL, 1 UN],
Kasoma Cave, 13 Aug. 1962, 18 Aug. 1962, leg. Félix / Michel Anciaux de Faveaux (MHNG: 1047.075
to 1047.079, RBINS: 14583); 4 ad ♀♀, 1 juv ♀, 10 juv ♂♂, 27 ♀♀, 12 ♂♂, 81 ?? [54 AL, 81 UN],
Keroma, 21 Apr. 1952, leg. H. de Saeger et al., 21 Apr. 1952, leg. H. de Saeger (RBINS: 13751.01 to
13751.10, 13752.01 to 13752.10, 13753.01 to 13753.10, 13754.01 to 13754.10, 13755.01 to 13755.10,
13756.01 to 13756.10, 13757.01 to 13757.10, 13758.01 to 13758.11, RMCA: 36746 to 36799); 1 ♀, 1 ♂
[2 SS], Kilemue, 25 Oct. 1966, between 25 Oct. 1966 and 24 May 1967, leg. Alexandre Prigogine
(RMCA: 34802, 34801); 2 ♀♀ [2 UN], Kinshasa, between 16 Jan. 1975 and 20 Jan. 1975, leg. Michael
D. Gallagher (HZM: 53.8025, 54.8026); 7 ?? [7 AL], Kisangani, between 1 Jun. 1982 and 30 Jun. 1982,
leg. Marc Colyn (RMCA: 82.031-M-0016 to 82.031-M-0022); 2 ?? [2 AL], Kisantu, prior to
17 Nov. 1913, leg. R.P. Vanderijst (RMCA: 2416a, 2416b); 1 ♀, 14 ♂♂, 2 ?? [11 SA, 6 AL], Kivu
172
VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
region, 8 Mar. 1964, 11 Mar. 1964, leg. P. Kunkel, 31 Jan. 1984, 2 Feb. 1984, 12 Feb. 1984, between
11 Aug. 1989 and 14 Aug. 1989, leg. unknown collector (SMF: 31831, 64983, 64982, 64981, 31832,
77087 to 77098); 1 juv ♂ [SA], Koloka, between 1 Jun. 1911 and 30 Jun. 1911, leg. unknown collector
(SMF: 6364); 1 ♀ [AL], Kumbi, 25 Nov. 1993, leg. Ngenge Masumbuko Kamitongo (ZMA: MAM.24900);
8 ♀♀, 2 ♂♂ [1 SA, 4 AL, 5 UN], Kyasala Cave [= Kiasala Cave], 15 Apr. 1956, 19 Apr. 1962,
22 Apr. 1962, leg. Félix / Michel Anciaux de Faveaux, 19 Apr. 1962, 22 Apr. 1962, leg. unknown
collector (MHNG: 922.097 to 922.099, 1047.065, RBINS: 14590, MHNG: 1047.066, RBINS: 14591 to
14594); 1 ♀ [AL], Kyasala Grande Grotte [= Kiasala Cave], 28 Jul. 1957, leg. Félix / Michel Anciaux
de Faveaux (MHNG: 1046.042); 1 ♂ [SS], Léopoldville [= Kinshasa], 10 Jul. 1909, leg. Herbert Lang,
James Paul Chapin and The American Museum Congo Expedition (AMNH: 49110); 1 ? [SS], Libenge,
10 Jun. 1937, leg. Leontovitch (RMCA: 13988); 1 ♀ [SA], Lubudi, 24 Apr. 1962, leg. Félix / Michel
Anciaux de Faveaux (SMF: 21287); 1 ♀ [AL], Luhohov (River), 18 Oct. 1990, leg. Wim Bergmans
(ZMA: MAM.24186); 4 ♀♀, 1 ♂, 14 ?? [6 SS, 13 AL], Lukolela, 19 Aug. 1930, 14 Sep. 1930, leg.
Franklin Edson, 18 Aug. 1930, 19 Aug. 1930, 15 Nov. 1930, 16 Nov. 1930, 21 Nov. 1930, leg. James
Paul Chapin (AMNH: 86772, 86773, 86929 to 86931, 86928, 86774 to 86776, 86881, 86918 to 86926);
16 ♀♀ [15 SS, 1 UN], Luluabourg [= Kananga], unknown date, leg. Richard Callewaert (BMNH:
1926.11.1.47 to 1926.11.1.53, 1926.7.6.62_1, 1926.7.6.62_2, 1926.7.6.64 to 1926.7.6.68, 1926.7.6.7,
1926.7.6.70); 1 ♀ [UN], Makpe, 8 Aug. 1951, leg. H. de Saeger et al. (RBINS: 13743); 6 ad ♀♀, 2
juv ??, 2 ♂♂, 23 ?? [33 UN], Maleli, 9 Feb. 1951, 14 Jun. 1951, 18 Mar. 1952, leg. H. de Saeger et al.
(RBINS: 13742.1, 13742.2, 13734.1, 13734.2, 13735.1, 13735.2, 13744.1 to 13744.4, 13745.1 to
13745.4, 13746.1 to 13746.9, 13747.1, 13747.2, 13748.1, 13748.2, 13741.1 to 13741.6); 4 ♀♀, 6 ♂♂,
6 ?? [1 SS, 11 AL, 4 SO], Malili, 13 Jun. 1951, 14 Jun. 1951, 18 Mar. 1952, between 18 Mar. 1952 and
28 Dec. 1972, leg. H. de Saeger (RMCA: 36706, 36707, 36712 to 36716, 36742, 36800, 36801, 36709,
36710, 36743 to 36745, 36708); 1 ♀ [AL], Masako, 11 Jun. 1989, leg. Ndey B. Ifuta (RMCA: 93.079M-0335); 1 ♀ [SS], Mbuji-Mayi, 32 km SE, 25 Feb. 1974, leg. Western New Mexico University
(FMNH: 152683); 1 juv ♀, 5 ♀♀, 4 ♂♂, 3 ?? [13 UN], Medje, 20 Jan. 1910, 21 Jan. 1910, 22 Jan. 1910,
24 Jan. 1910, 23 Mar. 1910, 27 Apr. 1910, 3 Aug. 1910, 11 Mar. 1914, leg. Herbert Lang, James Paul
Chapin and The American Museum Congo Expedition (AMNH: 49185, 49186, 49104, 49184, 49105 to
49107, 49187, 49108, 49380, 49378, 49379, 49109); 3 juv ??, 1 ♀, 1 ♂, 7 ?? [2 SS, 1 SA, 9 AL], Mount
Hoyo Caves, 4 May 1955, leg. Waldo LaSalle Schmitt and E. Baker (USNM: 301715 to 301726); 2 ♂♂
[2 UN], Mulungwishi, 27 Dec. 1959, 4 Jun. 1960, leg. unknown collector (RBINS: 14581, 14582); 1 ♀
[SA], Mulungwishi, Cave, 28 Jun. 1956, leg. Félix / Michel Anciaux de Faveaux (MHNG: 922.095);
1 ♀, 5 ♂♂ [6 AL], Musenge, 1 May 1992, 2 May 1992, 6 May 1992, leg. Ngenge Masumbuko Kamitongo
(ZMA: MAM.24550, MAM.24556, MAM.24552, MAM.24554, MAM.24551, MAM.24555); 2 ♀♀
[2 UN], Musosa, near, between 1 May 1941 and 31 May 1941, leg. unknown collector (RBINS: 14576,
14577); 16 ♀♀, 10 ♂♂, 17 ?? [10 SS, 25 AL, 1 SO, 7 UN], Nambiliki / Dungu, 18 Aug. 1950, leg. H. de
Saeger et al., 18 Aug. 1950, between 18 Aug. 1950 and 28 Dec. 1972, leg. H. de Saeger (RBINS:
13730.1, 13730.2, 13731.1, 13731.2, 13732.1, 13732.2, 13733, RMCA: 36701, 36717 to 36741, 36695
to 36700, 36702 to 36705); 1 ♀ [SS], N’gombe [= Ngombe], 10 Nov. 1921, leg. Henri Schouteden
(RMCA: 6561); 1 ♀, 1 ♂ [1 SS, 1 SA], Niangara, 19 Nov. 1910, 12 Dec. 1910, leg. Herbert Lang,
James Paul Chapin and The American Museum Congo Expedition (AMNH: 49388, 49183); 1 ad ♂,
1 ♂, 9 ?? [5 SA, 6 AL], Niapu, 27 Jan. 1914, 28 Jan. 1914, 27 Feb. 1914, leg. Herbert Lang, James Paul
Chapin and The American Museum Congo Expedition (AMNH: 49405 to 49408, 49410 to 49413,
49414 [holotype Hipposideros caffer niapu J.A. Allen, 1917], MCZ: 17224 [topotype Hipposideros
caffer niapu J.A. Allen, 1917], AMNH: 49409); 2 ♂♂ [2 AL], Nyabiondo, 28 Dec. 1990, 30 Dec. 1990,
leg. J.P. Lubula Bulambo and Ngenge Masumbuko Kamitongo (ZMA: MAM.24352, MAM.24353);
1 ♀ [UN], Obenge, 7 Feb. 2013, leg. Prescott Musaba and André Malekani (UNIKIS: TLL374); 1 ♀,
1 ♂ [2 SA], Oso River, between 1 Jan. 1957 and 31 Jan. 1957, leg. Félix / Michel Anciaux de Faveaux
(SMF: 22298, 22299); 3 ♀♀ [2 SA, 1 AS], Paoni, 5 Aug. 1976, 6 Aug. 1976, leg. Frieder Meier (MHNG:
1873.059 to 1873.061); 1 ? [UN], Pidigala River, 23 Apr. 1952, leg. H. de Saeger et al. (RBINS:
173
European Journal of Taxonomy 382: 1–327 (2017)
13750); 3 ?? [3 AL], Poko, 1 Aug. 1913, 30 Aug. 1913, leg. Herbert Lang, James Paul Chapin and The
American Museum Congo Expedition (AMNH: 49400 to 49402); 1 ♀, 1 ♂, 2 ?? [2 SS, 2 AL], Rutshuru,
14 Apr. 1927, leg. James Paul Chapin, Dewitt L. Sage, Frank P. Mathews and The Ruwenzori-Kivu
Expedition, 24 Feb. 1927, 25 Feb. 1927, leg. David H. Linder (AMNH: 82395, MCZ: 23378, 23379,
25057); 1 ? [SS], Saliboko, unknown date, leg. Alexandre Fain (BMNH: 1953.497); 1 ♀, 2 ♂♂ [2 SS,
1 UN], St. Jos. (Mission) [= Mikalaya], 31 May 1924, 29 Oct. 1924, leg. Richard Callewaert (AMNH:
86150, 86168, 86170); 3 ♀♀ [3 UN], Tantara Cave [= Shinkolobwe], 4 Jun. 1960, leg. unknown
collector (RBINS: 14578 to 14580); 1 ? [UN], Thysville [= Mbanza-Ngungu], 11 Feb. 1987, leg.
unknown collector (RBINS: 22430); 2 ?? [2 AL], Tshakala, between 1 Jan. 1982 and 31 Jan. 1982, leg.
Marc Colyn (RMCA: 82.019-M-0001, 82.019-M-0002); 1 ♂ [AL], Tshamalenge Cave, 25 Aug. 1957,
leg. Félix / Michel Anciaux de Faveaux (MHNG: 1046.043); 1 ♀ [SS], Tshikapa, 26 Nov. 1921, leg.
Henri Schouteden (RMCA: 6562); 2 ♀♀, 1 ♂ [3 AL], Uma, 10 Apr. 2014, 12 Apr. 2014, leg. GuyCrispin Gembu Tungaluna, Prescott Musaba and André Malekani (UNIKIS: UMA513, UMA622,
UMA625); 4 ♀♀, 6 ♂♂ [5 SS, 5 AL], Yalosemba, 17 Jun. 1979, 27 Jun. 1979, leg. C. Brian Robbins,
18 Jun. 1979, leg. Frieder Meier (USNM: 463482 to 463484, 537714, 537715, 537718, MHNG:
1871.015, 1871.016, USNM: 537716, 537717).
RWANDA: 1 ♀, 3 ♂♂ [4 SA], unknown locality, between 1 Jan. 1957 and 31 Dec. 1957, leg. unknown
collector (SMF: 29050 to 29053); 1 ? [SK], Kibumbu Crater, 9 Dec. 1991, leg. J.R. Wilson and Anthony
Michael Hutson (BMNH: 1992.389); 11 ad ♀♀, 2 ad ♂♂, 1 sad ♀, 12 juv ♀♀, 5 juv ♂♂ [29 SA, 2 AL],
Kidaho, 10 Jun. 1982, 11 Jun. 1982, 13 Jun. 1982, leg. Benny Baeten (RMCA: 87.046-M-0215 to
87.046-M-0245); 2 ?? [2 UN], Lugarama, unknown date, leg. unknown collector (INPBC: V3041.1,
V3041.2); 1 ? [UN], Nyakichozi, unknown date, leg. unknown collector (INPBC: V3118.1); 1 ad
♂ [SA], Nyakinama, 24 Oct. 1981, leg. Frits De Vree, Benny Baeten and Victor Van Cakenberghe
(RMCA: 82.006-M-0896); 15 ?? [15 SA], Nyundo, 22 Oct. 1980, leg. Danny Meirte (RMCA: 80.055M-0098 to 80.055-M-0112); 4 ♂♂ [1 SA, 3 AL], Ruhengeri, 20 Sep. 1976, leg. Frieder Meier (MHNG:
1873.068 to 1873.071); 1 ad ♀, 5 ad ♂♂, 2 juv ♀♀, 10 juv ♂♂ [14 SA, 4 AL], Rukira, 22 Nov. 1981,
24 Nov. 1981, 26 Nov. 1981, leg. Frits De Vree, Benny Baeten and Victor Van Cakenberghe (RMCA:
82.006-M-0897 to 82.006-M-0914); 1 ad ♂ [UN], Rwankalo, unknown date, leg. unknown collector
(RBINS: 13112).
Macronyctus gigas (Wagner, 1845)
DEMOCRATIC REPUBLIC OF THE CONGO: 3 ?? [3 AL], unknown locality, unknown date, leg.
Guy-Crispin Gembu Tungaluna (RMCA: a1.097-M-0550 to a1.097-M-0552); 1 ♂ [SS], Butembo,
12 Aug. 1947, leg. unknown collector (RBINS: 7062); 2 ♂♂ [2 AL], Île Mbiye [= Mbiye Island],
19 Feb. 2012, 31 May 2012, leg. Prescott Musaba (RMCA: a1.097-M-3213, a1.097-M-3212); 1 ♂ [SS],
Kakontwe, unknown date, leg. the Royal Museum for Central Africa (RMCA) (BMNH: 1954.820);
1 ♀, 1 ♂ [2 SS], Mbuji-Mayi, 27 Dec. 1973, leg. Western New Mexico University (FMNH: 152633,
152634); 1 ad ♀ [SS], Niangara, 2 Jun. 1913, leg. Herbert Lang, James Paul Chapin and The American
Museum Congo Expedition (AMNH: 49103 [holotype Hipposideros gigas niangaræ J.A. Allen, 1917]);
1 ? [SO], Nsiana, 1 Jan. 1927, leg. Cadgepole (MCZ: 23376); 1 ♀ [SS], Yalosemba, 17 Jun. 1979, leg.
C. Brian Robbins (USNM: 537710).
Macronycteris Gray, 1866
Macronycteris vittatus (Peters, 1852)
BURUNDI: 1 ♂ [UN], Bujumbura, 23 Apr. 1982, leg. Danny Meirte (RMCA: 82.016-M-0043); 1 ♂
[AL], Kigwena Hill, 24 Apr. 1982, leg. Danny Meirte (RMCA: 82.016-M-0068).
174
VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
DEMOCRATIC REPUBLIC OF THE CONGO: 1 ? [SO], unknown locality, unknown date, leg.
unknown collector (RBINS: 205); 1 ♀ [SS], Baliko, 14 Jun. 2012, leg. William (“Bill”) T. Stanley
(FMNH: 219682); 2 ?? [2 AL], Boende, unknown date, between 1 Feb. 1970 and 28 Feb. 1970,
leg. P. Lootens (RMCA: 75.035-M-0061, 75.035-M-0021); 3 ♀♀, 3 ♂♂ [5 SS, 1 SO], Bokalakala,
between 25 Apr. 1959 and 13 Jul. 1959, leg. N’Kele (RMCA: 27940 to 27945); 1 ♀ [UN], Bomane
[= Bomaneh], 21 May 2010, leg. Guy-Crispin Gembu Tungaluna, Prescott Musaba and José Akaibe
(UNIKIS: CRT2125); 1 ? [AL], Congo da Lemba, prior to 22 Jan. 1900, leg. Paul Van Bellighen
(RMCA: 410); 1 ? [AL], Ganda-Sundi, 29 Mar. 1964, leg. Alexandre Fain (RMCA: 33261); 7 ♀♀,
10 ♂♂ [17 UN], Goroumbwa Mine, 17 May 1999, 18 May 1999, 19 May 1999, 21 Oct. 1999,
22 Oct. 1999, 23 Oct. 1999, 25 Oct. 1999, leg. Robert (Bob) Swanepoel (TM: DRC0071, DRC0089,
DRC0124, DRC0148, DBA071, DBA072, DBA075, DBA112, DBA124, DBA185, DBA186, DBA194,
DBA199, DBA200, DBA280, DBA288, DBA289); 1 ? [UN], Kakontwe Cave, 5 Jul. 1948, leg.
Narcisse Leleup (RMCA: 22686); 1 ad (gravid) ♀ [AL], Kalomoni Cave [= Kakontwe], 31 Jan. 1964,
leg. Félix / Michel Anciaux de Faveaux (RMCA: 33992); 1 ? [AL], Kasaji, prior to 9 Jun. 1959, leg.
Fisher (RMCA: 27891); 1 ? [SS], Kiseny [= Kasenye], unknown date, leg. Henri Schouteden (RMCA:
16286); 1 ♀, 1 ♂ [1 SS, 1 SK], Kunungu, unknown date, leg. Henri Schouteden, prior to 25 Mar. 1929,
leg. Ngwe (RMCA: 7291b, 9707); 1 ? [AL], Lofoi [River], prior to 31 Mar. 1902, leg. Edgar Verdick
(RMCA: 495); 1 ? [SS], Lovanium University [= Kinshasa], between 1 May 1962 and 31 May 1962,
leg. De Bont (RMCA: 31200); 1 ♂ [UN], Marungu, unknown date, leg. Richard Böhm (ZMB: 6312);
1 ♀ [AL], Mosenge, 7 Aug. 1986, leg. WHO and Marc Colyn (RMCA: 88.011-M-0012); 1 ♂ [SS],
Mutsora, 3 Jul. 1951, leg. unknown collector (RBINS: 13325); 1 ? [SK], Mwanakusu Cave, between
1 Sep. 1959 and 30 Sep. 1959, leg. Benoit (RMCA: 28864); 1 ♀, 1 ? [1 SS, 1 AL], Panda [= Pande],
22 Dec. 1963, leg. Félix / Michel Anciaux de Faveaux, between 28 Jan. 1956 and 17 May 1957, leg.
De Smet (RMCA: 33991, 26211); 1 ? [SK], Tanganyika, Congo shore of Lake, prior to 19 Oct. 1889,
leg. unknown collector (RBINS: 205b).
Megadermatidae H. Allen, 1864
Cardioderma Peters, 1873
Cardioderma cor (Peters, 1872)
DEMOCRATIC REPUBLIC OF THE CONGO: 1 ♂ [AL], Rubi-Tele, 17 Aug. 2013, leg. Guy-Crispin
Gembu Tungaluna, Prescott Musaba and André Malekani (UNIKIS: RBTL493).
Lavia Gray, 1838
Lavia frons (E. Geoffroy St.-Hilaire, 1810)
BURUNDI: 1 ? [AL], Bujumbura, between 1 Jul. 1968 and 31 Jul. 1968, leg. Lewall (RMCA: 35152);
2 ♂♂ [2 SK], Resha, 11 Mar. 1979, 16 Mar. 1979, leg. unknown collector (SMF: 57496, 57497); 1 ♂
[AL], Russissi, bank [= Rusizi Delta], 27 Apr. 1982, leg. Danny Meirte (RMCA: 82.016-M-0146).
DEMOCRATIC REPUBLIC OF THE CONGO: 1 ♀, 1 ♂ [2 UN], unknown locality, prior to 8 Jan. 1940,
leg. unknown collector (RBINS: 6279.1, 6279.2); 1 ♀, 2 ?? [2 AL, 1 UN], Albertville [= Kalemie],
20 Nov. 1946, leg. unknown collector, prior to 18 Nov. 1959, leg. H. Bomans (RBINS: 9067, RMCA:
29298, 29299); 2 ♂♂ [2 SS], Bagbele Moko, 18 Nov. 1949, 19 Apr. 1950, leg. H. de Saeger et al.
(RBINS: 13632, 13640); 1 ? [SO], Bambesa, between 1 Apr. 1938 and 30 Apr. 1938, leg. J.M. Vrijdagh
(RMCA: 14679); 1 ♂, 1 ? [1 AL, 1 SK], Baraka, 5 mi [= 8 km] N, 10 Jan. 1957, leg. Robin E. Leech
and Edward S. Ross (CAS: 12487, 12526); 4 ?? [2 AL, 2 UN], Bauelendu, unknown date, leg. unknown
collector, 31 Jan. 1955, 2 Feb. 1955, leg. Gaston-François de Witte (INPBC: W1358, W1360, RMCA:
37141, 37140); 1 ? [SS], Beni, prior to 13 May 1939, leg. Lisfranc (RMCA: 15303); 1 ? [AL], Bokoro,
prior to 19 Nov. 1952, leg. E. Jans (RMCA: 21520); 1 ♀ [SO], Bomili, 16 Aug. 1938, leg. Duncan
175
European Journal of Taxonomy 382: 1–327 (2017)
M. Hodgson and William F. Coultas (MCZ: 38026); 1 ? [SS], Bosodula [= Boso-Dula], 19 Oct. 1941,
leg. J.M. Vrijdagh (RMCA: 17752); 1 ? [AL], Buta, unknown date, leg. Jozef Hutsebaut (RMCA:
16571); 1 ♀, 1 ♂, 1 ? [3 AL], Catchpole’s Farm, 28 Jul. 1924, 29 Jul. 1924, 27 Aug. 1924, leg. Edmund
Heller (FMNH: 30705, 30703, 30704); 2 ♀♀, 4 ♂♂, 3 ?? [5 SS, 1 AL, 1 SO, 2 UN], Cel I [= Bagbele],
unknown date, leg. unknown collector, 12 Feb. 1950, 5 Jun. 1950, 22 Nov. 1950, leg. H. de Saeger,
19 Jul. 1950, 31 Oct. 1950, 6 Dec. 1950, 12 Dec. 1950, leg. H. de Saeger et al. (INPBC: 1441, RMCA:
36938, RBINS: 13634, 13642, RMCA: 36939, 36940, RBINS: 13643, RMCA: 36945, RBINS: 13635);
1 ♀, 4 ♂♂, 3 ?? [1 SS, 2 SO, 5 UN], Cel II, unknown date, leg. unknown collector, 16 Jan. 1951,
17 Mar. 1951, 29 May 1951, 21 Apr. 1952, 8 May 1952, leg. H. de Saeger et al., 29 Jun. 1951, 7 Mar. 1952,
leg. H. de Saeger (INPBC: 4392, RBINS: 13646, 13645, 13652, 13647, RMCA: 36942, 36943, RBINS:
13655); 1 ? [AL], Dubulu, 14 Jan. 1932, leg. H.J. Bredo (RMCA: 11697); 1 ♀, 1 ♂ [2 SS], Edouard,
Lac [= Edward, Lake], 10 Jul. 1952, 13 Mar. 1953, leg. unknown collector (RBINS: 17298, 17297);
1 juv ♀, 1 ♀ [2 UN], Embe, 19 Apr. 1952, 20 Apr. 1952, leg. H. de Saeger et al. (RBINS: 13654, 13653);
1 ad ♀, 1 juv ♀, 2 juv ♂♂, 1 imm ♀, 1 foetus ?, 12 ♀♀, 9 ♂♂, 2 ?? [20 SS, 1 SA, 5 AL, 3 SO], Faradje,
18 Feb. 1910, 23 Apr. 1911, 26 Apr. 1911, 29 Sep. 1911, 5 Nov. 1911, 6 Nov. 1911, 15 Nov. 1911,
1 Jan. 1912, 2 Jan. 1912, 5 Mar. 1912, 26 Mar. 1912, 14 Oct. 1912, 21 Oct. 1912, 3 Dec. 1912,
15 Dec. 1912, 29 Dec. 1912, leg. Herbert Lang, James Paul Chapin and The American Museum Congo
Expedition, 21 Oct. 1912, leg. unknown collector, 7 Jul. 1925, between 7 Jul. 1925 and 23 Oct. 1970,
leg. Henri Schouteden (AMNH: 49080 to 49082, 49384, 49079, 49090, 49382, 49383, 49083, 49084,
49087, 49088, KU: 125198, AMNH: 49071 to 49073, 49086, 49284, 49074, RMCA: 12359, AMNH:
49091, 49089, 49420, 49075 to 49078, RMCA: 9050, 9049); 2 ♀♀, 2 ♂♂ [4 SO], Gangala [= Gangalana-Bodio], 24 Mar. 1957, 25 Mar. 1957, leg. Heinz Stephan (RMCA: 27431 to 27434); 3 ♀♀, 1 ♂, 5 ??
[3 SS, 1 SO, 5 UN], Gangala-na-Bodio, unknown date, 9 Apr. 1948, 13 Apr. 1948, 11 May 1948, leg.
unknown collector, 24 Dec. 1949, leg. H. de Saeger et al., 24 Dec. 1949, leg. H. de Saeger, 6 Dec. 1956,
between 1 Nov. 1956 and 30 Nov. 1956, leg. Max Poll (RBINS: 8206, 8184, 8183, 13629, 13631,
RMCA: 36937, 25607, RBINS: 8182, RMCA: 25606); 1 ♀ [AL], Garamba National Park, between
1 Jan. 1949 and 31 Dec. 1952, leg. H. de Saeger (RMCA: 36949); 1 ♂ [UN], Goma, 9 Dec. 1952, leg.
unknown collector (RBINS: 15453); 1 ♀, 1 ♂ [1 SS, 1 SO], Ishango River, 13 Mar. 1953, 17 Mar. 1953,
leg. Gaston-François de Witte (RMCA: 37129, 37130); 1 ? [UN], Ishango-Nyakazi, unknown date, leg.
unknown collector (INPBC: V96); 1 ♂, 2 ?? [3 SS], Kabalo, unknown date, leg. Henri Schouteden
(RMCA: 9173 to 9175); 1 ? [AL], Kalina, prior to 11 May 1949, leg. Dartevelle (RMCA: 18724); 1 ♀
[SS], Kamande, 19 Dec. 1934, leg. unknown collector (RBINS: 4774); 1 ? [AL], Kambukabakali,
5 Nov. 1954, leg. Gaston-François de Witte (RMCA: 37138); 2 ?? [1 AL, 1 UN], Kanyatsi, unknown
date, leg. unknown collector, 16 Sep. 1953, leg. Gaston-François de Witte (INPBC: W541, RMCA:
37133); 2 ?? [2 UN], Kasaka, unknown date, leg. Franz Ludwig Stuhlmann (ZMB: 67896, 67897); 1 ♀,
1 ♂ [2 AL], Kasengi [= Kasengu], 23 Aug. 1926, 2 Sep. 1926, leg. James Paul Chapin, Dewitt L. Sage,
Frank P. Mathews and The Ruwenzori-Kivu Expedition (AMNH: 82386, 82385); 1 ♀ [UN], Kasenye,
10 km N, 3 Mar. 1954, leg. unknown collector (RBINS: 15454); 2 ♀♀, 1 ♂, 1 ? [1 SS, 3 UN], Kasenyi
[= Kasenye], 31 Oct. 1939, leg. Lepersonne, between 26 Feb. 1953 and 30 Dec. 1955, between
1 Oct. 1953 and 6 Oct. 1953, leg. unknown collector (RMCA: 15958, RBINS: 10545, 10543, 10544);
3 ?? [1 AL, 2 UN], Kasindi, unknown date, leg. unknown collector, unknown date, leg. H. Schubolz,
13 May 1954, leg. Gaston-François de Witte (INPBC: W978, ZMB: 67860, RMCA: 37137); 2 ♂♂
[2 SO], Kasindi-port [= Kasindi], 18 Mar. 1953, 22 Sep. 1953, leg. Gaston-François de Witte (RMCA:
37131, 37132); 2 ♀♀, 1 ♂ [2 SS, 1 SO], Katanda, 14 Sep. 1932, leg. Burgeon, 16 Mar. 1953, leg.
unknown collector, 20 Apr. 1933, leg. Henri Martin Hackars and Georges Marlier (RMCA: 12028,
RBINS: 17296, RMCA: 12164); 1 ♀, 3 ♂♂ [4 SS], Katobwe, 1 Dec. 1926, 4 Dec. 1926, 5 Dec. 1926,
leg. John Todd Zimmer (FMNH: 29404 to 29407); 1 ? [AL], Kawa (Forest), between 1 Jan. 1951 and
31 Dec. 1951, leg. Alexandre Fain (RMCA: 21461); 1 ? [AL], Kiavikire [= Kiavikere], 24 Nov. 1954,
leg. Gaston-François de Witte (RMCA: 37142); 2 ?? [2 AL], Kishero, 3 Dec. 1954, leg. Gaston-François
de Witte (RMCA: 37143, 37144); 2 ♀♀ [2 AL], Kivu Mountains, unknown date, leg. Guy Babault
176
VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
(MNHN: ZM-MO-1985-278, ZM-MO-1985-279); 2 ?? [1 AL, 1 UN], Kyavinionge, unknown date, leg.
unknown collector, 11 May 1954, leg. Gaston-François de Witte (INPBC: W973, RMCA: 37136); 3 ♀♀,
1 ♂ [4 UN], Liki, 10 Nov. 1954, 18 Nov. 1954, leg. unknown collector (RBINS: 15456.1, 15456.2,
15462.1, 15462.2); 5 ?? [2 AL, 3 UN], Lomera, unknown date, leg. unknown collector, 9 Oct. 1953,
27 Oct. 1954, leg. Gaston-François de Witte (INPBC: W1305, W1307, W583, RMCA: 37139, 37135);
1 ? [UN], Lotakamela, unknown date, leg. unknown collector (RMCA: 18958); 1 ♂ [SO], Lulenga
[= Rugari], 13 Nov. 1925, leg. Henri Schouteden (RMCA: 9182); 1 ? [UN], Mabereni River, unknown
date, leg. unknown collector (RBINS: 8205); 6 ♀♀, 9 ♂♂ [14 SS, 1 UN], Mabwe, 5 Aug. 1947,
6 Aug. 1947, 7 Aug. 1947, 12 Aug. 1947, 4 Sep. 1947, 19 Nov. 1948, 23 Dec. 1948, 6 Jan. 1949,
16 Feb. 1949, 1 Mar. 1949, between 1 Jan. 1947 and 31 Dec. 1947, leg. unknown collector (RBINS:
10701, 10695, 10696, 10699, 10700, 10697, 10702, 10804, 10691, 10692, 10698, 10693, 10703, 10694,
10690); 6 ♀♀, 3 ♂♂, 2 ?? [10 SS, 1 SO], Mahagi Port, 5 May 1925, 10 May 1925, 13 May 1925,
14 May 1925, 16 May 1925, leg. Henri Schouteden, 28 Sep. 1955, leg. James Paul Chapin, prior to
5 Jan. 1940, leg. Lepersonne (RMCA: 9005 to 9007, 9009 to 9011, AMNH: 180906, RMCA: 9003,
9008, 9060, 15959); 1 ? [AL], Maka, 27 Jun. 1947, leg. Max Poll (RMCA: 18045); 1 juv ♂, 1 juv ?,
1 ♀, 2 ?? [5 AL], Masako, 14 Sep. 1988, 27 Aug. 1989, leg. Ndey B. Ifuta (RMCA: 93.079-M-0336 to
93.079-M-0340); 2 ♀♀ [2 SS], May ya Moto, between 5 Nov. 1934 and 15 Nov. 1934, leg. GastonFrançois de Witte (RMCA: 16628, 16629); 1 ♂ [SS], Molindi River, 1 May 1934, leg. unknown
collector (RBINS: 4773); 1 ? [AL], Mongbwalu, prior to 13 Feb. 1934, leg. Jordens (RMCA: 12246);
1 ♀, 1 ? [1 SO, 1 UN], Mulongo, unknown date, leg. Gerard, 28 Oct. 1974, leg. Michael D. Gallagher
(RMCA: 10516, HZM: 26.7791); 2 ?? [1 AL, 1 UN], Museya [= Mosenda], unknown date, leg. unknown
collector, 23 Sep. 1953, leg. Gaston-François de Witte (INPBC: W546, RMCA: 37134); 2 ?? [1 SS,
1 SO], Mutwanga, between 1 Jan. 1937 and 31 Jan. 1937, leg. Henri Martin Hackars, prior to
18 Sep. 1933, leg. Van Hoof (RMCA: 16601, 12170); 1 ♀, 1 ♂ [1 AL, 1 SO], Nagbarama, 17 Apr. 1950,
leg. H. de Saeger (RMCA: 36941, 36944); 1 ♂ [SO], Nagero, 4 Jun. 1954, leg. unknown collector
(RBINS: 14626); 1 ad ? [SI], Nasosurro River, unknown date, leg. Guy Babault (MNHN: ZMMO-1939-197); 1 ? [UN], Ndote, unknown date, leg. unknown collector (INPBC: W1382); 1 ? [UN],
Ndotwe, unknown date, leg. unknown collector (INPBC: V1438); 1 ♀, 1 ♂ [2 UN], Parc National de
Garamba [= Garamba National Park], 25 Jan. 1951, 29 Jul. 1951, leg. H. de Saeger et al. (RBINS:
13644, 13648); 1 ♀, 1 ♂ [2 AL], PFSK 8, 25 Mar. 1952, leg. H. de Saeger (RMCA: 36946, 36947); 1 ♀,
1 ♂ [2 UN], PFSK.8, 25 Mar. 1952, leg. H. de Saeger et al. (RBINS: 13650.1, 13650.2); 5 ♀♀, 3 ♂♂
[8 SS], Ruwenzori SE, prior to 31 Dec. 1911, leg. Rosenberg (RMCA: 935a to 935e, 936a to 936c);
1 ♀, 1 ♂ [2 SS], Rwindi, between 1 Nov. 1934 and 30 Nov. 1934, leg. Gaston-François de Witte (RMCA:
16630, 16631); 1 ♂ [UN], Sabe, between 21 Nov. 1953 and 30 Dec. 1955, leg. unknown collector
(RBINS: 10546); 1 ♀ [SA], Semmio [= Zemio], 28 Feb. 1883, leg. Bohndorff (RMNH: MAM.35983);
1 ? [UN], Senga, unknown date, leg. unknown collector (INPBC: W551); 1 ? [AL], Stanleyville
[= Kisangani], 4 Mar. 1928, leg. Collart (RMCA: 16263); 1 ♀, 1 ? [1 AL, 1 UN], Tore, unknown date,
leg. unknown collector, 22 Mar. 1952, leg. H. de Saeger (RBINS: 13651, RMCA: 36948); 2 ?? [2 AL],
Uvira, 12 Sep. 1951, leg. Narcisse Leleup (RMCA: 22753, 22754); 2 ♀♀, 3 ♂♂ [5 SS], Vitshumbi,
1 Aug. 1934, 7 Mar. 1938, leg. unknown collector, between 1 Oct. 1933 and 31 Oct. 1933, leg. Henri
Martin Hackars (RBINS: 4775 to 4777, RMCA: 12268, 12269); 1 ♂ [AL], Volcan du Kivu [= Nyirogongo
Volcano], unknown date, leg. unknown collector (MNHN: ZM-MO-1911-1019 A); 1 ♂ [UN], Yaekela,
3 May 2010, leg. Guy-Crispin Gembu Tungaluna, Prescott Musaba and José Akaibe (UNIKIS: CRT2363).
RWANDA: 2 ♂♂ [2 SS], Colline Musha [= Musha], 13 Dec. 1954, between 1 May 1955 and 31 May
1955, leg. Alexandre Fain (MHNG: 921.031, 921.032); 2 ♂♂ [2 SA], Gabiro, 14 Jan. 1938, leg. Serge
Frechkop (RMCA: 17085, 17086); 1 ? [SS], Gakoma (Mission), 12 Aug. 1949, leg. Aureliaan (RMCA:
18997); 3 ♀♀ [3 UN], Kagera National Park [= Akagera Park], 15 Jan. 1938, 19 Jan. 1938, leg.
unknown collector (RBINS: 4779, 4780, 4778); 1 ad ♀, 1 juv ? [1 SA, 1 AL], Karama, 11 Oct. 1981,
leg. Frits De Vree, Benny Baeten and Victor Van Cakenberghe (RMCA: 82.006-M-0928, 82.006-M177
European Journal of Taxonomy 382: 1–327 (2017)
0931); 1 ♀ [SA], Katjumbura, 10 Jun. 1938, leg. Serge Frechkop (RMCA: 17087); 1 ♂ [SA], Kibuye,
1 Nov. 1980, leg. Danny Meirte (RMCA: 80.055-M-0171); 1 ♀ [SS], Kimuna, 1 Dec. 1966, leg.
unknown collector (RBINS: 16004); 1 ? [UN], Kissenji [= Gisenyi], unknown date, leg. StegmannStein (ZMB: 67844).
Rhinolophidae Gray, 1825
Rhinolophus Lacépède, 1799
BURUNDI: 1 ? [AL], Rushubi Mine, 12 Nov. 1969, leg. P.L. Niort (MHNG: 1895.080).
DEMOCRATIC REPUBLIC OF THE CONGO: 2 ?? [2 AL], unknown locality, 23 Aug. 2012, leg.
Prescott Musaba (RMCA: a1.097-M-3214, a1.097-M-3215); 1 ? [AL], Boende, between 1 Jan. 1970
and 31 Jan. 1970, leg. P. Lootens (RMCA: 38819); 2 juv ?? [2 UN], Djomba [= Djemba], unknown
date, leg. unknown collector (INPBC: V1523.1, V1523.2); 1 juv ?, 3 ?? [4 UN], Garamba National
Park, between 1 Jan. 1948 and 31 Dec. 1948, leg. unknown collector (RBINS: 8129.1 to 8129.4); 1 juv
♀ [UN], Goma, between 26 Nov. 1952 and 30 Dec. 1955, leg. unknown collector (RBINS: 10548);
4 juv ♀♀, 7 juv ♂♂ [10 SS, 1 SO], Kafwe, 14 Jul. 1947, leg. unknown collector (RBINS: 10743
to 10752, 10772); 5 juv ♀♀ [3 SS, 2 SO], Kaswabilenga, 9 Oct. 1947, 4 Nov. 1947, leg. unknown
collector (RBINS: 10758, 10756, 10757, 10769, 10770); 4 juv ♂♂ [4 SS], Kibanga, 17 Nov. 1947, leg.
unknown collector (RBINS: 10759, 10760, 10849, 10850); 1 ♂ [AL], Kona, 12 May 2010, leg. GuyCrispin Gembu Tungaluna (RMCA: a1.097-M-3216); 1 ♂ [UN], Lieki, 1 Jun. 2010, leg. Guy-Crispin
Gembu Tungaluna, Prescott Musaba and José Akaibe (UNIKIS: CRT2038); 2 ?? [2 SS], Lovanium
University [= Kinshasa], 11 Jan. 1963, 6 Jul. 1963, leg. De Bont and Jo Van Orshoven (RMCA: 32632,
32631); 1 ♂ [MS], Lukobwe, 12 Oct. 1948, leg. unknown collector (RBINS: 1099); 8 juv ♀♀, 2 juv
♂♂ [10 SS], Masombwe, 10 Oct. 1948, 12 Oct. 1948, leg. unknown collector (RBINS: 10779, 10782,
10783, 10786, 10787, 10790, 10791, 10773, 10775, 10776); 1 ? [AL], Moenge, 15 May 2010, leg. GuyCrispin Gembu Tungaluna (RMCA: a1.097-M-3222); 1 ? [AL], Niambasha, 1 Aug. 1956, leg. Uwe
Rahm (RMCA: 33534); 1 juv ♂ [SS], Pelenge, Gorges, 1 Jun. 1947, leg. unknown collector (RBINS:
10736); 1 ♂ [UN], Yaekela, 4 May 2010, leg. Guy-Crispin Gembu Tungaluna, Prescott Musaba and
José Akaibe (UNIKIS: CRT2392).
Rhinolophus alcyone Temminck, 1853
DEMOCRATIC REPUBLIC OF THE CONGO: 1 ♂ [SK], Buluku River, 12 Jul. 1951, leg. H. de
Saeger et al. (RBINS: 13667); 1 ad ♂ [SS], Lukolela, 1 Aug. 1930, leg. James Paul Chapin (AMNH:
86880); 1 ♂ [SS], Makpe River, upper [= Makpe], 22 May 1952, leg. H. de Saeger et al. (RBINS:
13666); 1 ♂ [UN], Scierie Forest, 22 Nov. 1974, leg. Michael D. Gallagher (HZM: 1.8022).
Rhinolophus blasii Peters, 1867
DEMOCRATIC REPUBLIC OF THE CONGO: 1 ♂ [SA], Katanga (former) Province, 31 May 1962,
leg. Félix / Michel Anciaux de Faveaux (SMF: 21245); 1 ad ♀ [SA], Kivu region, unknown date, leg.
Guy Babault (MNHN: ZM-MO-1985-939); 1 ? [AL], Moba, 23 Apr. 1954, leg. H. Bomans (RMCA:
26280).
Rhinolophus clivosus Cretzschmar, 1828
BURUNDI: 1 ? [SB], Nyagatarugwa River Valley, 9 Aug. 1992, leg. L. Davenport (FMNH: 148366);
1 ♂ [SB], Nyamugari Hill, 10 Sep. 1990, leg. L. Davenport (FMNH: 148367).
178
VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
DEMOCRATIC REPUBLIC OF THE CONGO: 1 ♀ [AL], Defrenne Cave [= Mwanga Cave],
21 Jun. 1959, leg. Félix / Michel Anciaux de Faveaux (RMCA: 28212); 8 ?? [8 UN], Djomba
[= Djemba], unknown date, leg. unknown collector (INPBC: V1846.1 to V1846.8); 2 ♀♀ [2 AL],
Jadotville [= Likasi], 3 Apr. 1957, leg. Félix / Michel Anciaux de Faveaux (RMCA: 26256, 26257);
21 ♂♂ [21 AL], Jomba, 4 Sep. 1976, 5 Sep. 1976, 6 Sep. 1976, leg. Frieder Meier (MHNG: 1873.020
to 1873.040); 1 ♀ [SK], Kakanda Cave, 22 Sep. 1960, leg. unknown collector (RBINS: 14513); 1 ?
[SK], Kisantu, 11 Dec. 1920, leg. Henri Schouteden (RMCA: 7718); 1 sad ♀, 3 ?? [4 AL], Lubudi,
small Cave [= Lubudi], 1 Nov. 1948, leg. Narcisse Leleup (RMCA: 22667 to 22670); 2 ♂♂ [2 AL],
Lubuka Cave, 28 Aug. 1950, leg. Laurent (RMCA: 79.010-M-0005, 79.010-M-0006); 9 ?? [9 UN],
Musanze-Djomba, unknown date, leg. unknown collector (INPBC: V1760, V1764.1 to V1764.8); 1 ?
[SS], Mutwanga, between 1 Jan. 1937 and 31 Jan. 1937, leg. Henri Martin Hackars (RMCA: 16602);
1 ? [UN], Rutshuru, unknown date, leg. unknown collector (INPBC: W1163); 1 ♀ [AL], Swanepoel
I Cave, 1 May 1958, leg. Félix / Michel Anciaux de Faveaux (RMCA: 27538); 13 ?? [13 AL], Uvira,
12 Sep. 1951, leg. Narcisse Leleup (RMCA: 22740 to 22752).
RWANDA: 2 ♀♀, 1 ♂ [3 SS], Bugoye Kisenyi, between 1 Nov. 1933 and 30 Nov. 1933, leg. Henri
Martin Hackars (RMCA: 12265 to 12267); 1 ? [UN], Gisenyi, 11 Dec. 2004, leg. unknown collector
(SMF: 92962); 1 ? [SK], Kibumbu Crater, 9 Dec. 1991, leg. J.R. Wilson and Anthony Michael
Hutson (BMNH: 1992.390); 31 ad ♀♀, 49 ad ♂♂ [72 SA, 8 AL], Kidaho, 9 Jun. 1982, 10 Jun. 1982,
11 Jun. 1982, 12 Jun. 1982, 13 Jun. 1982, 16 Jun. 1982, 17 Jun. 1982, 18 Jun. 1982, leg. Benny Baeten
(RMCA: 87.046-M-0137 to 87.046-M-0214, 87.046-M-0135, 87.046-M-0136); 3 ad ♀♀, 6 ad ♂♂
[9 SK], Musanse [= Musanze], 7 Jan. 1966, 10 Jun. 1966, leg. unknown collector (RBINS: 15480.1
to 15480.3, 15479.1 to 15479.6); 111 ad ♀♀, 98 ad ♂♂, 2 sad ♀♀, 20 juv ♀♀, 18 juv ♂♂, 3 ?? [2 SS,
114 SA, 136 AL], Mutura, unknown date, 14 Dec. 1984, 15 Dec. 1984, 16 Dec. 1984, 18 Dec. 1984,
12 Jan. 1985, 13 Jan. 1985, leg. Benny Baeten and Luc Janssens, 22 Oct. 1981, 23 Oct. 1981,
25 Oct. 1981, 26 Oct. 1981, 27 Oct. 1981, leg. Frits De Vree, Benny Baeten and Victor Van Cakenberghe
(RMCA: 85.006-M-0537, 85.006-M-0538, 85.006-M-0076 to 85.006-M-0094, 85.006-M-0001 to
85.006-M-0037, 85.006-M-0465, 85.006-M-0038 to 85.006-M-0075, 82.006-M-0046 to 82.006-M0200); 1 ♂ [SA], Nyundo, 17 Oct. 1980, leg. Danny Meirte (RMCA: 80.055-M-0170); 2 ♂♂, 3 ??
[2 AL, 3 UN], Ruhengeri, 20 Sep. 1976, leg. Frieder Meier, 22 Jun. 2003, 25 Jun. 2003, 3 Jul. 2003,
leg. unknown collector (MHNG: 1873.041, 1873.042, SMF: 92438 to 92440); 1 ? [SS], Ubuvumo lava
hole, 11 Dec. 2004, leg. Michael Laumanns (SMF: 92961).
Rhinolophus eloquens K. Andersen, 1905
DEMOCRATIC REPUBLIC OF THE CONGO: 1 ♀ [SS], unknown locality, 1 Apr. 1927, leg. David
H. Linder (MCZ: 23377); 2 ♀♀, 1 ♂ [3 SS], Aba, 15 Dec. 1911, 16 Dec. 1911, 17 Dec. 1911, leg. Herbert
Lang, James Paul Chapin and The American Museum Congo Expedition (AMNH: 49100 to 49102);
1 ♂, 1 ? [1 AL, 1 UN], Bunia, unknown date, leg. Maristes, between 1 Apr. 1927 and 30 Apr. 1927, leg.
Harold Jefferson Coolidge (RMCA: 16277, MCZ: 28545); 1 ? [AL], Katanga, prior to 31 Aug. 1899,
leg. Charles Lemaire (RMCA: 387); 2 ♂♂ [2 SS], Kilo Mines [= Kilomines], 21 Apr. 1949, leg. Jean
Hiernaux (RMCA: 19095, 19096); 1 ? [AL], Kiseny [= Kasenye], between 1 Jan. 1926 and 31 Dec. 1926,
leg. Henri Schouteden (RMCA: 16276); 1 ♂ [AL], Luofu, 23 Mar. 1927, leg. James Paul Chapin, Dewitt
L. Sage, Frank P. Mathews and The Ruwenzori-Kivu Expedition (AMNH: 82393); 1 ♀, 2 ♂♂ [1 SS,
2 AL], Mboga [= Boga], unknown date, leg. Robert William Hayman, unknown date, leg. Howard de
Walden (BMNH: 1930.11.11.102 to 1930.11.11.104); 4 ♀♀ [1 SS, 1 SA, 2 AL], Rutshuru, 1 Apr. 1927,
leg. David H. Linder, 30 Jul. 1925, 23 Aug. 1925, leg. Edmund Heller, 26 Apr. 1926, leg. James Paul
Chapin, Dewitt L. Sage, Frank P. Mathews and The Ruwenzori-Kivu Expedition (MCZ: 25122, FMNH:
26500, AMNH: 82392, FMNH: 30716); 1 ♂ [SS], Vankerckhovenville [= Nzoro], unknown date, leg.
The Alexander-Gosling Expedition (BMNH: 1907.7.8.30).
179
European Journal of Taxonomy 382: 1–327 (2017)
RWANDA: 1 ad ? [SA], Kidaho, 8 Jun. 1982, leg. Benny Baeten (RMCA: 87.046-M-0349); 1 ♀ [SA],
Rwankwi, between 1 Apr. 1948 and 30 Apr. 1948, leg. J.V. Leroy (RMCA: 19069); 1 ♂, 1 ? [1 AL,
1 UN], Shangugu, between 1 Jan. 1926 and 31 Dec. 1926, between 1 Jan. 1926 and 31 Jan. 1926, leg.
Henri Schouteden (RMCA: 16218, 16275).
Rhinolophus fumigatus Rüppell, 1842
BURUNDI: 17 ♂♂ [17 AL], Karonge, 3 Oct. 1969, 12 Oct. 1969, leg. P.L. Niort (MHNG: 1895.072
to 1895.079, 1895.086, 1895.048 to 1895.050, 1895.052 to 1895.056); 1 ♂ [AL], Mpinga Cave
[= Mpinga Mission], 31 Dec. 1968, leg. P.L. Niort (MHNG: 1895.051); 2 ?? [2 AL], Rushubi Mine,
16 Nov. 1969, leg. P.L. Niort (MHNG: 1895.087, 1895.088); 1 ♂ [AL], Rwintare, 3 Apr. 1969, leg.
P.L. Niort (MHNG: 1895.047).
DEMOCRATIC REPUBLIC OF THE CONGO: 1 ad ♀, 8 ♀♀, 2 ♂♂ [10 SS, 1 SO], Aba, 15 Dec. 1911,
16 Dec. 1911, 17 Dec. 1911, leg. Herbert Lang, James Paul Chapin and The American Museum
Congo Expedition, 15 Dec. 1911, leg. unknown collector (AMNH: 49111, 49112, 49113 [holotype
Rhinolophus abæ J.A. Allen, 1917], 49114 to 49117, MCZ: 17375, RMCA: 12363, AMNH: 49118,
49197); 1 ? [UN], Albert National Park [= Virunga National Park], unknown date, leg. unknown
collector (INPBC: V264.2); 1 ? [AL], Bunia, prior to 20 Feb. 1939, leg. Maristes (RMCA: 15225);
1 ♀ [UN], Gangala-na-Bodio, 7 Nov. 1949, leg. H. de Saeger et al. (RBINS: 13657); 59 ♀♀,
164 ♂♂ [223 UN], Goroumbwa Mine, 17 May 1999, 18 May 1999, 19 May 1999, 19 Oct. 1999,
20 Oct. 1999, 21 Oct. 1999, 22 Oct. 1999, 23 Oct. 1999, 24 Oct. 1999, 25 Oct. 1999, 27 Oct. 1999, leg.
Robert (Bob) Swanepoel (TM: DRC0032, DRC0053, DRC0055, DRC0056, DRC0086, DRC0102 to
DRC0104, DRC0120, DRC0127 to DRC0131, DRC0134, DRC0135, DRC0142, DRC0146, DRC0147,
DRC0149, DRC0151, DRC0154, DRC0155, DRC0158, DRC0159, DBA008, DBA012 to DBA014,
DBA030 to DBA063, DBA085 to DBA106, DBA129, DBA130, DBA132 to DBA141, DBA143
to DBA156, DBA158 to DBA163, DBA178, DBA180 to DBA184, DBA195, DBA196, DBA201,
DBA203 to DBA206, DBA208 to DBA223, DBA228, DBA245 to DBA253, DBA256 to DBA260,
DBA262 to DBA265, DBA267, DBA269 to DBA273, DBA275, DBA276, DBA278, DBA283,
DBA290 to DBA299, DBA301 to DBA316, DBA318, DBA319, DBA322 to DBA326, DBA328 to
DBA330, DBA354, DBA360, DBA401 to DBA406, DBA408, DBA409, DBA411, DBA412); 2 ??
[2 AL], Ibambi, prior to 24 Jun. 1939, leg. Reitter (RMCA: 15388, 15389); 5 ♀♀, 4 ♂♂ [8 AL, 1 UN],
Kakanda Cave, 22 Apr. 1959, leg. Félix / Michel Anciaux de Faveaux (RMCA: 28169, 28171 to
28174, 28176, 28179, 28181, 28184); 3 ?? [3 AL], Lukatu Cave [= Ntoto Cave], prior to 2 Feb. 2007,
leg. A. Kimbembi-ma-Ibaka (RMCA: a7.006-M-0002 to a7.006-M-0004); 1 ? [AL], Mani Grot of
Nkela Grot van Mbumba-Nsi [= Mani Cave or Nkela Cave or Mbumba-Nsi], between 1 Jan. 2002
and 13 Jul. 2002, leg. A. Kimbembi-ma-Ibaka (RMCA: a2.040-M-0002); 1 ♂ [SK], Musosa, near,
between 1 Jun. 1940 and 30 Jun. 1940, leg. unknown collector (RBINS: 14489); 4 ♀♀, 1 ♂ [5 AL],
Mwadingusha, 18 Dec. 1957, leg. Félix / Michel Anciaux de Faveaux (RMCA: 27532 to 27536); 1 juv
♀, 2 ♀♀, 2 ♂♂, 7 ?? [1 SS, 1 AL, 6 SK, 1 SO, 3 UN], Ndelele, 3 Mar. 1952, 8 Mar. 1952, 11 Mar. 1952,
19 Nov. 1952, leg. H. de Saeger et al., 11 Mar. 1952, 19 Jun. 1952, leg. H. de Saeger (RBINS: 13659.1,
13659.2, 13661, RMCA: 36831, 36830, RBINS: 13662.1 to 13662.4, 13660.1, 13660.2, 13656); 1 ?
[AL], Rungu, between 1 Aug. 1925 and 31 Aug. 1925, leg. Henri Schouteden (RMCA: 16250); 2 ??
[2 UN], Thysville [= Mbanza-Ngungu], 11 Feb. 1987, leg. unknown collector (RBINS: 22428, 22429).
RWANDA: 1 ? [UN], Bilalu, unknown date, leg. unknown collector (INPBC: V3063); 1 ? [SO], Gabiro,
prior to 22 Jan. 1935, leg. Verhulst (RMCA: 12664); 22 ?? [22 UN], Lugarama, unknown date, leg.
unknown collector (INPBC: 3041.01 to 3041.12, V304.01 to V304.10); 9 ?? [9 UN], Magachi, unknown
date, leg. unknown collector (INPBC: V1903.1 to V1903.9); 1 ♂ [AL], Ruhengeri, 15 Nov. 1980, leg.
Hans H. de Iongh (ZMA: MAM.21229).
180
VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
Rhinolophus hildebrandtii Peters, 1878
BURUNDI: 1 ♂ [AL], Bujumbura, between 1 Jul. 1968 and 31 Jul. 1968, leg. Lewall (RMCA: 35153);
3 ♂♂ [3 AL], Kanyinya, 6 Jun. 1969, leg. P.L. Niort (MHNG: 1895.041, 1895.042, 1895.044); 3 ♀♀
[3 AL], Makamba, 10 Apr. 1969, between 23 Mar. 1970 and 24 Mar. 1970, leg. P.L. Niort (MHNG:
1895.046, 1895.081, 1895.082); 5 ♂♂ [5 AL], Mpinga Cave [= Mpinga Mission], 1 Jan. 1969,
25 Mar. 1970, leg. P.L. Niort (MHNG: 1895.043, 1895.045, 1895.083 to 1895.085); 1 ? [AL], Ruyigi,
between 19 Mar. 1952 and 22 Mar. 1952, leg. Laurent (RMCA: 79.010-M-0009).
DEMOCRATIC REPUBLIC OF THE CONGO: 1 ♀ [SK], unknown locality, prior to 1 May 1939, leg.
unknown collector (RBINS: 6282); 1 ♀, 1 ♂ [2 SA], Baga Cave [= Baya Cave], 12 Mar. 1956, leg.
Félix / Michel Anciaux de Faveaux (HNHM: 95.52.1, 95.52.2); 7 ?? [7 AL], Baya Carrière [= Baya
Quarry], 17 May 1965, leg. Walter N. Verheyen, 26 Jun. 1963, leg. Parent (RMCA: 34055 to 34059,
31505, 31506); 1 ♂, 2 ?? [3 AL], Baya Cave [= Baya Quarry], unknown date, leg. Félix / Michel
Anciaux de Faveaux (MNHN: ZM-MO-1957-130 B, ZM-MO-1957-130 C, ZM-MO-1957-130 D);
1 ♂ [AL], Defrenne Cave [= Mwanga Cave], 21 Jun. 1959, leg. Félix / Michel Anciaux de Faveaux
(RMCA: 28211); 4 ♀♀ [4 SA], Dethiox Caves [= Kambove], 4 Jun. 1960, leg. Félix / Michel Anciaux de
Faveaux (SMF: 18914, 18915, 18917, 18918); 10 ♀♀, 9 ♂♂ [19 SK], Elisabethville [= Lubumbashi],
25 Mar. 1966, leg. unknown collector (RBINS: 17205.01 to 17205.19); 1 juv ?, 1 ? [2 UN], Gangalana-Bodio, prior to 23 Nov. 1957, leg. Station de la chasse et domestication des éléphants (RMCA:
26664, 26665); 8 ♀♀, 2 ♂♂, 1 ? [4 SS, 6 AL, 1 SO], Grotte de Baya [= Baya Cave], 12 Mar. 1956,
18 Mar. 1956, 14 May 1956, leg. Félix / Michel Anciaux de Faveaux, 14 May 1956, 12 Jul. 1956,
13 Jul. 1956, leg. M. Lips (MHNG: 922.093, 922.094, RMCA: 23801, 23802, 23805, MHNG: 922.090,
922.091, RMCA: 23804, 23806, MHNG: 922.087, 922.092); 1 ♂ [AL], Ibembo, 25 Jan. 1951, leg. Jozef
Hutsebaut (RMCA: 89.004-M-0002); 1 ♂ [AL], Jadotville [= Likasi], 31 Aug. 1958, leg. Félix / Michel
Anciaux de Faveaux (RMCA: 27905); 1 ♂, 1 ? [1 AL, 1 UN], Kaboyaboya Cave, unknown date, leg.
unknown collector, 23 Jul. 1962, leg. Félix / Michel Anciaux de Faveaux (MHNG: 1046.001, 1046.100);
16 ♀♀, 2 ♂♂, 2 ?? [12 AL, 6 SK, 2 UN], Kakanda Cave, 21 Sep. 1960, 29 Sep. 1960, 21 Jul. 1962, leg.
unknown collector, 22 Apr. 1959, leg. Félix / Michel Anciaux de Faveaux, between 21 Apr. 1955 and
22 Apr. 1955, leg. R.P. Th. de Caters (RBINS: 14448, 14451, 14428, 14436 to 14438, 14441, RMCA:
28156 to 28165, RBINS: 14433, RMCA: 22889, 22897); 1 ad ♂, 4 ♀♀, 7 ♂♂, 8 ?? [1 SS, 2 SA, 15 AL,
2 UN], Kakontwe Cave, unknown date, 3 Jul. 1956, 4 Jul. 1956, 9 Jul. 1956, 13 Feb. 1957, 8 Jul. 1957,
12 Jul. 1957, leg. Félix / Michel Anciaux de Faveaux, 5 Jul. 1948, leg. Narcisse Leleup, 6 Feb. 1957,
leg. G. Verhoustraete, prior to 24 Sep. 1962, leg. L. Poelman (MNHN: ZM-MO-1957-130 A, MHNG:
1046.066a, 1046.066b, RMCA: 26742, 26743, 26851, 26250, 23866, 23816, 22687, 26215, 26781,
26833, 26233, 31209 to 31214); 1 ♂ [AL], Kalumbu Cave, 1 Jul. 1959, leg. Félix / Michel Anciaux
de Faveaux (RMCA: 28318); 5 ?? [5 UN], Kando Cave, unknown date, leg. unknown collector
(RMCA: 22062 to 22066); 1 ♂ [AL], Kandu (Cave), 25 Jul. 1957, leg. Félix / Michel Anciaux de
Faveaux (RMCA: 26876); 2 ♂♂ [2 SS], Kipangaribwe River, 3 Jan. 1948, 17 Jan. 1948, leg. unknown
collector (RBINS: 10852, 10851); 1 ? [AL], Kiwakishi Cave [= Kiamokoto], between 1 Jul. 1958 and
31 Jul. 1958, leg. Narcisse Leleup (RMCA: 27292); 1 ♀, 1 ♂ [1 SA, 1 UN], Kondo Cave [= Kakanda
Cave], 21 Sep. 1960, leg. Félix / Michel Anciaux de Faveaux (SMF: 19571, 19572); 10 ♀♀, 7 ♂♂
[4 SA, 10 AL, 3 UN], Kyasala Cave [= Kiasala Cave], 31 Mar. 1956, 28 Jul. 1957, 21 Jul. 1962, leg.
Félix / Michel Anciaux de Faveaux, 21 Jul. 1962, leg. unknown collector (MHNG: 1046.099, RBINS:
14447, 14450, SMF: 21273, 21274, MHNG: 1046.067, 1046.068, RMCA: 26784, 26791, 26795, 26824,
26825, 26829, 26835, 26867, 23810, 23853); 3 ♀♀ [3 SA], Kyasala Grande Grotte [= Kiasala Cave],
28 Jul. 1957, leg. Félix / Michel Anciaux de Faveaux (MHNG: 1046.069 to 1046.071); 5 ?? [5 AL],
Lubudi, small Cave [= Lubudi], 1 Nov. 1948, leg. Narcisse Leleup (RMCA: 22662 to 22666); 1 ?
[UN], Marungu, unknown date, leg. unknown collector (NMZL: 3396); 48 ?? [48 AL], M’baya Cave
[= Baya Cave], 25 Mar. 1966, leg. Xavier Misonne (RMCA: 34571 to 34618); 1 ♀ [SK], Musosa, near,
181
European Journal of Taxonomy 382: 1–327 (2017)
between 1 Jun. 1943 and 30 Jun. 1943, leg. unknown collector (RBINS: 14432); 2 ♀♀, 1 ♂ [3 AL],
Mwanga Cave, 28 Jun. 1964, leg. Félix / Michel Anciaux de Faveaux (RMCA: 33999 to 34001); 1 juv
♀, 4 ♀♀, 7 ♂♂ [1 SA, 5 SK, 6 UN], Mwela-Pande Cave [= Tenke], 11 Nov. 1959, leg. Félix / Michel
Anciaux de Faveaux, 5 Dec. 1959, 6 Dec. 1959, leg. unknown collector (SMF: 19068, RBINS: 14449,
14427, 14430, 14431, 14435, 14439, 14440, 14443 to 14446); 1 ♀, 2 ♂♂ [2 SA, 1 AL], Pempere
Cave, 19 Jun. 1957, 11 Jun. 1958, leg. Félix / Michel Anciaux de Faveaux (SMF: 18940, 18941,
MHNG: 1046.065); 18 ♀♀, 8 ♂♂ [16 SA, 10 AL], Pierkat Quarry [= Baya Quarry], 12 Mar. 1956,
18 Mar. 1956, 14 May 1956, leg. Félix / Michel Anciaux de Faveaux (RMCA: 23807, 23808, 23827 to
23850); 1 ♂ [AL], Swanepoel II Abyss, 1 May 1958, leg. Félix / Michel Anciaux de Faveaux (RMCA:
27539); 6 ♀♀, 1 ? [1 AL, 2 SK, 4 UN], Tantara Cave [= Shinkolobwe], unknown date, 4 Jun. 1960,
leg. unknown collector, 11 Nov. 1957, leg. Félix / Michel Anciaux de Faveaux (RBINS: 14442, RMCA:
27531, RBINS: 14424 to 14426, 14429, 14434).
Rhinolophus hilli Aellen, 1973
RWANDA: 1 ad ♀ [SA], Routabansougera, 19 Oct. 1981, leg. Frits De Vree, Benny Baeten and Victor
Van Cakenberghe (RMCA: 82.006-M-0001); 1 ad ♀ [SA], Uwinka, 25 Aug. 1964, leg. U. Goepel
(ZMZ: 126639 [holotype Rhinolophus hilli Aellen, 1973]).
Rhinolophus kahuzi Fahr & Kerbis Peterhans, 2013
DEMOCRATIC REPUBLIC OF THE CONGO: 1 ad ♂ [SA], Mount Kahuzi, W slope, 22 Jul. 2007,
leg. R. Kizungu (FMNH: 219793 [holotype Rhinolophus kahuzi Fahr & Kerbis Peterhans, 2013]).
Rhinolophus landeri Martin, 1838
BURUNDI: 1 ♀, 1 ♂ [1 SA, 1 SK], Resha, 16 Mar. 1979, 30 Mar. 1979, leg. Heinz Stephan (SMF:
57523, 57524).
DEMOCRATIC REPUBLIC OF THE CONGO: 1 ad ♀, 1 ♀, 3 ♂♂ [5 SS], Aba, 13 Dec. 1911,
15 Dec. 1911, 17 Dec. 1911, leg. Herbert Lang, James Paul Chapin and The American Museum Congo
Expedition (AMNH: 49132, 49173, 49175 [holotype Rhinolophus axillaris Allen, 1917], 49177,
RMCA: 12361); 2 ♂♂ [1 AL, 1 SK], Aka, 15 May 1952, leg. H. de Saeger et al., 15 May 1952,
leg. H. de Saeger (RBINS: 13664, RMCA: 36832); 2 ?? [2 AL], Banana, between 1 Jan. 1952 and
31 Jan. 1952, leg. Is. Mesmaekers (RMCA: 20746, 20747); 11 ?? [5 AL, 6 UN], Banana - Boma
Road, km 74, 25 km from coast in Forest, 16 Dec. 1951, leg. Is. Mesmaekers (RMCA: 20643 to
20653); 2 ♀♀ [2 AL], Defrenne Cave [= Mwanga Cave], 21 Jun. 1959, leg. Félix / Michel Anciaux
de Faveaux (RMCA: 28213, 28214); 1 ♀, 1 ♂ [2 SK], Gandajika, 1 Oct. 1948, leg. unknown collector
(RBINS: 14505, 14506); 1 ♂ [UN], Goroumbwa Mine, 19 May 1999, leg. Robert (Bob) Swanepoel
(TM: DRC0141); 2 ♀♀, 1 ♂ [3 AL], Jadotville [= Likasi], 8 Jul. 1957, leg. Félix / Michel Anciaux de
Faveaux (RMCA: 26722 to 26724); 1 ? [AL], Kabinda, between 1 Jan. 1945 and 31 Dec. 1946, leg.
Hautmann (RMCA: 18149); 2 ♀♀, 1 ♂, 2 ?? [1 SA, 2 AL, 1 SK, 1 UN], Kakanda Cave, 21 Sep. 1960,
leg. unknown collector, 21 Sep. 1960, leg. Félix / Michel Anciaux de Faveaux, between 21 Apr. 1955
and 22 Apr. 1955, leg. R.P. Th. de Caters (RBINS: 14504, SMF: 19576, 19577, RMCA: 22886, 22887);
1 ? [AL], Kakanda Mutaka, between 6 Feb. 1954 and 15 Feb. 1954, leg. R.P. Th. de Caters (RMCA:
22380); 3 ?? [3 SA], Kakondo, 13 Aug. 1963, 10 Sep. 1963, leg. P. Kunkel (SMF: 31825, 31828,
31827); 2 ?? [2 AL], Kakontwe, 15 Aug. 1948, leg. Narcisse Leleup (RMCA: 22652, 22653); 5 ♀♀,
9 ♂♂ [11 AL, 3 UN], Kakontwe Cave, 24 Jun. 1956, 4 Jul. 1956, 8 Jul. 1957, 12 Jul. 1957, between
3 Feb. 1956 and 6 Jul. 1956, leg. Félix / Michel Anciaux de Faveaux (RMCA: 26747 to 26753, 26756,
26773, 23860, 23867, 26877, 26878, 23868); 1 ♀, 1 ♂ [2 AL], Kalumbu Cave, 1 Jul. 1959, leg. Félix
/ Michel Anciaux de Faveaux (RMCA: 28319, 28320); 1 ? [AL], Kaniama, prior to 14 Sep. 1954, leg.
182
VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
Vachaudez (RMCA: 22370); 2 ♂♂ [2 SK], Kanzenze Mission, 1 Jun. 1948, leg. unknown collector
(RBINS: 15455.1, 15455.2); 1 ? [AL], Katanga, unknown date, leg. Félix / Michel Anciaux de Faveaux
(MNHN: ZM-MO-1957-131); 2 ♀♀ [2 SA], Katanga (former) Province, 18 Dec. 1957, leg. Félix /
Michel Anciaux de Faveaux (SMF: 18938, 18939); 1 ? [AL], Kikango, 16 Jan. 1954, leg. H. Bomans
(RMCA: 22127); 10 ?? [10 AL], Kisangani, between 1 Jun. 1982 and 30 Jun. 1982, leg. Marc Colyn
(RMCA: 82.031-M-0023 to 82.031-M-0032); 1 ♀, 7 ♂♂ [8 AL], Kyasala Cave [= Kiasala Cave],
15 Apr. 1956, 28 Jul. 1957, leg. Félix / Michel Anciaux de Faveaux (RMCA: 23813, 26785, 26802,
26812, 26827, 26843, 26844, 26866); 1 ? [AL], Lake Kivu, near, between 1 Jan. 1934 and 31 Dec. 1937,
leg. Guy Babault (MCZ: 46096); 1 ? [AL], Lubondai-Tshimbulu [= Lubondaie-Tshimbulu], between
1 Jan. 1955 and 31 Dec. 1955, leg. S. Marks (RMCA: 26279); 1 ♀ [SS], Luluabourg [= Kananga],
unknown date, leg. Richard Callewaert (BMNH: 1926.7.6.111); 11 ♀♀, 9 ♂♂ [18 AL, 2 UN], Lusolo
Cave [= Lubudi], 28 Jul. 1957, leg. Félix / Michel Anciaux de Faveaux (RMCA: 26793, 26794, 26811,
26814, 26815, 26819, 26821, 26822, 26828, 26830, 26831, 26836 to 26840, 26852, 26853, 26868,
26869); 2 ♂♂, 1 ? [2 SA, 1 AL], Lwiro [= Luiro], 13 Jul. 1956, leg. Alvin Novick, 28 Oct. 1965, leg.
P. Kunkel (MCZ: 48215, 48216, SMF: 31826); 1 ♂ [SK], Maleli, 13 May 1951, leg. H. de Saeger et al.
(RBINS: 13663); 2 ♀♀, 4 ♂♂ [5 AL, 1 UN], Mulonga Cave, 30 Jul. 1957, 1 May 1959, leg. Félix /
Michel Anciaux de Faveaux (RMCA: 28203, 28204, 26783, 26786, 26800, 26817); 2 ♀♀, 7 ♂♂ [3 SK,
6 UN], Mulungwishi, 27 Dec. 1959, leg. unknown collector (RBINS: 14495 to 14503); 2 ♀♀, 1 ♂
[2 AL, 1 UN], Mulungwishi, Cave, 28 Jun. 1956, leg. Félix / Michel Anciaux de Faveaux (RMCA:
23815, 23865, 26231); 1 ad+foetus ♀, 7 ?? [8 AL], Mwanakusu Cave, 23 Aug. 1959, 28 Aug. 1959,
leg. Segers and Benoit Mission, 24 Oct. 1954, leg. Narcisse Leleup (RMCA: 29207, 22724 to 22729,
29208); 2 juv ??, 2 ♀♀, 1 ♂ [2 AL, 1 SK, 2 UN], Ndelele, 11 Mar. 1952, leg. H. de Saeger, 31 Jul. 1952,
leg. H. de Saeger et al. (RMCA: 36828a, 36828b, 36829a, 36829b, RBINS: 13665); 1 ? [AL], Pawa,
unknown date, leg. Reitter (RMCA: 15727); 1 ♂ [SS], Pelenge, Gorges, 4 Jun. 1947, leg. Gaston-François
de Witte (RBINS: 10740); 1 ♀ [SA], Sankuru Province, between 1 Jan. 1952 and 31 Dec. 1952, leg.
Félix / Michel Anciaux de Faveaux (MHNG: 1047.080); 1 ♀ [AL], Swanepoel II Abyss, 1 May 1958,
leg. Félix / Michel Anciaux de Faveaux (RMCA: 27540); 1 ♀ [AL], Tantara Cave [= Shinkolobwe],
29 Sep. 1957, leg. Félix / Michel Anciaux de Faveaux (RMCA: 26806); 1 ? [AL], Tshakala, between
1 Jan. 1982 and 31 Jan. 1982, leg. Marc Colyn (RMCA: 82.019-M-0012).
RWANDA: 1 ? [UN], Nyakichozi, unknown date, leg. unknown collector (INPBC: V3118.2).
Rhinolophus ruwenzorii J. Eric Hill, 1942
DEMOCRATIC REPUBLIC OF THE CONGO: 1 ad ♀ [SA], Butahu Valley, Cave, 24 Dec. 1926, leg.
James Paul Chapin, Dewitt L. Sage, Frank P. Mathews and The Ruwenzori-Kivu Expedition (AMNH:
82394 [holotype Rhinolophus ruwenzorii J.Eric Hill, 1942]); 1 ♂ [SS], Butembo, 16 Aug. 1947,
leg. Jean Hiernaux (RBINS: 7047); 4 ♀♀, 3 ♂♂, 3 ?? [7 SA, 3 AL], Kibwe Lya Mikako Cave, 28
Dec. 1966, leg. M.J. Celis and M. Lejeune (RMCA: 35170, 35173, 35174, 35206, 35208, 35211, 35216
to 35219); 1 ♀, 3 ♂♂ [4 SA], Komanda, 21 Aug. 1976, leg. Frieder Meier (MHNG: 1873.011 to
1873.014); 1 ♂ [SK], Matata Cave, 10 Aug. 1947, leg. Jean Hiernaux (RBINS: 7048); 3 ♂♂ [2 SS,
1 SA], Matupi Cave, between 14 Sep. 1959 and 16 Sep. 1959, leg. A.E. Wright and The Cambridge
Africa Expedition 1959 (BMNH: 1960.100, 1960.101, 1960.99); 1 ♀ [SK], Mount Hoyo [= Hoyo,
Mount], 10 Aug. 1947, leg. Jean Hiernaux (RBINS: 7049); 1 ♀ [SA], Paoni, 6 Aug. 1976, leg. Frieder
Meier (MHNG: 1873.010).
RWANDA: 2 ad ♂♂ [2 SA], Mutura, 16 Dec. 1982, leg. Benny Baeten and Luc Janssens (RMCA:
85.006-M-0447, 85.006-M-0448).
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European Journal of Taxonomy 382: 1–327 (2017)
Rhinolophus simulator K. Andersen, 1904
DEMOCRATIC REPUBLIC OF THE CONGO: 2 ♀♀, 18 ?? [20 AL], Baya Carrière [= Baya Quarry],
17 May 1965, between 1 Dec. 1962 and 31 Dec. 1962, leg. Walter N. Verheyen (RMCA: 34044 to
34054, 35022, 35028, 35042, 35043, 34060 to 34064); 1 ♀, 1 ♂ [2 SA], Defrenne Cave [= Mwanga
Cave], 7 Aug. 1960, leg. Félix / Michel Anciaux de Faveaux (SMF: 19080, 19081); 2 ♀♀, 4 ♂♂ [6 SA],
Kaboyaboya Cave, 18 Jul. 1962, 23 Jul. 1962, leg. Félix / Michel Anciaux de Faveaux (SMF: 21253
to 21258); 1 ♂ [SA], Kalomoni Cave [= Kakontwe], 11 Jun. 1960, leg. Félix / Michel Anciaux de
Faveaux (SMF: 19079); 4 ♀♀, 4 ♂♂ [8 SA], Kasoma Cave, 13 Aug. 1962, leg. Félix / Michel Anciaux
de Faveaux (SMF: 21265 to 21272); 1 ♂ [SA], Lubudi, 23 Jul. 1962, leg. Félix / Michel Anciaux de
Faveaux (SMF: 21259); 5 ♀♀, 1 ♂ [6 SA], Lusolo Cave [= Lubudi], 16 Jul. 1962, leg. Félix / Michel
Anciaux de Faveaux (SMF: 21247 to 21252); 3 ♀♀, 2 ♂♂ [5 SA], Mulong Cave [= Mulonga Cave],
26 Jul. 1962, leg. Félix / Michel Anciaux de Faveaux (SMF: 21260 to 21264); 2 ♀♀ [2 AL], Mwanga
Cave, 28 Jun. 1964, leg. Félix / Michel Anciaux de Faveaux (RMCA: 33997, 33998); 1 ♂ [SA], Trou
aux Serpents [= Shinkolobwe], 30 Jun. 1962, leg. Félix / Michel Anciaux de Faveaux (SMF: 21246).
Rhinolophus swinnyi Gough, 1908
DEMOCRATIC REPUBLIC OF THE CONGO: 1 ad ♂, 1 ♂ [1 SA, 1 AL], Dethioux Hoeve, Grot I
[= Shinkolobwe], 29 Sep. 1957, leg. Félix / Michel Anciaux de Faveaux (MHNG: 1046.063, 1046.064);
1 ♀, 1 ♂ [2 UN], Jadotville [= Likasi], 26 Jul. 1962, leg. unknown collector (RBINS: 14478, 14479);
2 ♀♀, 5 ♂♂ [4 AL, 3 SK], Kaboyaboya Cave, 23 Jul. 1962, leg. Félix / Michel Anciaux de Faveaux,
23 Jul. 1962, leg. unknown collector (MHNG: 1047.059 to 1047.062, RBINS: 14481, 14482, 14485);
5 ♀♀ [2 SA, 3 AL], Kakontwe Cave, 8 Jul. 1957, 12 Jul. 1957, leg. Félix / Michel Anciaux de Faveaux
(MHNG: 1046.048, 1046.049, 1046.045 to 1046.047); 1 ? [UN], Kando Cave, unknown date, leg.
unknown collector (MHNG: 1046.005); 1 ♂ [AL], Kandu (Cave), 25 Jul. 1957, leg. Félix / Michel
Anciaux de Faveaux (MHNG: 1046.050); 1 ♀, 1 ♂ [2 SA], Kasoma Cave, 13 Aug. 1962, leg. Félix /
Michel Anciaux de Faveaux (HNHM: 95.50.1, 95.50.2); 1 ♀ [AL], Kyamakonde Cave, 18 Jul. 1962,
leg. Félix / Michel Anciaux de Faveaux (MHNG: 1047.058); 1 ♂ [AL], Kyantapo Cave, 29 Jul. 1957,
leg. Félix / Michel Anciaux de Faveaux (MHNG: 1046.059); 1 ♀ [SA], Kyasala Cave [= Kiasala
Cave], 15 Apr. 1956, leg. Félix / Michel Anciaux de Faveaux (MHNG: 922.086); 2 ♀♀ [2 AL], Kyasala
Grande Grotte [= Kiasala Cave], 28 Jul. 1957, leg. Félix / Michel Anciaux de Faveaux (MHNG:
1046.051, 1046.052); 6 ♀♀, 15 ♂♂ [2 SA, 10 AL, 3 SK, 6 UN], Lusolo Cave [= Lubudi], 28 Jul. 1957,
16 Jul. 1962, leg. Félix / Michel Anciaux de Faveaux, 16 Jul. 1962, leg. unknown collector (MHNG:
1047.052 to 1047.057, RBINS: 14473 to 14475, 14477, 14480, 14483, 14484, 14487, 14488, MHNG:
1046.053 to 1046.058); 1 ♀, 2 ♂♂, 1 ? [3 AL, 1 UN], Mulonga Cave, 30 Jul. 1957, leg. Félix / Michel
Anciaux de Faveaux (MHNG: 1046.006, 1046.060 to 1046.062); 1 ♀ [UN], Mulungwishi, 27 Dec. 1959,
leg. unknown collector (RBINS: 14476); 1 ♀ [SA], Pempere Cave, 19 Jun. 1957, leg. Félix / Michel
Anciaux de Faveaux (MHNG: 1046.044); 2 ♀♀ [1 SK, 1 UN], Salomoni Cave, 11 Jun. 1960, 20 May
1962, leg. unknown collector (RBINS: 14472, 14486).
Rhinolophus willardi Kerbis Peterhans & Fahr, 2013
DEMOCRATIC REPUBLIC OF THE CONGO: 3 ad ♀♀ [3 SB], Kabobo Forest, 14 Feb. 2007, leg.
B.D. Marks, 16 Feb. 2007, leg. Prince Kiswele Kaleme (FMNH: 195082 [paratype Rhinolophus willardi
Kerbis Peterhans and Fahr, 2013], 195083 [paratype Rhinolophus willardi Kerbis Peterhans and
Fahr, 2013], 195084 [paratype Rhinolophus willardi Kerbis Peterhans and Fahr, 2013]); 1 ad ♂
[SA], Misotschi-Kabogo Highlands, 28 Feb. 2007, leg. A.J. Plumptre and E.A. Mulungu (FMNH:
195182 [holotype Rhinolophus willardi Kerbis Peterhans and Fahr, 2013]).
184
VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
Rhinonycteridae Gray, 1866
Cloeotis Thomas, 1901
Cloeotis percivali Thomas, 1901
DEMOCRATIC REPUBLIC OF THE CONGO: 1 ad ♀, 1 ad ♂ [2 AL], Dethioux Farm, Cave I
[= Shinkolobwe], 15 Jun. 1958, leg. Félix / Michel Anciaux de Faveaux (RMCA: 27907, 27908); 1 ♀,
1 ♂ [2 SA], Dethiox Caves [= Kambove], 4 Jun. 1960, leg. Félix / Michel Anciaux de Faveaux (SMF:
18920, 18921); 1 ♂ [UN], Elisabethville [= Lubumbashi], 25 Mar. 1966, leg. unknown collector
(RBINS: 17208); 1 ♂ [AL], Kaboyaboya Cave, 23 Jul. 1962, leg. Félix / Michel Anciaux de Faveaux
(MHNG: 1046.098); 1 ♂ [SA], Kasoma Cave, 13 Aug. 1962, leg. Félix / Michel Anciaux de Faveaux
(SMF: 21275); 1 ad ♀, 1 ad ♂ [2 SS], Kiamokoto, 4 Oct. 1948, leg. unknown collector (RBINS: 10792,
10793); 1 ♀, 1 ♂ [1 SA, 1 AL], Lubudi, 23 Jul. 1962, leg. Félix / Michel Anciaux de Faveaux (SMF:
21276, 21277); 5 ad ♀♀, 1 ♂, 2 ?? [5 SS, 1 SK, 2 UN], Masombwe, unknown date, 10 Oct. 1948,
19 Oct. 1948, leg. unknown collector (RBINS: 10802, 10803, 10777, 10789, 10794, 10796, 10797,
10800).
Vespertilioniformi Hutcheon & Kirsch, 2006
Emballonuridae Gervais, 1855
Coleura Peters, 1868
Coleura afra (Peters, 1852)
DEMOCRATIC REPUBLIC OF THE CONGO: 5 ♀♀, 4 ♂♂, 7 ?? [4 SS, 7 SK, 5 SO], Aba, 16 Dec. 1911,
leg. Herbert Lang, James Paul Chapin and The American Museum Congo Expedition, 16 Dec. 1911, leg.
unknown collector (AMNH: 48809, 48811 to 48815, 49033, 51250 to 51256, MCZ: 17369, RMCA:
12394); 3 ♀♀, 1 ♂, 4 ?? [8 AL], Cape Cassa Mabongo [= Kasambongo], unknown date, leg. the
Belgian Hydrobiological Expedition to Tanganyika (BMNH: 1953.10 to 1953.12, 1953.5 to 1953.9);
6 ♀♀, 3 ♂♂ [9 UN], Tanganyika, Congo shore of Lake, between 1 Jan. 1946 and 31 Dec. 1947, leg.
unknown collector (RBINS: 9062.1 to 9062.9); 1 ♀, 1 ♂, 3 ?? [4 SS, 1 AL], Wago, Mount, 7 Nov. 1951,
between 1 Nov. 1951 and 30 Nov. 1951, between 1 Jul. 1951 and 31 Jul. 1951, between 1 Jan. 1951 and
31 Dec. 1951, leg. Alexandre Fain (RMCA: 23282, 21466, 23291, 23292, 23283).
Saccolaimus Temminck, 1838
Saccolaimus peli (Temminck, 1853)
DEMOCRATIC REPUBLIC OF THE CONGO: 3 ♀♀, 1 ♂, 2 ?? [5 SS, 1 AL], Avakubi, 18 Nov. 1909,
17 Jan. 1914, 15 Jan. 1915, leg. Herbert Lang, James Paul Chapin and The American Museum Congo
Expedition, 16 Jan. 1914, leg. J. Bequaert, between 17 Jan. 1914 and 17 Dec. 1914, leg. unknown
collector (RMCA: 12377, 3420, AMNH: 48772, 49350, 48757, MCZ: 17366); 1 ♂ [SS], Bafwaboka
[= Bafwabaka], 28 Dec. 1909, leg. Herbert Lang, James Paul Chapin and The American Museum
Congo Expedition (AMNH: 48761); 1 ? [SO], Beni, prior to 24 Jul. 1939, leg. Lisfranc (RMCA: 15404);
1 ? [AL], Beno, prior to 3 Jan. 1950, leg. P. Bayet (RMCA: 19472); 1 juv ? [AL], Bokoro, prior to
19 Nov. 1952, leg. E. Jans (RMCA: 21521); 1 ? [UN], Djeka, prior to 15 Jun. 1956, leg. Roiseux (RMCA:
23640); 2 ♂♂ [2 SO], Ikela, between 1 Jan. 1946 and 31 Dec. 1946, leg. L. Dupuis (RMCA: 17907,
17908); 1 ♀ [SS], Inkongo [= Inkongo-Kakese], unknown date, leg. H. Wilson (BMNH: 1916.5.15.5);
2 ?? [2 AL], Irumu, 28 Sep. 1926, leg. James Paul Chapin, Dewitt L. Sage, Frank P. Mathews and The
Ruwenzori-Kivu Expedition (AMNH: 82363, 82364); 2 ?? [1 SS, 1 SO], Kikwit, between 1 Nov. 1922
and 30 Nov. 1922, leg. J. Rodhain (RMCA: 14913, 14914); 1 ad ♀ [SS], Kinawa Passage, 26 Sep. 1953,
leg. unknown collector (RBINS: 21460); 1 ♀ [SS], Kisiki, 22 Dec. 1938, leg. Duncan M. Hodgson and
William F. Coultas (MCZ: 38027); 1 ♀ [SS], Kunungu, prior to 25 Mar. 1929, leg. Ngwe (RMCA: 9708);
185
European Journal of Taxonomy 382: 1–327 (2017)
2 ?? [2 SS], Lesse, 4 Nov. 1913, prior to 2 Jun. 1914, leg. Bonnevie (RMCA: 3241, 3242); 1 embryo ?,
4 ♀♀ [1 SS, 2 SA, 2 AL], Lukolela, 28 Jul. 1930, 3 Aug. 1930, 4 Sep. 1930, leg. James Paul Chapin,
4 Sep. 1930, leg. Franklin Edson (AMNH: 86904, 86902, 86768, 86903, 86915); 1 ♀ [SS], Luluabourg
[= Kananga], unknown date, leg. Richard Callewaert (BMNH: 1926.7.6.98); 2 ♀♀, 2 ♂♂ [1 SS, 3 UN],
Medje, 14 Jan. 1910, 15 Jan. 1910, 18 Jan. 1910, 19 Jan. 1910, leg. Herbert Lang, James Paul Chapin
and The American Museum Congo Expedition (AMNH: 48762, 48763, RMCA: 12378, AMNH: 48765);
2 ♂♂ [1 AL, 1 SK], Moera’s Village, 15 Jun. 1924, leg. Edmund Heller (FMNH: 30709, 30710); 2 ♂♂
[2 SK], Musenge, 23 Mar. 1924, 24 Mar. 1924, leg. Edmund Heller (FMNH: 30707, 30708); 2 ♀♀, 1 ♂,
2 ?? [2 SS, 1 SA, 2 AL], N’Gayu area, 18 Dec. 1909, leg. Herbert Lang, James Paul Chapin and The
American Museum Congo Expedition (AMNH: 48758 to 48760, 49281, 49282); 3 ♀♀, 2 ♂♂ [5 SS],
Niangara, 27 May 1913, 31 May 1913, leg. Herbert Lang, James Paul Chapin and The American Museum
Congo Expedition, 31 May 1913, leg. unknown collector (AMNH: 48766, 48768, 48769, 48767, MCZ:
17365); 1 ? [AL], Omaniumdu, 27 Jul. 1959, leg. Laurent (RMCA: 29535); 1 ? [AL], Panga, between
1 Sep. 1925 and 30 Sep. 1925, leg. Henri Schouteden (RMCA: 16258); 3 ?? [2 AL, 1 UN], Pawa, prior
to 23 Jan. 1939, leg. Degotte (RMCA: 14990 to 14992); 1 ♀ [SS], Rungu, 28 Jul. 1913, leg. Herbert
Lang, James Paul Chapin and The American Museum Congo Expedition (AMNH: 48770); 1 ? [SO],
St. Jos. (Misson) [= Mikalaya], unknown date, leg. Richard Callewaert (AMNH: 86261); 1 ♂ [SS],
St. Joseph de Luluabourg Mission [= Mikalaya], 18 Jul. 1922, leg. Richard Callewaert (RMCA: 7541);
2 ?? [2 AL], Sturi Forest [= Ituri Forest], unknown date, leg. The Ruwenzori Expedition (BMNH:
1907.1.2.10, 1907.1.2.9); 2 ♂♂ [2 SS], Upper Congo, unknown date, leg. Alexander Douglas Mitchell
Carruthers (BMNH: 1907.6.14.1, 1907.6.14.2); 1 ♂ [AL], Walikale, 28 Feb. 1924, leg. Edmund Heller
(USNM: 260135); 1 juv ♀, 1 ♀, 2 ♂♂ [3 SS, 1 AL], Yalosemba, 17 Jun. 1979, 28 Jun. 1979, leg.
C. Brian Robbins, 30 Jun. 1979, leg. Lynn W. Robbins (USNM: 537702, 463476, 537701, TTU: 36355);
1 ? [SO], Yokamba Elongo, 27 Jan. 1952, leg. Stevenart (RMCA: 21129).
Taphozous E. Geoffroy St.-Hilaire, 1818
Taphozous mauritianus E. Geoffroy St.-Hilaire, 1818
BURUNDI: 1 ad ♀, 1 ♀ [2 SK], Bujumbura, 7 km NW, 21 Mar. 1979, leg. Heinz Stephan (SMF:
57544, 57545); 1 ? [AL], Russissi, bank [= Rusizi Delta], 27 Apr. 1982, leg. Danny Meirte (RMCA:
82.016-M-0148).
DEMOCRATIC REPUBLIC OF THE CONGO: 1 ♂, 2 ?? [1 SS, 2 AL], unknown locality, unknown
date, leg. unknown collector, prior to 4 Jul. 1958, leg. E. Jans (BMNH: 1906.7.1.18, RMCA: 27373,
27374); 1 ? [UN], Aba, 12 Apr. 1948, leg. unknown collector (RBINS: 8188); 2 ♀♀, 1 ♂, 1 ? [4 AL],
Avakubi, 3 Nov. 1911, 10 Dec. 1913, 10 Dec. 1914, leg. Herbert Lang, James Paul Chapin and The
American Museum Congo Expedition (AMNH: 49349, MCZ: 17227, AMNH: 49367, 49366); 1 ad ♀,
4 ad ♂♂, 1 juv ♀, 1 juv ♂, 7 ?? [7 SS, 1 SA, 1 AL, 4 SO, 1 SN], Bambesa, 23 Nov. 1939, 22 Apr. 1940,
24 Aug. 1940, 27 Aug. 1940, 29 Apr. 1941, 2 May 1941, 14 May 1941, between 1 Jun. 1937 and 30
Jun. 1937, leg. J.M. Vrijdagh (RMCA: 18104, 17768, 17763, 17765, 16106 to 16108, 17749, 17750,
17764, 17751, 14072, 14074, 14079); 5 ?? [5 AL], Banana, between 1 May 1952 and 31 May 1952, leg.
Is. Mesmaekers, prior to 28 Feb. 1899, leg. Weyns (RMCA: 21372, 21373, 148, 148a, 148b); 1 ? [SS],
Baudouinville [= Moba], 5 May 1948, leg. Jozef Hutsebaut (RMCA: 18567); 1 ♂, 1 ? [2 SS], Baye via
Bondo, 13 Dec. 1956, leg. Rooyakkers (RMCA: 25861, 25862); 1 ? [AL], Beno, prior to 3 Jan. 1950,
leg. P. Bayet (RMCA: 19473); 1 ? [AL], Binga, prior to 27 Feb. 1935, leg. Van den Put (RMCA: 12678);
1 ? [SS], Blukwa, 9 Jul. 1952, leg. unknown collector (RBINS: 21664); 1 ? [AL], Boende, between
1 Feb. 1970 and 28 Feb. 1970, leg. P. Lootens (RMCA: 75.035-M-0020); 4 ?? [4 AL], Bokuma, 30
Jan. 1952, prior to 5 Mar. 1953, prior to 3 Feb. 1955, leg. P. Lootens (RMCA: 21396, 23246, 23247,
21682); 2 ♀♀, 5 ?? [5 SS, 2 SO], Bokungu, 14 Jun. 1949, 19 Jan. 1952, 28 Apr. 1952, prior to 22
Oct. 1949, prior to 13 Oct. 1950, leg. L. Dupuis (RMCA: 18845, 20847, 20848, 20846, 20084, 18929,
186
VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
18930); 1 ♀ [SS], Bolobo, prior to 23 Jul. 1946, leg. N’Kele (RMCA: 17402); 1 ♀ [UN], Bomane
[= Bomaneh], 22 May 2010, leg. Guy-Crispin Gembu Tungaluna, Prescott Musaba and José Akaibe
(UNIKIS: CRT2227); 1 ♂, 12 ?? [6 SS, 5 AL, 2 SO], Buta, unknown date, 12 Sep. 1939, 12 Mar. 1945,
20 Jun. 1949, prior to 31 May 1939, prior to 29 Mar. 1940, prior to 28 Dec. 1938, prior to 24 Jan. 1949,
prior to 16 Jun. 1937, prior to 1 Apr. 1932, between 1 Dec. 1927 and 31 Dec. 1927, leg. Jozef Hutsebaut,
29 Jan. 1925, leg. Henri Schouteden (RMCA: 14565, 16282, 17303, 15777, 18842, 8929, MCZ: 23934,
RMCA: 11590, 13855, 19079, 14947, 15954, 15324); 1 ? [AL], Butembo, prior to 26 May 1959, leg.
P. Dyleff (RMCA: 27885); 3 ♀♀, 2 ♂♂ [2 SS, 2 AL, 1 UN], Cel II, 12 Dec. 1951, leg. H. de Saeger
et al., 14 Mar. 1951, 26 Nov. 1951, 12 Dec. 1951, leg. H. de Saeger (RBINS: 13533, RMCA: 36841,
36840, 36838, 36839); 2 ♀♀, 1 ♂, 1 ? [1 SS, 3 AL], Elisabethville [= Lubumbashi], 10 Mar. 1956, leg.
Félix / Michel Anciaux de Faveaux, 14 Feb. 1956, leg. service de l’hygiene publique, 14 Apr. 1930, leg.
J. De Riemaecker (MHNG: 922.100, RMCA: 23824, 26281, 19417); 2 juv ♀♀, 2 juv ♂♂, 1 juv ?,
22 ♀♀, 17 ♂♂, 1 ? [33 SS, 9 AL, 3 SO], Faradje, 6 Feb. 1911, 1 Apr. 1911, 11 Apr. 1911, 17 May 1911,
20 May 1911, 28 May 1911, 7 Jun. 1911, 11 Jul. 1911, 5 Sep. 1911, 26 Oct. 1911, 22 Nov. 1911,
24 Nov. 1911, 6 Mar. 1912, 21 May 1912, 22 Oct. 1912, 25 Oct. 1912, 12 Nov. 1912, 13 Nov. 1912,
28 Nov. 1912, 3 Dec. 1912, 5 Dec. 1912, 7 Dec. 1912, 15 Dec. 1912, 16 Dec. 1912, 5 Jan. 1913,
5 Oct. 1913, 14 Oct. 1913, 5 Nov. 1913, between 5 Jan. 1913 and 5 Dec. 1913, leg. Herbert Lang, James
Paul Chapin and The American Museum Congo Expedition (AMNH: 48786, 48787, 48798, 48779,
48793, 48806, 48794, 49355, 48783, 48796, 48808, MCZ: 17367, AMNH: 48788, 48774, 48775, 48789,
48797, MCZ: 17368, AMNH: 49362 to 49364, 48804, 48801 to 48803, 48776 to 48778, 48805, 48800,
48790, 48795, 48780, 49360, 48799, 49353, 49361, 48781, 48782, RMCA: 12383, AMNH: 49352,
48791, RMCA: 12384, AMNH: 48807, MCZ: 17228); 1 ♀, 1 ♂ [2 UN], Fataki, between 1 Jan. 1948
and 6 Apr. 1964, leg. unknown collector (RBINS: 18530.1, 18530.2); 2 ♀♀, 1 ♂ [3 SO], Gangala
[= Gangala-na-Bodio], 24 Mar. 1957, 27 Mar. 1957, leg. Heinz Stephan (RMCA: 27461, 27463,
27462); 1 juv ?, 1 neonate ♀, 1 neonate ♂, 3 neonate ??, 3 ?? [2 SS, 1 AL, 1 SO, 5 UN], Gangala-naBodio, unknown date, 2 Jan. 1947, 9 Apr. 1948, 5 May 1948, leg. unknown collector, 14 Nov. 1956,
between 1 Nov. 1956 and 30 Nov. 1956, leg. Max Poll (RBINS: 12942, RMCA: 25601, RBINS: 12941,
8190, 11655, 8189.1, 8189.2, RMCA: 25600, 25806); 1 ? [AL], Ibambi, 11 Nov. 1938, leg. Degotte
(RMCA: 15215); 7 ?? [7 AL], Ibembo, 20 Sep. 1949, prior to 17 Jul. 1952, prior to 15 Dec. 1950, leg.
Jozef Hutsebaut (RMCA: 19462, 20213 to 20217, 21277); 2 ?? [2 AL], Ikela, prior to 4 Jan. 1958, leg.
P. Lootens (RMCA: 27018, 27019); 1 ♂ [SS], Inkongo [= Inkongo-Kakese], 2 Feb. 1923, leg. H. Wilson
(RMCA: 6312); 1 ♀ [SS], Kabwe, 12 Nov. 1973, leg. Chris E. Heidenreich (WNMU: 2944); 2 ??
[2 AL], Kakanda Cave, between 21 Apr. 1954 and 22 Apr. 1954, leg. R.P. Th. de Caters (RMCA: 22898,
22899); 2 ♀♀, 3 ♂♂ [4 SS, 1 SO], Kamituga, 2 Dec. 1950, 11 Jul. 1958, 16 Jul. 1964, 23 Aug. 1966,
11 Jul. 1974, leg. Alexandre Prigogine (RMCA: 27729, 76.022-M-0024, 32800, 20466, 34803); 1 ♀
[AL], Kapolowe, 16 Apr. 1956, leg. Félix / Michel Anciaux de Faveaux (RMCA: 23826); 1 ♂ [AL],
Kapolowe Mission [= Kapolowe], 16 Apr. 1956, leg. Félix / Michel Anciaux de Faveaux (RMCA:
23919); 1 ♀ [AL], Kasengi [= Kasengu], 22 Aug. 1926, leg. James Paul Chapin, Dewitt L. Sage, Frank
P. Mathews and The Ruwenzori-Kivu Expedition (AMNH: 82382); 1 ♂ [AL], Katana, unknown date,
leg. unknown collector (MNHN: ZM-MO-1985-273); 2 ♀♀, 4 ♂♂ [6 SS], Katobwe, 28 Nov. 1926,
1 Dec. 1926, 7 Dec. 1926, leg. John Todd Zimmer (FMNH: 29411, 29412, 29410, 29413 to 29415);
4 ♀♀, 1 ♂ [2 SS, 3 SO], Kilwezi, 9 Aug. 1948, 20 Aug. 1948, leg. unknown collector (RBINS: 10857,
10853 to 10856); 1 ♂ [SS], Koteli [= Kotili], 23 Jan. 1925, leg. Henri Schouteden (RMCA: 8920); 1 ♂
[SS], Kunungu, prior to 22 Oct. 1930, leg. N’Kele (RMCA: 10617); 2 ?? [2 SS], Leverville [= Lusanga],
between 1 Feb. 1929 and 28 Feb. 1929, leg. Tinant (RMCA: 9990, 9991); 1 ? [AL], LubondaiTshimbulu [= Lubondaie-Tshimbulu], prior to 31 Jan. 1958, leg. S. Marks (RMCA: 26928); 1 ? [AL],
Lubumbashi, unknown date, leg. J. Bequaert (RMCA: 7699); 1 ? [AL], Lubutu, 16 Sep. 1985, leg.
Marc Colyn (RMCA: 86.024-M-0005); 1 ♀ [SS], Lulenga [= Rugari], 28 Nov. 1925, leg. Henri
Schouteden (RMCA: 9153); 4 ♀♀, 1 ♂, 1 ? [4 SS, 1 SA, 1 AL], Luluabourg [= Kananga], unknown
date, leg. Richard Callewaert, 16 Nov. 1965, leg. Antoon Emeric Marcel De Roo, between 1 Jan. 1932
187
European Journal of Taxonomy 382: 1–327 (2017)
and 31 Dec. 1932, leg. Jean Romieux, prior to 17 Apr. 1913, leg. Cambier (BMNH: 1926.11.1.24,
1926.11.1.25, 1926.7.6.97, RMCA: 33981, MHNG: 850.045, RMCA: 1549); 1 ? [SS], Luputa, prior to
22 Jun. 1933, leg. Bouvier (RMCA: 12131); 1 ♂ [SO], Lusambo, unknown date, leg. J. Ghesquière
(RMCA: 7333); 1 ? [SO], Macaco, 21 Sep. 1921, leg. Henri Schouteden (RMCA: 6553); 1 ad ♀ [SS],
Mahagi, 20 Apr. 1942, leg. J.M. Vrijdagh (RMCA: 17747); 1 ? [AL], Makaw, prior to 15 Nov. 1958,
leg. E. Jans (RMCA: 27498); 1 ? [AL], Makengo, 11 Dec. 1946, leg. Dartevelle (RMCA: 18162); 1 ?
[AL], Masako, unknown date, leg. Ndey B. Ifuta (RMCA: 93.079-M-0355); 1 ad ♂ [SO], Mondombe,
1 Jan. 1953, leg. L. Dupuis (RMCA: 21716); 1 ♀ [UN], Mpaza, 22 Jan. 1952, leg. H. de Saeger et al.
(RBINS: 13536); 1 ♀ [SO], Murambi, 23 May 1949, leg. unknown collector (RBINS: 12940); 3 ♀♀,
1 ♂, 1 ? [3 SS, 1 AL, 1 SO], Mutsora, 30 Dec. 1946, 20 Feb. 1947, 16 Jun. 1951, 11 Jul. 1952, leg.
unknown collector, 2 Aug. 1954, leg. Gaston-François de Witte (RBINS: 17294, 12938, RMCA: 37162,
RBINS: 12939, 12937); 1 ♂ [AL], Nagera [= Nagero], 26 Mar. 1952, leg. H. de Saeger (RMCA:
36842); 3 ♂♂ [2 SO, 1 UN], Nagero, 1 Dec. 1953, 31 Dec. 1953, leg. unknown collector, 27 Dec. 1951,
leg. H. de Saeger et al. (RBINS: 14627, 13534, 14628); 2 ♀♀ [1 SS, 1 SO], Niangara, 18 Nov. 1910,
25 Nov. 1910, leg. Herbert Lang, James Paul Chapin and The American Museum Congo Expedition
(AMNH: 48773, 48784); 1 juv ? [AL], Niarembe, between 1 Jun. 1925 and 30 Jun. 1925, leg. Henri
Schouteden (RMCA: 16251); 1 ? [AL], Nyangwe, 6 Apr. 1920, leg. J. Ghesquière (RMCA: 16247); 1 ♂
[SS], Pania-Mutombo [= Pania-Mutombo], 20 Jun. 1934, leg. R.S. Pierrepont (MCZ: 31736); 2 ♀♀,
1 ♂ [3 UN], Parc National de Garamba [= Garamba National Park], 23 Jul. 1951, 8 May 1952, leg.
H. de Saeger et al. (RBINS: 13531.1, 13531.2, 13537); 9 ♀♀, 7 ♂♂, 2 ?? [12 SS, 2 AL, 1 SK, 1 SO,
2 UN], Rutshuru, unknown date, 10 Mar. 1938, leg. unknown collector, 31 Jul. 1925, 1 Aug. 1925,
3 Aug. 1925, leg. Edmund Heller, 3 Jan. 1938, 20 Jan. 1938, 3 Mar. 1938, 4 Mar. 1938, 5 Mar. 1938,
24 Mar. 1938, between 1 Mar. 1938 and 31 Mar. 1938, between 1 Feb. 1938 and 28 Feb. 1938, leg.
J. Ghesquière, between 1 Mar. 1938 and 31 Mar. 1938, leg. Serge Frechkop (INPBC: V1347, V375,
FMNH: 26498, RBINS: 4770, RMCA: 14542, 14607, 14608, 14444, 14595, FMNH: 30725, 30726,
30724, RMCA: 14593, 14594, 14592, 14596, 14606, 16745); 1 ♂ [SS], St. Jos. (Mission) [= Mikalaya],
29 Jul. 1924, leg. Richard Callewaert (AMNH: 86171); 2 ♀♀, 1 ♂ [1 SS, 2 SO], St. Joseph de
Luluabourg Mission [= Mikalaya], 2 May 1939, leg. Jean Jacques Deheyn, 6 Jun. 1922, leg. Richard
Callewaert (RMCA: 15371, 15372, 7551); 2 ♀♀, 1 ♂, 1 ? [2 SS, 1 AL, 1 SO], Stanleyville [= Kisangani],
5 Jul. 1912, leg. Cuthbert Christy, between 1 Mar. 1929 and 31 Mar. 1929, leg. Collart, between
1 Apr. 1924 and 30 Apr. 1924, leg. J. Ghesquière (RMCA: 1430, 16246, 9215, 9216); 1 juv ♂, 1 ♀,
2 ♂♂, 1 ? [4 SS, 1 SO], Tshikapa, 25 Oct. 1921, 20 Nov. 1921, leg. Henri Schouteden (RMCA: 6549
to 6552, 6554); 1 ♂ [SS], Upper Congo, unknown date, leg. Alexander Douglas Mitchell Carruthers
(BMNH: 1907.6.14.3); 9 ?? [9 AL], Wafanya [= Wafania], unknown date, 17 Sep. 1991, 20 Mar. 1992,
15 Apr. 1992, between 1 Jan. 1992 and 31 Jan. 1992, leg. P. Lootens (RMCA: 92.079-M-0119 to 92.079M-0122, 92.079-M-0151, 92.079-M-0002, 91.076-M-0086, 92.079-M-0001, 92.079-M-0003); 1 ?
[AL], Yakuluku, 3 Nov. 1911, leg. Herbert Lang, James Paul Chapin and The American Museum Congo
Expedition (AMNH: 49365); 1 ♀, 2 ♂♂, 1 ? [1 SS, 1 SK, 2 UN], Yangambi, 13 Mar. 1947, 24 Feb. 1948,
leg. unknown collector, 5 Aug. 1942, leg. J.M. Vrijdagh (RBINS: 14464, 14466, 6986, RMCA: 17748);
1 ? [AL], Yaosuka, 5 Jul. 1949, leg. R.P. Van Boven (RMCA: 79.012-M-0001); 3 ♀♀, 2 ♂♂ [5 UN],
Zobia, prior to 18 Apr. 1946, leg. unknown collector (RBINS: 14463, 14465, 14467 to 14469).
RWANDA: 1 ? [SA], Astrida [= Butare], between 1 Jan. 1948 and 31 Dec. 1948, leg. Alexandre Fain
(RMCA: 18992); 3 ?? [3 SS], Gabiro, 16 Jan. 1938, leg. unknown collector, 16 Jan. 1938, leg. Serge
Frechkop (RBINS: 4769, RMCA: 17074, 17075); 1 ? [UN], Kagera National Park [= Akagera Park],
prior to 31 Dec. 1943, leg. unknown collector (RBINS: 4771).
188
VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
Taphozous nudiventris Cretzschmar, 1830
DEMOCRATIC REPUBLIC OF THE CONGO: 1 ♀ [UN], Cel II, 8 Mar. 1952, leg. H. de Saeger et al.
(RBINS: 13542).
Taphozous perforatus E. Geoffroy St.-Hilaire, 1818
DEMOCRATIC REPUBLIC OF THE CONGO: 5 ?? [5 AL], unknown locality, prior to 14 Sep. 1957,
leg. Félix / Michel Anciaux de Faveaux (RMCA: 27919 to 27923); 3 ♀♀ [3 UN], Bamangwa
[= Bamungwa [River]], 23 Feb. 1951, leg. H. de Saeger et al. (RBINS: 13539.1 to 13539.3); 2 ♀♀
[2 AL], Bamongwa [= Bamungwa [River]], 23 Feb. 1951, leg. H. de Saeger (RMCA: 36843, 36844);
1 ♀ [UN], Biadimbi, 21 Feb. 1951, leg. H. de Saeger et al. (RBINS: 13538); 4 ♀♀, 2 ♂♂, 15 ?? [2 SS,
17 AL, 2 UN], Defrenne Cave [= Mwanga Cave], 19 Jun. 1957, 31 Aug. 1958, 25 Aug. 1962, leg.
Félix / Michel Anciaux de Faveaux, 19 Sep. 1948, leg. Narcisse Leleup (RMCA: 26707 to 26710,
22487, 22488, 22602 to 22614, MHNG: 1046.096, RMCA: 27898); 6 ♀♀, 2 ♂♂, 4 ?? [3 SS, 7 AL,
2 SK], Dungu, 1 Mar. 1913, leg. Herbert Lang, James Paul Chapin and The American Museum Congo
Expedition, 1 May 1913, leg. Herbert Lang (AMNH: 48816, 48817, 49251, 49348, 49354, 49356 to 49358,
MCZ: 17225, RMCA: 12395, FMNH: 43874, 43875); 1 ♂ [SS], Gangala-na-Bodio, 8 Apr. 1945, leg.
unknown collector (RBINS: 14631); 1 ? [AL], Kasongo, 25 Jul. 1959, leg. Segers and Benoit Mission
(RMCA: 29209); 5 ♂♂ [5 AL], Kasowena Cave, 28 Jun. 1964, leg. Félix / Michel Anciaux de Faveaux
(RMCA: 34002 to 34006); 2 ♀♀, 2 ♂♂, 3 ?? [1 SS, 1 AL, 1 SK, 1 SO, 3 UN], Ndelele, 3 Mar. 1952,
11 Mar. 1952, leg. H. de Saeger et al., 2 Mar. 1952, 11 Mar. 1952, leg. H. de Saeger (RBINS: 13535,
13540.1, 13540.2, RMCA: 36845 to 36847, RBINS: 13541); 9 ♀♀, 1 ♂ [4 SA, 5 AL, 1 UN], Pempere
Cave, 19 Jun. 1957, 11 Jun. 1958, 25 Aug. 1962, leg. Félix / Michel Anciaux de Faveaux, 25 Aug. 1962,
leg. unknown collector (RMCA: 27587 to 27590, SMF: 18944, 18945, MHNG: 1046.030, 1046.031,
RBINS: 14512, SMF: 21298); 1 ♀, 1 ♂ [2 AL], Pimpin Cave [= Pempere Cave], unknown date, leg.
the Royal Museum for Central Africa (RMCA) (BMNH: 1959.506, 1959.507).
Nycteridae Van der Hoeven, 1855
Nycteris G. Cuvier & E. Geoffroy, 1795
DEMOCRATIC REPUBLIC OF THE CONGO: 1 ad ♀, 4 ad ♂♂ [3 SS, 2 SO], unknown locality,
29 Nov. 1955, prior to 29 Nov. 1955, between 21 Nov. 1955 and 29 Nov. 1955, leg. Alexandre
Fain (RMCA: 23305, 23306, 23304, 23302, 23303); 1 ? [AL], Bamanya, between 1 Jan. 1974 and
31 Dec. 1974, leg. R.P. Hulsaert (RMCA: 75.012-M-0001); 1 ? [AL], Gumba, 10 Oct. 1937, leg.
Jean Jacques Deheyn (RMCA: 14263b); 3 ♂♂ [3 SN], Kateke [= Kateke [River]], 4 Dec. 1947, leg.
unknown collector (RBINS: 10687 to 10689); 2 ad ?? [2 SO], Keyberg, 20 Jan. 1952, leg. Uwe Rahm
(RMCA: 28897, 28898); 1 ♀ [SS], Luberezi [= Luberizi], 9 Mar. 1971, leg. Alexandre Prigogine
(RMCA: 36262); 2 ♂♂ [2 SS], Make, 17 Nov. 1992, 26 Jun. 1993, leg. Tshinyenye (RMCA: 92.148M-0001, 93.097-M-0023); 1 ♀ [SK], Musisi swamp area, 21 Jul. 2005, leg. Project Beatra (FMNH:
189123); 1 ? [AL], Nioka, 19 May 1953, leg. Célestin Hecq (RMCA: 21899b); 1 ? [UN], Rutshuru,
unknown date, leg. unknown collector (INPBC: W3.1); 1 ad ♀ [SA], Vieux Beni, 24 Aug. 1953, leg.
Gaston-François de Witte (RMCA: 37128); 1 juv ♂, 2 ?? [1 SS, 2 SO], Vumbi, between 1 Jan. 1952 and
31 Dec. 1952, leg. Alexandre Fain (RMCA: 81.012-M-0035, 81.012-M-0039, 81.012-M-0040).
Nycteris arge Thomas, 1903
BURUNDI: 1 ♂ [SB], Ruhinga Hill, 11 Mar. 1993, leg. Jay L. Udelhoven (FMNH: 155972).
DEMOCRATIC REPUBLIC OF THE CONGO: 1 ♀, 2 ♂♂, 1 ? [3 AL, 1 UN], unknown locality,
between 1 Jun. 1956 and 31 Jul. 1956, leg. Alvin Novick, prior to 1 Aug. 1957, leg. Heinz Stephan
189
European Journal of Taxonomy 382: 1–327 (2017)
(MCZ: 48241, 48242, 48244, RMCA: 27435); 1 ad ♂ [AL], Abialosa River, unknown date, leg. Jacques
Verschuren (INPBC: V1289); 1 ♂ [UN], Aka River, 22 May 1952, leg. H. de Saeger et al. (RBINS:
13547); 1 ad ♂, 1 juv ♂, 1 ♂ [1 AL, 1 SK, 1 UN], Aka-Inimvua conluence, 15 May 1952, leg. H. de
Saeger et al., 15 May 1952, leg. H. de Saeger (RBINS: 13546, RMCA: 36556, 36557); 1 ad ♀ [SK],
Alima, 15 Feb. 1959, leg. P. Dyleff (RMCA: 28155a); 1 ad ♂ [SA], Amangundu, unknown date, leg.
Jacques Verschuren (INPBC: V1294); 1 ad ♀, 1 ? [1 AL, 1 UN], Amatupi River [= Abatupi River],
unknown date, leg. Jacques Verschuren, unknown date, leg. unknown collector (INPBC: V1291.1,
V1291.2); 3 ad ♀♀, 3 ad ♂♂ [4 SS, 1 AL, 1 SK], Avakubi, 13 Oct. 1909, 16 Nov. 1909, 21 Nov. 1909,
24 Nov. 1909, 28 Nov. 1909, leg. Herbert Lang, James Paul Chapin and The American Museum Congo
Expedition (AMNH: 49140, 49371, 49133, RMCA: 12443, AMNH: 49136, 49137); 2 ?? [2 SA],
Babongena, between 1 May 1982 and 31 May 1982, leg. Marc Colyn (RMCA: 82.031-M-0009, 82.031M-0010); 1 ♂ [AL], Banalia, 17 Jan. 1939, leg. Duncan M. Hodgson and William F. Coultas (MCZ:
38010); 2 ad ♂♂ [2 SS], Beni, 8 Aug. 1906, leg. R.E. Dent (BMNH: 1906.12.4.61, 1906.12.4.62); 2 ad
♂♂ [2 SA], Boende, between 1 Oct. 1969 and 31 Oct. 1969, leg. P. Lootens, between 15 Mar. 1940 and
16 Mar. 1940, leg. R.P. Wynants (RMCA: 75.035-M-0028, 15950); 3 ad ♀♀, 1 ad ♂, 1 ad ? [1 SS, 4 SA],
Bokuma, 6 Jun. 1951, 30 Jan. 1952, between 30 Jan. 1952 and 8 May 1952, between 1 Jan. 1931 and
31 Dec. 1931, leg. P. Lootens (RMCA: 21409, 20546, 23243, 23244, 21410); 1 ♀ [UN], Bomane
[= Bomaneh], 22 May 2010, leg. Guy-Crispin Gembu Tungaluna, Prescott Musaba and José Akaibe
(UNIKIS: CRT2221); 1 juv ♂ [AL], Buluku, 22 May 1952, leg. H. de Saeger (RMCA: 36558); 1 ad ♀,
1 ad ?, 2 ?? [2 SA, 1 SO, 1 UN], Buta, 20 Feb. 1928, prior to 16 Jun. 1937, leg. Jozef Hutsebaut (RMCA:
9485, 13856 to 13858); 1 ♂ [SA], Bwandza, 30 Jun. 1979, leg. Frieder Meier (MHNG: 1870.092); 3 ??
[3 UN], Bwirina, unknown date, leg. unknown collector (INPBC: V1661.1 to V1661.3); 1 ad ♀ [SS],
Djamba, 20 Dec. 1924, leg. Henri Schouteden (RMCA: 8874); 2 ad ♀♀ [2 SA], Djelube River, middle,
unknown date, leg. Jacques Verschuren (INPBC: V1078.1, V1078.2); 1 ad ♀ [SA], Djumma River,
middle, unknown date, leg. Jacques Verschuren (INPBC: V1279); 1 ? [UN], Durba, 18 Oct. 1999, leg.
Robert (Bob) Swanepoel (TM: DBA001); 1 ♀, 1 ♂ [1 AL, 1 SB], Epulu, 20 Jun. 1981, leg. John
A. Hart, 27 Jun. 1991, leg. Julian C. Kerbis Peterhans (FMNH: 138137, 149405); 1 ♂ [AL], Equateur
(former) Province, 8 Apr. 1996, leg. unknown collector (SMF: 84410); 1 ad ♀, 2 ad ♂♂ [1 SA, 2 AL],
Equateur Province, 22 Jun. 1956, between 1 Jun. 1956 and 31 Jul. 1956, leg. Alvin Novick, between
1 Jan. 1921 and 31 Dec. 1921, leg. Verlaine (MCZ: 48238, 48243, RMCA: 80.034-M-0001); 3 ad ♂♂
[1 SA, 2 UN], Ibembo, unknown date, 13 Feb. 1950, leg. Jozef Hutsebaut (BMNH: 1954.791, 1954.792,
RMCA: 19632); 1 ad ♀, 1 ad ♂ [2 SA], Ikela, prior to 4 Jan. 1958, between 1 Jan. 1931 and 31 Dec. 1931,
leg. P. Lootens (RMCA: 25830, 27020); 1 ? [AL], Imbonga, between 1 Jan. 1987 and 31 Dec. 1987, leg.
P. Lootens (RMCA: 87.043-M-0003); 1 ad ? [AL], Inangongo, unknown date, leg. Jacques Verschuren
(INPBC: V1095.1); 1 ad ♀, 2 ad ♂♂, 2 ♂♂ [1 SS, 1 SA, 1 SK, 2 UN], Irangi, 21 Aug. 1959, 23 Jun. 1961,
11 Jul. 1961, leg. Uwe Rahm, 20 Mar. 1990, leg. Klaus-Gerhard Heller and Marianne Volleth,
24 Jan. 1984, leg. Heinz Stephan (BMNH: 1962.1852, SMF: 78331, RMCA: 28380, BMNH: 1962.1851,
SMF: 65029); 1 ad ♀, 2 ad ♂♂ [3 SA], Irsac Labo [= Luiro], unknown date, leg. H.J. Bredo, unknown
date, leg. Th. Andersen (ZFMK: 1964.1318 to 1964.1320); 1 ♀ [SS], Irumu, 30 Nov. 1924, leg. Edmund
Heller (FMNH: 26238); 1 ♂ [UN], Isai River, 18 Dec. 1974, leg. Michael D. Gallagher (HZM: 2.7845);
3 ad ♀♀, 2 ad ♂♂ [3 SS, 1 SA, 1 UN], Ituri Forest, unknown date, leg. R.E. Dent and R.B. Woosnam,
24 Apr. 1930, leg. Howard de Walden (BMNH: 1907.1.2.12, 1930.11.11.84 to 1930.11.11.87); 1 ♂ [SS],
Kasende, 5 Jul. 1934, leg. R.S. Pierrepont (MCZ: 31739); 1 ad ?, 1 ? [2 SS], Kawa (Forest), 1 Jan. 1952,
leg. unknown collector, 16 Jul. 1952, leg. Alexandre Fain (RBINS: 21663, BMNH: 1953.480); 1 ad ♀
[SS], Keri River, 26 Aug. 1926, leg. D. Fraser (BMNH: 1927.3.1.18); 1 ad ♀ [SS], Kiamokoto, 7 May
1948, leg. Gaston-François de Witte et al. (RBINS: 10838); 1 ad ♀, 1 ? [1 SK, 1 UN], Kilembe, 52 mi
[= 84 km] S, 8 Aug. 1957, leg. Robin E. Leech and Edward S. Ross (CAS: 12488, 12521); 1 ad ♀ [SA],
Kilo Mines [= Kilomines], between 1 Jan. 1930 and 31 Dec. 1930, leg. du Soleil (RMCA: 16200); 1 ad
♂ [SO], Kingulube, 14 Dec. 1968, leg. Alexandre Prigogine (RMCA: 35507); 2 ad ♀♀ [2 UN], Kisanga,
22 Apr. 1960, 25 Apr. 1960, leg. Uwe Rahm (BMNH: 1962.1853, 1963.1169); 1 ♂ [SS], Kisiki,
190
VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
19 Dec. 1938, leg. Duncan M. Hodgson and William F. Coultas (MCZ: 38025); 2 ad ♀♀ [1 SA, 1 UN],
Koloka, unknown date, leg. unknown collector, between 1 Jun. 1911 and 30 Jun. 1911, leg. H. Schubolz
(ZMH: 3, SMF: 6380); 1 ad ♂ [SA], Koteli [= Kotili], between 1 Dec. 1924 and 31 Dec. 1924, leg.
Henri Schouteden (RMCA: 16199); 1 ♀ [AL], Kumbi, 25 Nov. 1993, leg. Ngenge Masumbuko
Kamitongo (ZMA: MAM.24899); 1 ad ♂ [SS], Kunungu, 19 Nov. 1937, leg. N’Kele (RMCA: 14168);
1 ad ♀ [SS], Luebo, 21 Aug. 1921, leg. Henri Schouteden (RMCA: 6555); 1 ad ♂, 1 ♀, 2 ♂♂, 6 ??
[2 SS, 7 AL, 1 SO], Lukolela, 11 Aug. 1930, 7 Nov. 1930, 29 Nov. 1930, 27 Dec. 1930, 17 Mar. 1931,
leg. James Paul Chapin, 15 Nov. 1932, leg. Lecocq, 4 Sep. 1930, leg. Franklin Edson (AMNH: 86910,
RMCA: 12176, AMNH: 86864, 86865, 86867, 86913, 86912, 86769, 86770, 86911); 1 ♀ [AL], Mabali,
22 Jun. 1956, leg. Alvin Novick (MCZ: 48239); 1 ad ♂ [UN], Makala, 4 Dec. 1912, leg. J. Christie
(BMNH: 1914.4.8.5); 2 ?? [2 UN], Makpe River, upper [= Makpe], 13 Jul. 1951, leg. H. de Saeger
et al. (RBINS: 13543.1, 13543.2); 2 ♀♀ [2 SO], Medje, 2 May 1909, 16 Jan. 1910, leg. Herbert Lang,
James Paul Chapin and The American Museum Congo Expedition (AMNH: 49194, 49138); 2 ?? [2 UN],
Mogbwamu River, 8 Mar. 1952, leg. H. de Saeger et al. (RBINS: 13545.1, 13545.2); 1 ad ? [SS],
Mount Wago [= Wago, Mount], 1 Aug. 1952, leg. Alexandre Fain (BMNH: 1953.481); 1 ♀, 3 ♂♂
[4 SS], Muhulu, 15 Jan. 1957, leg. Heinz Stephan (RMCA: 27436 to 27439); 2 ad ♀♀, 1 ad ♂ [3 SS],
Munoi conluence, 2 Jun. 1948, leg. Gaston-François de Witte et al. (RBINS: 10835 to 10837); 2 ??
[2 AL], Niangara, 27 Mar. 1913, leg. Herbert Lang, James Paul Chapin and The American Museum
Congo Expedition (AMNH: 49386, 49387); 1 ad ♂ [SS], Panga, between 1 Sep. 1925 and 30 Sep. 1925,
leg. Henri Schouteden (RMCA: 9127); 1 ad ? [SS], Paulis [= Isiro], 5 Jan. 1950, leg. Abbeloos (RMCA:
19819); 1 ad ♀, 2 ad ♂♂, 1 juv ♂ [3 SA, 1 AL], Pawa, unknown date, leg. Reitter, 8 Nov. 1938, between
31 Dec. 1936 and 8 Nov. 1938, leg. Degotte (RMCA: 15728, 15216 to 15218); 2 ad ♀♀, 1 ad ♂ [1 SA,
2 AL], Ruanoli-Mavea conluence, unknown date, leg. Jacques Verschuren (INPBC: V1668.2,
V1668.3.1, V1668.3.2); 1 ad ♀, 1 ad ♂, 1 juv ♂, 1 ♀ [1 SS, 2 SA, 1 AL], Stanleyville [= Kisangani],
6 Oct. 1910, leg. Weyns, between 1 May 1926 and 31 May 1926, leg. Henri Schouteden, between
11 Jan. 1939 and 11 Dec. 1939, leg. Duncan M. Hodgson and William F. Coultas, between 5 Feb. 1928
and 5 Nov. 1928, leg. Collart (RMCA: 893, 16198, MCZ: 38032, RMCA: 16194); 1 ♂, 1 ? [2 UN],
Taere River, unknown date, leg. unknown collector, 10 Mar. 1952, leg. H. de Saeger et al. (INPBC:
4380.2, RBINS: 13544); 1 juv ♂ [AL], Tayele (River) [= Taere River], 10 Mar. 1952, leg. H. de Saeger
(RMCA: 36559); 1 ? [AL], Tshakala, between 1 Jan. 1982 and 31 Dec. 1982, leg. Marc Colyn (RMCA:
82.019-M-0010); 1 ad ♂ [UN], Ukaika, unknown date, leg. Rudolf Grauer (NMW: 10301); 5 ad ♀♀,
3 ad ♂♂ [4 SA, 4 AL], Virunga National Park, unknown date, leg. Jacques Verschuren
(INPBC: V1853A, V1853B, V1854A, V1854B, V1855.1A, V1855.1B, V1855.1C, RBINS: 1289B);
17 ?? [17 AL], Wafanya [= Wafania], 8 Oct. 1988, 29 Aug. 1991, 12 Sep. 1991, 25 Sep. 1991,
10 Feb. 1992, 20 Mar. 1992, 8 Apr. 1992, between 1 Dec. 1986 and 31 Dec. 1987, between 1 Aug. 1987
and 31 Aug. 1987, between 1 Jun. 1988 and 30 Jun. 1988, between 1 May 1988 and 31 May 1988,
between 1 Mar. 1992 and 31 Mar. 1992, between 1 Mar. 1991 and 31 Mar. 1991, between 1 Feb. 1991
and 28 Feb. 1991, between 1 Jan. 1992 and 31 Jan. 1992, leg. P. Lootens (RMCA: 92.079-M-0034,
92.079-M-0035, 91.076-M-0081, 92.079-M-0112, 92.079-M-0115, 91.076-M-0092, 91.076-M-0059,
92.079-M-0117, 89.020-M-0040, 92.079-M-0114, 91.076-M-0007, 91.076-M-0014, 92.079-M-0113,
89.020-M-0041, 89.020-M-0043, 88.005-M-0008, 87.043-M-0005); 1 ad ♀ [SS], Wago, Mount,
21 Sep. 1952, leg. Alexandre Fain (RMCA: 81.012-M-0037); 1 ad ? [SS], Wasa [= Wasa Wago Forest],
between 1 Jan. 1951 and 31 Dec. 1952, leg. Alexandre Fain (RMCA: 81.012-M-0038); 1 ad ♀ [SA],
Yalusaka, 16 Mar. 1958, leg. F. Carbone (RMCA: 27163); 1 ad ♂ [SA], Yambata, 11 Mar. 1914, leg.
De Giorgi (RMCA: 5443); 1 ad ♀ [SA], Yangambi, 7 Feb. 1942, leg. R.P. Van Boven (RMCA: 80.033M-0001).
191
European Journal of Taxonomy 382: 1–327 (2017)
Nycteris grandis Peters, 1865
DEMOCRATIC REPUBLIC OF THE CONGO: 1 ? [AL], Babongena, between 1 May 1982 and
31 May 1982, leg. Marc Colyn (RMCA: 82.031-M-0006); 1 ♀ [SK], Baliko, 7 Jun. 2012, leg. William
(“Bill”) T. Stanley (FMNH: 219603); 1 ad ? [SS], Beni, prior to 2 Jun. 1914, leg. Bonnevie (RMCA:
3240); 1 ad? ♀ [SO], Biyonga [= Bionga], 17 Aug. 1953, leg. L. Dupuis (RMCA: 21915); 1 ad ♂ [SA],
Boende, between 1 Jan. 1966 and 31 Jan. 1966, leg. P. Lootens (RMCA: 34162); 1 ad ♂ [SA], Bokuma,
18 May 1951, leg. P. Lootens (RMCA: 20679); 1 ? [AL], Boteka, 10 Oct. 1985, leg. P. Lootens (RMCA:
85.052-M-0001); 1 ad ♂ [SA], Camp Putnam, between 1 May 1953 and 31 May 1953, leg. Uwe
Rahm (RMCA: 28902); 1 ad ♂ [SA], Djuma River, upper, unknown date, leg. Jacques Verschuren
(INPBC: V1277.1); 1 ad ♀ [SA], Epulu, between 1 Oct. 1956 and 31 Oct. 1956, leg. Max Poll (RMCA:
25652); 1 ♂ [AL], Irangi, 18 Jan. 1984, leg. Heinz Stephan (SMF: 65028); 1 juv ♂, 1 ♀ [2 AL],
Irangi, 3 mi [= 4.8 km] from, 16 Oct. 1990, leg. Wim Bergmans (ZMA: MAM.24178, MAM.24179);
1 ad ♂ [SS], Ituri Forest, 24 Apr. 1930, leg. Howard de Walden (BMNH: 1930.11.11.89); 1 ad ♂
[SA], Kalikimvua, 11 Jun. 1952, leg. H. de Saeger et al. (RBINS: 13621); 1 ad ♂ [SO], Kamituga,
6 Jan. 1964, leg. Alexandre Prigogine (RMCA: 32393); 1 ♀ [AL], Kasiba, 15 Oct. 1990, leg. Wim
Bergmans (ZMA: MAM.24177); 1 ♀, 2 ♂♂ [2 SA, 1 AL], Kivu region, between 11 Aug. 1989 and
14 Aug. 1989, between 1 Jan. 1988 and 31 Dec. 1988, leg. unknown collector (SMF: 73700, 77099,
77100); 1 ad ♀, 1 ad ♂, 1 ♀ [3 SS], Luluabourg [= Kananga], 28 May 1922, 11 Mar. 1926, leg.
Richard Callewaert (BMNH: 1927.12.21.10, 1927.12.21.8, 1926.7.6.72); 2 ♀♀ [2 AL], Mabali,
between 18 Jun. 1956 and 19 Jul. 1956, leg. Alvin Novick (MCZ: 48236, 48237); 1 ad ♂, 2 ?? [1 SA,
2 UN], Makayova River, lower, unknown date, leg. unknown collector, unknown date, leg. Jacques
Verschuren (INPBC: V1256.1 to V1256.3); 2 ?? [2 AL], Masako, unknown date, leg. Ndey B. Ifuta
(RMCA: 93.079-M-0341, 93.079-M-0342); 1 ♀, 1 ? [2 UN], Menga, 25 Jan. 1996, leg. Robert (Bob)
Swanepoel (TM: 45568, 45569); 1 ad ♀, 1 ? [1 SA, 1 UN], Mogbwamu River, 8 Jul. 1951, leg. H. de
Saeger et al., 8 Jul. 1951, leg. H. de Saeger (RBINS: 13620, RMCA: 36560); 1 ad ♂ [UN], Mulolo,
15 Mar. 1960, leg. Uwe Rahm (BMNH: 1962.1849); 1 ad ♀ [SA], Netonna, 8 Jun. 1890, leg. P. Hesse
(SMF: 2514); 1 ad ♂ [SA], Omaniumdu, 27 Jul. 1959, leg. Laurent (RMCA: 29536); 1 ad ? [SS], Poko,
between 25 Feb. 1914 and 25 Jul. 1914, leg. Cuthbert Christy (RMCA: 4340); 1 ♀, 1 ♂ [2 AL], Quatorze,
1 Jul. 2013, leg. William (“Bill”) T. Stanley (FMNH: 222427, 222428); 1 ad ♂ [UN], Ukaika, unknown
date, leg. Rudolf Grauer (NMW: 10300); 1 ♀, 5 ?? [5 AL, 1 UN], Wafanya [= Wafania], 10 Oct. 1991,
12 Oct. 1991, between 12 Oct. 1991 and 18 Oct. 1991, between 1 Nov. 1991 and 30 Nov. 1991, between
1 Aug. 1987 and 31 Aug. 1987, leg. P. Lootens (RMCA: 91.076-M-0106, 91.076-M-0109, 92.079-M0033, 88.005-M-0009, 92.079-M-0110, 91.076-M-0102); 1 ad ♀ [UN], Yalosemba, 27 Jun. 1979, leg.
C. Brian Robbins (USNM: 537706); 1 ♂ [AL], Yayama, 24 Jun. 1986, leg. WHO and Marc Colyn
(RMCA: 88.011-M-0013).
Nycteris hispida (Schreber, 1774)
BURUNDI: 1 ? [AL], unknown locality, between 1 Jan. 1965 and 31 Dec. 1970, leg. P.L. Niort (MHNG:
1896.015); 1 ad ♀ [SK], Kayanza, 12 Mar. 1979, leg. Heinz Stephan (SMF: 57536); 1 ♂ [AL], Mweya,
29 Apr. 1971, leg. P.L. Niort (MHNG: 1896.018).
DEMOCRATIC REPUBLIC OF THE CONGO: 1 ad ♂, 1 ad ?, 1 juv ?, 5 ?? [1 AL, 1 SK, 1 SO, 5 UN],
unknown locality, unknown date, leg. Cuthbert Christy, 13 Aug. 1958, leg. N’Kele, between 1 Jan. 1931
and 31 Dec. 1931, leg. E. Jans, between 15 Jul. 1999 and 18 Jul. 1999, between 11 Aug. 1999 and
15 Aug. 1999, between 1 Jul. 1999 and 4 Jul. 1999, leg. unknown collector (RMCA: 3463, 27666,
27371, SMF: 90192 to 90194, 90190, 90191); 1 ad ♀ [AL], Aka, 14 May 1952, leg. H. de Saeger et al.
(RBINS: 13612); 1 ? [UN], Albert National Park [= Virunga National Park], unknown date, leg.
unknown collector (INPBC: V1838.2); 1 ♀ [UN], Ankoro, 3 Nov. 1974, leg. Michael D. Gallagher
(HZM: 45.7852); 2 ad ♀♀, 2 ad ??, 1 juv ?, 1 ♀, 1 ♂, 1 ? [6 SS, 1 SA, 1 AL], Avakubi, 15 Nov. 1909,
192
VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
17 Nov. 1909, 28 Jan. 1914, 15 Aug. 1914, leg. Herbert Lang, James Paul Chapin and The American
Museum Congo Expedition, 17 Sep. 1912, 26 Sep. 1912, leg. Cuthbert Christy, 30 Oct. 1912, leg.
unknown collector, between 1 Jan. 1914 and 31 Jan. 1914, leg. J. Bequaert (RMCA: 12386, AMNH:
49141, RMCA: 1426, AMNH: 49134, RMCA: 1423, AMNH: 49368, BMNH: 1914.4.8.4, RMCA:
5445); 1 ♂ [UN], Avakubi, near, 10 Dec. 1974, leg. Michael D. Gallagher (HZM: 44.7851); 1 ? [SA],
Babongena, between 1 May 1982 and 31 May 1982, leg. Marc Colyn (RMCA: 82.031-M-0007); 1 ad ?,
1 juv ♀, 1 juv ♂ [1 SS, 2 SO], Bambesa, 22 Jun. 1938, between 30 Jul. 1940 and 30 Aug. 1940, between
12 Apr. 1941 and 22 Apr. 1941, leg. J.M. Vrijdagh (RMCA: 14798, 17754, 17767); 1 ad ♀, 1 ad ♂
[2 SA], Banana, between 1 May 1952 and 31 May 1952, between 1 Jan. 1952 and 31 Jan. 1952, leg. Is.
Mesmaekers (RMCA: 20750, 21377); 1 ad ♀, 1 ad ♂, 1 juv ♂ [2 SA, 1 AL], Boende, between 1 Dec. 1969
and 31 Dec. 1969, between 1 Jan. 1931 and 31 Dec. 1931, leg. P. Lootens (RMCA: 35639, 75.035-M0022, 75.035-M-0023); 2 ad ♂♂ [2 SA], Bokuma, between 1 Jan. 1931 and 31 Dec. 1931, leg. P. Lootens
(RMCA: 22134, 23245); 1 ad ♀ [SS], Bolobo, between 1 Jan. 1931 and 31 Dec. 1931, leg. N’Kele
(RMCA: 20258); 1 ad ♀, 3 ?? [3 SA, 1 AL], Boma, 27 Jan. 1915, leg. Herbert Lang, James Paul Chapin
and The American Museum Congo Expedition, 3 Sep. 1913, leg. Styczynski (AMNH: 49392 to 49394,
RMCA: 2486); 1 ? [AL], Boteka, 14 Oct. 1985, leg. P. Lootens (RMCA: 85.052-M-0002); 1 ad ♀ [AL],
Bukavu, unknown date, leg. Jacques Verschuren (INPBC: V394); 1 ? [AL], Bunia, between 1 Apr. 1927
and 30 Apr. 1927, leg. Harold Jefferson Coolidge (MCZ: 28546); 6 ad ♀♀, 1 ad ♂ [6 SA, 1 AL], Buta,
unknown date, prior to 9 Nov. 1938, prior to 3 Nov. 1939, leg. Jozef Hutsebaut (RMCA: 14562a, 14562b,
14585, 15737 to 15739, 14903); 1 ♀, 2 ?? [1 SS, 1 AL, 1 SK], Catchpole’s Farm, 27 Jul. 1924,
28 Aug. 1924, leg. Edmund Heller, 29 Aug. 1924, leg. Alfred Morris Collins (FMNH: 30711, 30712,
26239); 27 ad ♀♀, 21 ad ♂♂, 4 ad ??, 2 sad ♂♂, 2 juv ♀♀, 3 juv ??, 2 foetus ??, 2 ♀♀, 4 ?? [3 SS,
22 SA, 36 AL, 2 SK, 4 UN], Cel II, unknown date, leg. unknown collector, 16 Feb. 1951, 1 Mar. 1951,
7 Mar. 1951, 17 Mar. 1951, 30 Mar. 1951, 2 Apr. 1951, 21 May 1951, 22 May 1951, 26 May 1951,
25 Jun. 1951, 19 Jul. 1951, 21 Jul. 1951, 25 Sep. 1951, 2 Nov. 1951, 15 Jan. 1952, 31 Jan. 1952,
1 Feb. 1952, 16 Feb. 1952, 18 Feb. 1952, 26 Feb. 1952, 27 Feb. 1952, 2 Apr. 1952, 3 May 1952, 8 May
1952, 16 Jul. 1952, leg. H. de Saeger et al., 1 Mar. 1951, 17 Mar. 1951, 30 Mar. 1951, 2 Apr. 1951,
12 May 1951, 21 May 1951, 31 May 1951, 19 Jul. 1951, 21 Jul. 1951, 31 Jan. 1952, 1 Feb. 1952,
26 Feb. 1952, 7 Apr. 1952, 8 May 1952, 16 Jul. 1952, between 12 Feb. 1952 and 18 Feb. 1952, leg. H. de
Saeger (INPBC: 3305, 3427, RBINS: 13597, RMCA: 36597, RBINS: 13574, RMCA: 36565, 36568,
RBINS: 13595, 13572, 13596, 13617, RMCA: 36594, RBINS: 13575, RMCA: 36566, RBINS: 13587,
13600.1, 13600.2, 13585, 13592.1, 13592.2, RMCA: 36577, RBINS: 13576, RMCA: 36567, RBINS:
13609.1, 13609.2, 13594, 13579.1, 13579.2, RMCA: 36569, RBINS: 13591.1, 13591.2, RMCA: 36575,
36576, RBINS: 13580.1, 13580.2, 13583, 13593.1, 13593.2, 13602.1, 13602.2, RMCA: 36584, RBINS:
13582, 13601, 13616, 13571, RMCA: 36563, 36564, RBINS: 13598.1 to 13598.4, RMCA: 36578 to
36581, 36587, 36570, 36571a, 36571b, RBINS: 13573, RMCA: 36591, 36592a, 36592b, RBINS:
13615, RMCA: 36593, 36582, 36583); 1 ♂ [SS], Cinputu, 13 Sep. 1983, leg. Betu Nsankulu (RMCA:
84.035-M-0024); 1 ad ♀, 1 ad ♂ [1 SS, 1 SA], Djamba, 19 Dec. 1924, between 1 Dec. 1924 and
31 Dec. 1924, leg. Henri Schouteden (RMCA: 8873, 16195); 1 ad ♀ [SA], Djeka, 29 May 1955, leg.
Roiseux (RMCA: 23634); 1 ad ♂, 1 ad ?, 1 ? [1 SA, 2 SK], Duma, 12 Oct. 1910, leg. unknown collector,
12 Oct. 1910, leg. H. Schubolz, between 1 Sep. 1911 and 30 Sep. 1911, leg. A.T.M. Herzog (SMF: 6381,
6381??6382, 6382); 1 ad ? [SO], Eala, 24 Jan. 1921, leg. Henri Schouteden (RMCA: 6558); 1 ♂ [UN],
Edouard, Lac [= Edward, Lake], 11 Nov. 1954, leg. unknown collector (RBINS: 15465); 1 ♂ [AL],
Epulu, 13 km E, 26 Apr. 1981, leg. John A. Hart (FMNH: 138138); 2 ad ♂♂, 1 ♀, 3 ♂♂, 3 ?? [3 SS,
1 SA, 5 AL], Faradje, 22 Nov. 1911, 25 Feb. 1912, 1 Mar. 1912, 9 Mar. 1912, 13 Jan. 1913, leg. Herbert
Lang, James Paul Chapin and The American Museum Congo Expedition (AMNH: 49085, 49144
[holotype Nycteris pallida J.A. Allen, 1917], 49418, 49417, RMCA: 12390, AMNH: 49419, 49424,
49421, MCZ: 17226 [topotype Nycteris pallida J.A. Allen, 1917]); 1 ad ♂ [SS], Fundi, between
16 Feb. 1912 and 16 Nov. 1912, leg. Cuthbert Christy (RMCA: 1608); 2 ?? [2 UN], Gangala-na-Bodio,
unknown date, 9 Apr. 1948, leg. unknown collector (INPBC: HV.11b, RBINS: 8123); 4 ad ♀♀, 2 ad ♂♂,
193
European Journal of Taxonomy 382: 1–327 (2017)
1 ad ?, 1 juv ?, 1 ? [6 AL, 1 SO, 2 UN], Garamba National Park, unknown date, leg. unknown
collector, 18 Jan. 1951, 16 Feb. 1951, 30 Mar. 1951, 21 Jul. 1951, 18 Mar. 1952, between 1 Jan. 1950
and 31 Dec. 1952, leg. H. de Saeger et al., between 1 Jan. 1950 and 31 Dec. 1952, leg. H. de Saeger
(INPBC: 5120, RBINS: 13578, 13570, 13605, 13590, 13569.1, 13569.2, 13618, RMCA: 36599); 1 ad
♀, 1 ad ♂ [2 SA], Goma, 10 Mar. 1953, 5 Dec. 1953, leg. The KEA-Mission (RBINS: 14457, 14458);
1 ad ♂ [SK], How…, unknown date, leg. J.J. Laarman (RMNH: MAM.16343); 2 ad ♀♀ [1 SA, 1 AL],
Ikela, 3 Jun. 1955, between 1 Jan. 1931 and 31 Dec. 1931, leg. P. Lootens (RMCA: 25826, 25828); 1 ad
♀ [SS], Irangi, 15 Aug. 1956, leg. J.J. Laarman (RMNH: MAM.16365); 1 ♀, 2 ?? [1 SK, 2 UN],
Irumu, unknown date, leg. unknown collector, 10 Dec. 1924, leg. Edmund Heller, 14 Sep. 1926, leg.
James Paul Chapin, Dewitt L. Sage, Frank P. Mathews and The Ruwenzori-Kivu Expedition
(INPBC: W508, FMNH: 30713, AMNH: 82388); 1 ad ♂, 2 ?? [1 SA, 2 UN], Ishango River, unknown
date, leg. Jacques Verschuren, unknown date, leg. unknown collector (INPBC: V1297.1, V1297.2,
W1391); 1 ad ♂ [SS], Kadia, between 1 Jul. 1931 and 31 Jul. 1931, leg. Gaston-François de Witte
(RMCA: 11402); 2 ♀♀, 1 ♂ [3 AL], Kadjidji [= Kajuju], unknown date, leg. Guy Babault (MNHN: ZMMO-1984-1337, ZM-MO-1984-1345, ZM-MO-1984-1346); 1 ad ♀ [SA], Kakungu, 23 Nov. 1953, leg.
Gaston-François de Witte (RMCA: 37121); 1 ad ♂ [AL], Kamande, unknown date, leg. Jacques
Verschuren (INPBC: V1057.1); 1 ad ♂ [SA], Kamituga, between 1 Dec. 1956 and 31 Dec. 1956, leg.
Alexandre Prigogine (RMCA: 26189); 1 ? [AL], Kampala, between 1 Jan. 1935 and 31 Dec. 1935, leg.
Guy Babault (MCZ: 46087); 1 ♂ [SS], Kananga, 4 Jan. 1974, leg. Western New Mexico University
(FMNH: 152456); 1 ad ♀ [SA], Karawa, prior to 24 Nov. 1936, leg. R.P. Wallin (RMCA: 14568); 1 ad
♂ [SA], Kasaji, 13 Mar. 1953, leg. Fisher (RMCA: 21807); 1 ? [UN], Kasaka, unknown date, leg.
unknown collector (INPBC: V1144.35); 1 ad ♀ [SA], Kasenga, between 1 Jul. 1936 and 31 Aug. 1936,
leg. Schwetz (RMCA: 13699); 1 ? [AL], Kasengi [= Kasengu], 27 Aug. 1926, leg. James Paul Chapin,
Dewitt L. Sage, Frank P. Mathews and The Ruwenzori-Kivu Expedition (AMNH: 82387); 2 ad ♀♀
[1 AL, 1 UN], Kasenyi [= Kasenye], 2 Dec. 1957, leg. Robin E. Leech and Edward S. Ross, 25 Feb. 1954,
leg. The KEA-Mission (CAS: 12484, RBINS: 15464); 1 ad ♂, 1 ? [1 SK, 1 UN], Kassi River, unknown
date, leg. unknown collector, 5 Apr. 1951, leg. H. de Saeger (INPBC: 3253, RMCA: 36595); 1 ad ♀
[AL], Kaswabilenga, 1 Oct. 1947, leg. Gaston-François de Witte et al. (RBINS: 10808); 1 ? [UN],
Katanda, unknown date, leg. unknown collector (INPBC: V1838.1); 2 ♀♀ [2 SS], Katobwe,
27 Nov. 1926, 30 Nov. 1926, leg. John Todd Zimmer (FMNH: 29408, 29409); 3 ad ?? [3 SS], Kawa
(Forest), 16 Jul. 1952, 17 Jul. 1952, 5 Aug. 1952, leg. Alexandre Fain (RMCA: 81.012-M-0014, BMNH:
1953.482, 1953.483); 1 ad ♀ [SS], Kiambi, 7 May 1931, leg. Gaston-François de Witte (RMCA: 11323);
1 ? [UN], Kiavikire [= Kiavikere], unknown date, leg. unknown collector (INPBC: W587); 1 ad ♂
[SO], Kingabwa, 19 Dec. 1975, leg. Tshinyenye (RMCA: 76.010-M-0003); 1 ? [AL], Kisangani,
between 1 Jun. 1982 and 30 Jun. 1982, leg. Marc Colyn (RMCA: 82.031-M-0034); 1 ad ? [SS], Kiseny
[= Kasenye], between 1 Jan. 1942 and 31 Dec. 1942, leg. R. Van Saceghem (RMCA: 17339); 1 ? [UN],
Kisisile, unknown date, leg. unknown collector (INPBC: W844); 1 ad ♂ [SA], Kitonga, unknown date,
leg. Jacques Verschuren (INPBC: V1302); 1 ad ♀ [AL], Kivu, Lake [= Lake Kivu], between 1 May
1953 and 31 May 1953, leg. The KEA-Mission (RBINS: 14462); 1 ad ? [SS], Korovi, Mount,
25 Oct. 1951, leg. Alexandre Fain (RMCA: 23280); 1 ad ♂ [SS], Koteli [= Kotili], 2 Jan. 1925, leg.
Henri Schouteden (RMCA: 8876); 1 ad ♂ [SS], Kunungu, 11 Dec. 1927, leg. Ngwe (RMCA: 9391);
1 ad ♂ [SS], Kwamouth, 17 Jun. 1921, leg. Henri Schouteden (RMCA: 6559); 2 ad ♀♀ [1 SA, 1 AL],
Kyavinionge, 20 May 1954, leg. Gaston-François de Witte, 27 Nov. 1953, leg. The KEA-Mission
(RMCA: 37124, RBINS: 14459); 1 juv ♂ [AL], Léopoldville [= Kinshasa], 12 Mar. 1913, leg. Cuthbert
Christy (RMCA: 1536); 1 ad ? [SO], Leverville [= Lusanga], between 1 Feb. 1929 and 28 Feb. 1929,
leg. Tinant (RMCA: 9986); 1 ad ♂ [SS], Lukonzolwa, 16 Feb. 1931, leg. Gaston-François de Witte
(RMCA: 11470); 2 ad ♀♀, 5 ad ♂♂, 2 ♀♀ [7 SS, 1 SA, 1 UN], Luluabourg [= Kananga], unknown
date, 17 Jul. 1922, 19 Oct. 1923, 2 Dec. 1925, 25 Feb. 1926, leg. Richard Callewaert, 4 Jun. 1964, leg.
Antoon Emeric Marcel De Roo (BMNH: 1926.11.1.27, 1926.7.6.73, 1926.7.6.74, 1934.10.24.18,
RMCA: 7568, BMNH: 1926.7.6.76, 1926.11.1.26, 1927.12.21.9, RMCA: 33601); 1 ad ♀, 1 ad ♂ [1 SA,
194
VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
1 UN], Lwiro [= Luiro], 1 Aug. 1964, leg. U. Goepel, 8 Jul. 1958, leg. Uwe Rahm (ZMZ: 124600,
RMCA: 33535); 5 ad ♀♀, 2 ad ♂♂ [5 SS, 1 SK, 1 SO], Mabwe River, 11 Feb. 1949, 1 Mar. 1949, leg.
Gaston-François de Witte et al. (RBINS: 10839 to 10844, 10846); 1 ad ♀, 1 ad ♂ [1 SS, 1 SK], Makala,
3 Dec. 1912, 5 Dec. 1912, leg. Cuthbert Christy (RMCA: 1607, 1609); 1 ad ♂ [SA], Makengo,
11 Dec. 1946, leg. Dartevelle (RMCA: 18161); 2 ad ♀♀, 1 ad ♂ [2 SA, 1 SO], Makpe, 12 Jul. 1951,
14 Jul. 1951, leg. H. de Saeger (RMCA: 36573, 36574, 36596); 1 ad ♀, 1 ad ♂ [2 SA], Makpe River,
upper [= Makpe], 10 Jul. 1951, 12 Jul. 1951, leg. H. de Saeger et al. (RBINS: 13586, 13589); 1 ad ♀,
1 ? [1 SA, 1 SK], Maleli, 11 Jun. 1951, leg. H. de Saeger et al. (RBINS: 13584.1, 13584.2); 1 ad ♀ [SA],
Malili, 14 Jun. 1951, leg. H. de Saeger (RMCA: 36572); 1 ♀, 1 ? [1 AL, 1 UN], Mayumbe, near,
unknown date, prior to 2 Apr. 1902, leg. unknown collector (MNHN: ZM-MO-1884-751, RBINS:
200b); 1 ad ♀, 1 ad ? [1 SA, 1 UN], Mbwambala, 21 Jun. 1995, 29 Jun. 1995, leg. Herwig Leirs et al.
(RMCA: 97.021-M-0837, MSB: 274621); 1 ad ♀, 1 ♂ [1 SO, 1 UN], Medje, 20 Jan. 1910, leg. Herbert
Lang, James Paul Chapin and The American Museum Congo Expedition, 9 Apr. 1914, leg. Cuthbert
Christy (AMNH: 49143, RMCA: 4343); 1 ad ♀ [AL], Meridi River, 20 Apr. 1952, leg. H. de Saeger
et al. (RBINS: 13611); 1 ad ♀, 1 ad ♂ [2 AL], Moko River, 20 Jun. 1952, leg. H. de Saeger et al.
(RBINS: 13614.1, 13614.2); 1 ad ♂ [SA], Moliro, prior to 28 Feb. 1899, leg. Charles Lemaire (RMCA:
162); 1 ad ♀, 3 ad ♂♂, 1 ad ? [4 SA, 1 SO], Mongbwalu, 30 May 1939, leg. Janssens, prior to
16 Feb. 1938, leg. Harford (RMCA: 15560, 14382a to 14382d); 1 ? [AL], Mosenge, 15 Aug. 1986, leg.
WHO and Marc Colyn (RMCA: 88.011-M-0014); 1 ad ♂ [SA], Mukishi [Station], between 29 Dec. 1927
and 9 Feb. 1928, leg. Becquet (RMCA: 16193); 1 juv ♂ [AL], Mulungu, prior to 1 Feb. 1940, leg.
F.L. Hendricks (RMCA: 15872); 1 ad ♀, 1 ad ♂ [2 SA], Murambi, 25 Aug. 1955, leg. Gaston-François
de Witte (RMCA: 37126, 37127); 1 ad ♂ [SA], Mushie, between 1 Jan. 1931 and 31 Dec. 1931, leg.
Cuthbert Christy (RMCA: 1537); 1 ad ♂ [AL], Musosa, near, between 1 Jun. 1940 and 30 Jun. 1940,
leg. H.J. Bredo (RBINS: 14461); 2 ad ♀♀, 2 ad ♂♂, 3 ?? [3 SA, 1 SO, 3 UN], Mutsora, unknown date,
leg. unknown collector, 15 Dec. 1952, 13 Feb. 1954, 4 Aug. 1954, between 11 Dec. 1952 and 11 Dec. 1953,
leg. Gaston-François de Witte (INPBC: V489, W1020, W814, RMCA: 37123, 37120, 37125, 37122);
1 ad ♀ [SO], Mutsora Station, 11 Apr. 1950, leg. J. de Wilde (RBINS: 12943); 1 ad ♀, 1 ad ? [2 SS],
Mutwanga, between 1 Nov. 1937 and 30 Nov. 1937, between 1 Sep. 1937 and 30 Sep. 1937, leg. Henri
Martin Hackars (RMCA: 16603, 21546); 1 ad ? [SA], Mweka Kasai [= Mweka], between 1 Jan. 1930
and 31 Dec. 1930, leg. Heusgens (RMCA: 11550); 2 ad ♂♂ [2 SA], Nadegbe River, 10 Mar. 1952, leg.
H. de Saeger (RMCA: 36585, 36586); 1 ad ♀, 1 ad ? [2 AL], Nampume, 6 Mar. 1952, leg. H. de Saeger
et al. (RBINS: 13606.1, 13606.2); 1 ad ♀ [AL], Nampune [= Nampume], 6 Mar. 1952, leg. H. de
Saeger (RMCA: 36588); 1 ad ♂ [AL], Ndelele, 13 Mar. 1952, leg. H. de Saeger et al. (RBINS: 13603);
1 juv ? [SO], Ndwa, between 1 Oct. 1950 and 30 Nov. 1950, leg. Ngwe (RMCA: 20255); 1 ad ♀, 3 ad
♂♂ [3 SS, 1 SA], Niambasha, 30 Jan. 1959, leg. Uwe Rahm (RMCA: 28026, 28899 to 28901); 1 ♀,
1 ♂ [2 AL], Niangara, 29 Feb. 1948, leg. Neal A. Weber (AMNH: 150183, 150184); 1 ad ? [SS], Paulis
[= Isiro], 10 Dec. 1950, leg. Abbeloos (RMCA: 20511); 2 ad ♀♀, 1 ad ♂ [1 SS, 2 SA], PFSK 8,
25 Mar. 1952, between 15 Mar. 1952 and 25 Mar. 1952, leg. H. de Saeger (RMCA: 36598, 36589,
36590); 1 ad ♀ [AL], PFSK.17, 26 Mar. 1952, leg. H. de Saeger et al. (RBINS: 13608); 1 ad ♂ [AL],
PFSK.8, 25 Mar. 1952, leg. H. de Saeger et al. (RBINS: 13607); 3 ad ?? [2 SS, 1 SK], Poko, 17 Jul. 1914,
21 Jul. 1914, 2 Aug. 1914, leg. Cuthbert Christy (RMCA: 4336b, 4342, 4341); 1 juv ♀ [AL], Pweto,
unknown date, leg. Gerard (RMCA: 5441b); 2 ad ♀♀, 1 ad ?, 1 juv ♀ [1 SA, 2 AL, 1 UN], Rutshuru,
unknown date, leg. Rudolf Grauer, 30 Apr. 1938, 29 May 1938, 31 May 1938, leg. Serge Frechkop
(NMW: 10304, RMCA: 17088, RBINS: 4783, RMCA: 17089); 4 ad ♀♀ [4 UN], Ruwenzori
SE, unknown date, leg. unknown collector, 11 May 1906, leg. R.B. Woosnam, 17 Jun. 1906, leg.
R.E. Dent (BMNH: 1906.12.4.75, 1906.12.4.76, 1906.12.4.17, 1906.12.4.16); 1 ? [UN], Samboko
River, unknown date, leg. unknown collector (INPBC: V1034.2); 1 ♀ [UN], Scierie Forest,
22 Nov. 1974, leg. Michael D. Gallagher (HZM: 43.7850); 3 ♀♀, 1 ♂ [3 SS, 1 UN], St. Jos. (Mission)
[= Mikalaya], 2 Aug. 1924, 15 Aug. 1924, 8 Oct. 1924, leg. Richard Callewaert (AMNH: 86177, 86172,
86175, 86143); 1 ad ♀ [SS], St. Joseph de Luluabourg Mission [= Mikalaya], 17 Jul. 1922, leg.
195
European Journal of Taxonomy 382: 1–327 (2017)
Richard Callewaert (RMCA: 7545); 2 ad ♀♀, 1 ♀, 1 ♂, 1 ? [2 SA, 2 AL, 1 SO], Stanleyville [=
Kisangani], 8 Aug. 1909, 1 Jan. 1915, leg. Herbert Lang, James Paul Chapin and The American Museum
Congo Expedition, between 16 Jul. 1928 and 16 Aug. 1928, leg. Collart, prior to 29 Nov. 1933, leg.
Schwetz (AMNH: 49390, 49139, 49531, RMCA: 16203, 12223); 2 ad ♂♂ [2 UN], Tandala, 16 Jul. 1967,
leg. Lynn W. Robbins, 18 Jul. 1979, leg. C. Brian Robbins (USNM: 537703, 537705); 1 ad ♀ [SK],
Teturi, 12 Sep. 1946, leg. Jean Hiernaux (RBINS: 7045); 1 ? [UN], Thysville [= Mbanza-Ngungu],
11 Feb. 1987, leg. unknown collector (RBINS: 22431); 1 ? [AL], Tshakala, between 1 Jan. 1982 and
31 Jan. 1982, leg. Marc Colyn (RMCA: 82.019-M-0009); 1 ♂ [SO], Tshambi, 10 Nov. 1933, leg.
unknown collector (RBINS: 4782); 1 ad ♂ [SO], Utu [River], 6 Feb. 1959, leg. Uwe Rahm (RMCA:
33531); 1 ad ♀ [SA], Utukuru, 10 Jun. 1952, leg. H. de Saeger et al. (RBINS: 13613); 2 ad ♂♂, 1 ♀
[1 AL, 2 UN], Uvira, unknown date, leg. Rudolf Grauer, 26 Jul. 1956, leg. Alvin Novick (NMW: 10302,
10303, MCZ: 48240); 1 juv ♂, 1 ♀, 2 ?? [4 AL], Vankerckhovenville [= Nzoro], 18 Apr. 1912, leg.
Herbert Lang, James Paul Chapin and The American Museum Congo Expedition (AMNH: 49395 to
49398); 1 ad ♀ [AL], Vitshumbi, 2 Jan. 1953, leg. The KEA-Mission (RBINS: 14456); 1 ad ♀ [SS],
Vumbi, between 1 Jan. 1952 and 31 Dec. 1952, leg. Alexandre Fain (RMCA: 81.012-M-0036); 2 ??
[1 SA, 1 AL], Wafanya [= Wafania], 25 Sep. 1991, 26 May 1992, leg. P. Lootens (RMCA: 91.076-M0090, 92.079-M-0111); 1 ♂ [SS], Yalosemba, 20 Jun. 1979, leg. C. Brian Robbins (USNM: 537704);
1 ad ♂ [SS], Yangambi, 10 Nov. 1947, leg. G. Gilbert (RBINS: 6987).
RWANDA: 1 juv ? [SS], Astrida [= Butare], between 1 Jan. 1951 and 31 Jan. 1951, leg. Aureliaan
(RMCA: 20490); 1 ad ♂ [SS], Butare, unknown date, leg. Alexandre Fain (RMCA: 81.012-M-0010);
1 ad ♀ [SA], Cyili Ntyano [= Cyili], 2 May 1985, leg. Théoneste (RMCA: 87.047-M-0001); 1 ♂ [AL],
Lac Ihema [= Ihema, Lake], 9 Sep. 1980, leg. Hans H. de Iongh (ZMA: MAM.21230); 1 ad ♂, 1 ♀
[1 SS, 1 SA], Nyarutovu, 18 May 1955, leg. Alexandre Fain (MHNG: 921.035, RMCA: 81.012-M0029); 2 ad ♀♀, 2 ad ♂♂, 1 juv ♂ [5 SS], Rukara, between 1 Jan. 1955 and 31 Dec. 1955, leg.
Alexandre Fain (RMCA: 81.012-M-0030 to 81.012-M-0034).
Nycteris intermedia Aellen, 1959
DEMOCRATIC REPUBLIC OF THE CONGO: 1 ad ♂ [SA], unknown locality, between 1 Jun. 1911
and 30 Jun. 1911, leg. H. Schubolz (SMF: 6383); 4 ad ♂♂ [2 SA, 2 AL], Djelube River, middle,
unknown date, leg. Jacques Verschuren (INPBC: V1076.1, V1076.2, V1077.1, V1077.2); 1 ad ♀ [SS],
Ituri Forest, unknown date, leg. Howard de Walden (BMNH: 1930.11.11.82); 1 ad ♂ [SS], Karambi,
27 Sep. 1953, leg. J.J. Laarman (RMNH: MAM.16340); 1 ad ♀ [UN], Kisanga, 22 Apr. 1960, leg. Uwe
Rahm (BMNH: 1962.1850); 1 ad ♀ [SA], Ruanoli-Mavea conluence, unknown date, leg. Jacques
Verschuren (INPBC: V1668.4); 1 ad ♂ [SS], Teturi, 25 May 1948, leg. Jean Hiernaux (RBINS: 7046);
1 ad ♂ [SA], Virunga National Park, unknown date, leg. Jacques Verschuren (INPBC: V1655).
Nycteris macrotis Dobson, 1876
BURUNDI: 1 ad ♀, 1 ad ♂ [2 SK], Resha, 28 Feb. 1979, leg. Heinz Stephan (SMF: 57532, 57533).
DEMOCRATIC REPUBLIC OF THE CONGO: 1 ad ♀ [SO], Alima, 15 Feb. 1959, leg. P. Dyleff
(RMCA: 28155b); 1 ? [AL], Babongena, between 1 May 1982 and 31 May 1982, leg. Marc Colyn
(RMCA: 82.031-M-0008); 1 ad ♀ [SO], Bokalakala, 23 Oct. 1959, leg. N’Kele (RMCA: 28542);
1 imm ♀ [SS], Bokungu, 11 Dec. 1948, leg. L. Dupuis (RMCA: 18715); 5 ad ♀♀, 3 ad ??, 2 imm ??
[2 SS, 4 SA, 2 AL, 2 SO], Boma, 15 Aug. 1922, leg. J. Rodhain, between 1 Apr. 1936 and 30 Sep. 1936,
between 1 Apr. 1936 and 30 Apr. 1936, leg. Schwetz (RMCA: 6066 to 6069, 13700a to 13700c, 13702
to 13704); 1 ad ♂ [SS], Eastern Congo, unknown date, leg. J.J. Laarman (RMNH: MAM.16342); 1 ♀
[SA], Faradje, 4 Apr. 1911, leg. Herbert Lang, James Paul Chapin and The American Museum Congo
Expedition (AMNH: 49422); 1 ad ♀, 2 ad ♂♂ [2 SA, 1 AL], Garamba National Park, 1 Jun. 1912, leg.
196
VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
Herbert Lang, James Paul Chapin and The American Museum Congo Expedition, 27 Dec. 1950, 12 May
1951, leg. H. de Saeger et al. (AMNH: 49399, RBINS: 13624, 13623); 1 ad ♂ [SO], Grotte de Baya
[= Baya Cave], 12 Jul. 1956, leg. M. Lips (RMCA: 23796); 2 ad ♀♀, 1 ad ♂, 1 juv ♂ [4 SS], Kakyelo,
13 Nov. 1930, leg. Gaston-François de Witte and de Crawhez (RMCA: 10845, 10848, 10849, 10852);
1 ad ♀ [SA], Kalina, prior to 16 Nov. 1950, leg. Jean Jacques Deheyn (RMCA: 20101); 1 ad ♂ [SS],
Kamueshi [= Kamuesha], 5 Apr. 1979, leg. Betu Nsankulu (RMCA: 79.032-M-0013); 1 juv ? [AL],
Kasongo, 23 Oct. 1959, leg. Segers and Benoit Mission (RMCA: 29164); 1 ? [AL], Katanga, near,
unknown date, leg. Félix / Michel Anciaux de Faveaux (MNHN: ZM-MO-1957-128); 16 ad ♀♀, 15 ad
♂♂, 1 ♀ [20 SS, 9 SA, 3 SO], Kateke [= Kateke [River]], 25 Nov. 1947, 26 Nov. 1947, 27 Nov. 1947,
1 Dec. 1947, 4 Dec. 1947, leg. Gaston-François de Witte et al. (RBINS: 10827, 10828, 10830 to 10834,
10845, 10813 to 10826, 10829, 10811.1 to 10811.9); 1 ♀ [UN], Katupila, 22 Oct. 1974, leg. Michael
D. Gallagher (HZM: 37.7849); 1 ad ♂ [SS], Kawezi-Uvira Road, 3 Jul. 1957, leg. J.J. Laarman
(RMNH: MAM.16369); 1 ad ♀, 3 ad ♂♂, 1 juv ♀, 1 juv ♂ [3 SA, 2 AL, 1 SO], Keyberg, 27 Oct. 1955,
20 Dec. 1955, 4 Jun. 1956, leg. M. Lips, 27 Oct. 1955, leg. Félix / Michel Anciaux de Faveaux (RMCA:
26277, 26278, 26275, 26276, SMF: 16875, RMCA: 23800); 1 ad ♀ [SS], Kidada, 17 Feb. 1922, leg.
Henri Schouteden (RMCA: 6556); 1 ad ♂ [SA], Kimilolo, between 1 Jan. 1931 and 31 Dec. 1931, leg.
De Smet (RMCA: 22441); 1 ♀, 2 ♂♂ [3 UN], Kinshasa, between 15 Jan. 1975 and 16 Jan. 1975, leg.
Michael D. Gallagher (HZM: 34.7846 to 36.7848); 1 sad ♂, 1 juv ♂ [2 AL], Léopoldville [= Kinshasa],
20 Jan. 1947, leg. Dartevelle, prior to 23 Jul. 1946, leg. Lepersonne (RMCA: 18163, 18129); 1 ad
♀ [SS], Luiko [= Lwiko], 13 Sep. 1950, leg. Alexandre Prigogine (RMCA: 20430); 1 ad ♂ [SO],
Luluabourg [= Kananga], 28 Nov. 1923, leg. Richard Callewaert (BMNH: 1926.7.6.75); 2 juv ??
[2 UN], Mabwe, 1 Mar. 1949, leg. unknown collector (RBINS: 10810.1, 10810.2); 1 ad ♂ [SK], Mabwe
River, 11 Feb. 1949, leg. Gaston-François de Witte et al. (RBINS: 10847); 1 ad ♀ [SS], Macaco,
29 Sep. 1921, leg. Henri Schouteden (RMCA: 7296); 1 juv ♂ [AL], Mahagi Port, 29 Mar. 1953, leg.
Célestin Hecq (RMCA: 21898); 5 ?? [5 SK], Maleli, 13 Jun. 1951, leg. H. de Saeger et al. (RBINS:
13625.1 to 13625.5); 1 ad ♀ [SA], Malili, 13 Jun. 1951, leg. H. de Saeger (RMCA: 36561); 1 ad
♀ [SK], Marari, 13 Jun. 1951, leg. H. de Saeger et al. (RBINS: 13626); 1 ad ♀ [SO], Morubia,
13 Jun. 1951, leg. H. de Saeger (RMCA: 36562); 1 ad ♀, 1 ad ♂ [2 SA], Musosa, near, 6 Jul. 1944,
between 1 Jun. 1943 and 30 Jun. 1943, leg. H.J. Bredo (RBINS: 14493, 14490); 2 ♀♀, 2 ♂♂ [4 UN],
Mwebe Bridge, 24 Jan. 1996, leg. Robert (Bob) Swanepoel (TM: 45614, 45615, 45617, 45618); 1 ad ?
[SS], Né, Mount, 25 Sep. 1952, leg. Alexandre Fain (RMCA: 23166); 1 ? [UN], Rutshuru, unknown
date, leg. unknown collector (INPBC: V1769); 3 ad ♀♀, 2 ad ♂♂ [5 SA], Shinkolobwe, 2 Aug. 1960,
leg. Félix / Michel Anciaux de Faveaux (RBINS: 14491, 14492, 14494, SMF: 19066, 19067); 2 ad ♀♀
[1 SS, 1 SK], St. Joseph de Luluabourg Mission [= Mikalaya], 19 Jun. 1923, between 19 Feb. 1922
and 19 Jun. 1922, leg. Richard Callewaert (RMCA: 7108, 7546); 1 ad ♀ [SS], Temvo, 8 Mar. 1922, leg.
Henri Schouteden (RMCA: 6557).
RWANDA: 1 ad ♀, 8 ad ♂♂ [8 SS, 1 SO], Astrida [= Butare], 6 Aug. 1954, 7 Aug. 1954, between
1 Jan. 1954 and 31 Dec. 1954, leg. Alexandre Fain (RMCA: 81.012-M-0005 to 81.012-M-0009, 81.012M-0001 to 81.012-M-0004); 1 ad ♀, 2 ad ♂♂ [3 SS], Bugundi, 19 Nov. 1955, leg. Alexandre Fain
(RMCA: 81.012-M-0011 to 81.012-M-0013); 1 ad ♂ [SA], Butare, 18 Jun. 1966, leg. Félix / Michel
Anciaux de Faveaux (RBINS: 15493); 1 ad ♀ [SA], Karama, 11 Oct. 1981, leg. Frits De Vree, Benny
Baeten and Victor Van Cakenberghe (RMCA: 82.006-M-0929); 1 ad ♀, 1 ad ♂, 1 sad ♀, 1 juv ♀ [3 SS,
1 SO], Musha, 17 Oct. 1955, between 1 May 1955 and 31 May 1955, leg. Alexandre Fain (RMCA:
81.012-M-0015 to 81.012-M-0018); 11 ad ♀♀, 7 ad ♂♂, 1 ad ?, 2 juv ♀♀, 1 juv ? [10 SS, 9 SA,
3 AL], Nyakibanda, 14 Oct. 1955, between 15 Oct. 1955 and 25 Oct. 1955, between 14 Oct. 1955 and
17 Oct. 1955, between 1 Apr. 1956 and 30 Apr. 1956, leg. Alexandre Fain, between 1 Mar. 1962 and
31 Mar. 1962, leg. Van Parijs (RMCA: 81.012-M-0019 to 81.012-M-0021, 81.012-M-0023 to 81.012M-0025, 31048 to 31059, 81.012-M-0027, 81.012-M-0028, 81.012-M-0022, 81.012-M-0026); 10 ad
♀♀, 10 ad ♂♂, 1 ad ?, 2 juv ♂♂, 2 ?? [12 SS, 7 SA, 2 AL, 2 SK, 2 SO], Nyumba Cave, unknown date,
197
European Journal of Taxonomy 382: 1–327 (2017)
between 1 Jan. 1954 and 31 Dec. 1954, leg. Alexandre Fain, 1 Feb. 1966, 7 Dec. 1966, 17 Dec. 1966, leg.
Félix / Michel Anciaux de Faveaux, 7 Dec. 1966, leg. unknown collector (BMNH: 1953.566, 1953.567,
1953.677 to 1953.680, RBINS: 15483.1 to 15483.4, 15484.1 to 15484.4, 15939.1, 15939.2, 15994,
15989 to 15993, 15995, MHNG: 921.036, 921.037); 1 ad ♀, 1 ad ♂ [2 SA], Simbi Cave, 10 Apr. 1968,
between 1 Mar. 1968 and 31 Mar. 1968, leg. Félix / Michel Anciaux de Faveaux (RBINS: 15931, 15930).
Nycteris major (K. Andersen, 1912)
DEMOCRATIC REPUBLIC OF THE CONGO: 1 ad ♂ [SA], Avakubi, 1 Sep. 1913, leg. Herbert Lang,
James Paul Chapin and The American Museum Congo Expedition (AMNH: 49403 [holotype Nycteris
avakubia J.A. Allen, 1917]); 1 ♂ [UN], Kona, 14 May 2010, leg. Guy-Crispin Gembu Tungaluna,
Prescott Musaba and José Akaibe (UNIKIS: CRT1959).
Nycteris nana (K. Andersen, 1912)
BURUNDI: 1 ♀ [SB], Giserama, Colline, 12 Aug. 1991, leg. Julian C. Kerbis Peterhans (FMNH:
148980).
DEMOCRATIC REPUBLIC OF THE CONGO: 1 ? [SK], unknown locality, unknown date, leg.
unknown collector (BMNH: 1930.11.11.83); 1 ad ♀, 1 ? [1 SA, 1 SK], Aka, 16 May 1952, leg. H. de
Saeger et al. (RBINS: 13565.1, 13565.2); 1 ad ♂ [SA], Bagisana, 19 Apr. 1952, leg. H. de Saeger et al.
(RBINS: 13563); 1 ad ♂ [SA], Bagunda, 18 Jan. 1951, leg. H. de Saeger et al. (RBINS: 13549); 1 ♀,
1 ♂ [1 AL, 1 SK], Batala’s Village, 13 Apr. 1925, 16 Apr. 1925, leg. Edmund Heller (FMNH: 30714,
30715); 1 ad ♀ [SA], Buluku River, 22 May 1952, leg. H. de Saeger et al. (RBINS: 13566); 1 ad ♀
[SS], Byangore River, 27 Apr. 1952, leg. Gaston-François de Witte et al. (RBINS: 17291); 1 ad ♂
[SA], Djugu Forest, between 1 Jan. 1948 and 31 Dec. 1948, leg. Jacques Verschuren (RBINS: 18546);
1 ? [SA], Duma, between 1 Oct. 1910 and 31 Oct. 1910, leg. unknown collector (SMF: 6390); 1 ad ♀,
1 ad ♂ [1 SS, 1 SA], Garamba National Park, 7 Feb. 1952, leg. H. de Saeger et al. (RBINS: 13554,
13556); 1 ad ♂ [AL], Ibembo, 20 Jul. 1950, leg. Jozef Hutsebaut (RMCA: 20092); 1 ad ♀ [SS], Irangi,
31 Jan. 1957, leg. J.J. Laarman (RMNH: MAM.16366); 1 ? [UN], Kambemba, unknown date, leg.
unknown collector (INPBC: V1065); 1 ad ? [SS], Kawa (Forest), 7 Aug. 1952, leg. Alexandre Fain
(BMNH: 1953.484); 1 ad ♀, 1 ad ♂ [2 SA], Kokodo River, 19 Apr. 1952, leg. H. de Saeger et al.
(RBINS: 13561.1, 13561.2); 1 juv ♀ [AL], Kokodu, 19 Apr. 1952, leg. H. de Saeger (RMCA: 36604);
1 ♂ [AL], Kumbi, 17 Nov. 1993, leg. Ngenge Masumbuko Kamitongo (ZMA: MAM.24898); 1 ad ♂
[SA], Luluabourg [= Kananga], 22 Apr. 1964, leg. Antoon Emeric Marcel De Roo (RMCA: 33600);
1 juv ♂ [AL], Makpe, 13 Jul. 1951, leg. H. de Saeger (RMCA: 36600); 1 ad ♀, 2 ad ♂♂ [3 SA], Makpe
River, upper [= Makpe], 13 Jul. 1951, leg. H. de Saeger et al. (RBINS: 13552.1, 13552.2, 13553); 2
ad ♀♀, 1 juv ? [1 SA, 2 SK], Maleli, 12 Apr. 1951, 18 Mar. 1952, leg. H. de Saeger et al. (RBINS:
13551, 13555.1, 13555.2); 1 ♀, 1 ♂, 1 ? [3 AL], Masako, unknown date, 16 Jul. 1989, leg. Ndey B. Ifuta
(RMCA: 93.079-M-0345, 93.079-M-0343, 93.079-M-0344); 1 ad ♂ [AL], Meridi River, 20 Apr. 1952,
leg. H. de Saeger (RMCA: 36605); 1 ♀, 1 ? [2 UN], Mongbwalu, 21 Jan. 2011, leg. unknown collector
(DNSM: 12754, 12755); 1 ad ♀, 1 ad ♂ [1 SA, 1 AL], Morubia, 14 Jun. 1951, 18 Mar. 1952, leg. H. de
Saeger (RMCA: 36601, 36603); 1 ad ♂, 1 ? [1 SA, 1 UN], Nakpanga, unknown date, leg. unknown
collector, 18 Apr. 1952, leg. H. de Saeger (INPBC: 4779.2.3, RMCA: 36608); 1 ad ♀ [SA], Nakpanga
River, 18 Apr. 1952, leg. H. de Saeger et al. (RBINS: 13564); 1 ad ♂ [SA], Pawa, between 1 Jan. 1931
and 31 Dec. 1931, leg. Radna (RMCA: 15314); 1 ad ♀ [SA], PFNK, 7 Feb. 1952, leg. H. de Saeger
(RMCA: 36602); 1 ad ♀ [SA], PFNK.15, 7 Feb. 1952, leg. H. de Saeger et al. (RBINS: 13557); 2 ad
♀♀, 1 ad ♂, 2 juv ??, 1 ♀, 1 ♂ [1 SA, 1 AL, 5 SK], Pidigala Nord River, 23 Apr. 1952, 24 Apr. 1952,
leg. H. de Saeger et al. (RBINS: 13558.1, 13558.2, 13560, 14258B, 19258A, 13559.1, 13559.2); 2 ad
♀♀, 1 ad ♂ [3 SA], Pidigala River, 23 Apr. 1952, 24 Apr. 1952, leg. H. de Saeger (RMCA: 36606,
36607, 36609); 1 ad ♀, 1 ad ♂, 1 ad ? [1 SS, 2 SA], PPK.10, 2 Aug. 1952, leg. H. de Saeger et al.
198
VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
(RBINS: 13562, 13567.1, 13567.2); 1 ♂ [UN], Rubi-Tele, 24 Aug. 2013, leg. Guy-Crispin Gembu
Tungaluna, Prescott Musaba and André Malekani (UNIKIS: RBTL554); 1 ? [AL], Tshakala, between
1 Jan. 1982 and 31 Dec. 1982, leg. Marc Colyn (RMCA: 82.019-M-0011); 2 ?? [2 AL], Wafanya
[=Wafania], 15 Apr. 1988, 21 Apr. 1992, leg. P. Lootens (RMCA: 89.020-M-0042, 92.079-M-0118);
1 ad ♀ [SA], Wilibadi River, 30 Jan. 1951, leg. H. de Saeger et al. (RBINS: 13550); 1 ♂ [SA],
Yalosemba, 30 Jun. 1979, leg. Frieder Meier (MHNG: 1870.097).
Nycteris parisii (de Beaux, 1924)
DEMOCRATIC REPUBLIC OF THE CONGO: 1 ? [SS], Né, Mount, 29 Sep. 1952, leg. unknown
collector (RBINS: 21668).
Nycteris thebaica E. Geoffroy St.-Hilaire, 1818
DEMOCRATIC REPUBLIC OF THE CONGO: 1 ? [UN], Albert National Park [= Virunga National
Park], unknown date, leg. unknown collector (INPBC: V1804); 1 ad ♀ [SA], Bulaga [= Kasenga
Road, km 36], 26 Mar. 1956, leg. Félix / Michel Anciaux de Faveaux (RMCA: 26230); 1 ad ♂ [SA],
Dethioux Hoeve, Grot I [= Shinkolobwe], 29 Sep. 1957, leg. Félix / Michel Anciaux de Faveaux
(MHNG: 1046.036); 7 ♀♀, 3 ♂♂, 1 ? [11 AL], Kadjidji [= Kajuju], unknown date, leg. Guy Babault
(MNHN: ZM-MO-1984-1347 to ZM-MO-1984-1357); 2 ad ♂♂ [1 SA, 1 UN], Kakanda Cave, unknown
date, between 21 Apr. 1955 and 22 Apr. 1955, leg. R.P. Th. de Caters (BMNH: 1955.146, RMCA: 22890);
3 ad ♀♀ [3 SS], Kakyelo, 13 Nov. 1930, leg. Gaston-François de Witte and de Crawhez (RMCA: 10844,
10846, 10847); 1 ad ♂ [SO], Kasenga Road, km 67, 8 Jun. 1956, leg. M. Lips (RMCA: 23798); 1 ad
♂ [SS], Kaswabilenga, 8 Oct. 1947, leg. Gaston-François de Witte et al. (RBINS: 10812); 1 ad ♂ [SA],
Lubudi-Lusolo, 16 Jul. 1962, leg. Félix / Michel Anciaux de Faveaux (SMF: 21243); 1 ? [AL], Lukatu
Cave [= Ntoto Cave], prior to 2 Feb. 2007, leg. A. Kimbembi-ma-Ibaka (RMCA: a7.006-M-0001);
1 ad ♂ [UN], Mahagi Port, unknown date, leg. J. Heck (BMNH: 1954.797); 1 ad ? [SA], Makayova
River, lower, unknown date, leg. Jacques Verschuren (INPBC: V1345); 1 ad ♂ [SA], Mount Ne [= Né,
Mount], 25 Sep. 1952, leg. Alexandre Fain (MHNG: 921.034); 1 ad ♂ [SO], Mukween [= Mukuen],
12 Jul. 1956, leg. M. Lips (RMCA: 23797); 1 ad ? [SS], Né, Mount, 25 Sep. 1952, leg. Alexandre Fain
(BMNH: 1953.485); 1 ad ? [AL], Rutshuru, unknown date, leg. Jacques Verschuren (INPBC: V1354);
1 ad ♀, 2 ad ?? [3 AL], Sinda River, unknown date, leg. Jacques Verschuren (INPBC: V481.1 to V481.3);
3 ad ♀♀, 1 ad ♂, 2 juv ♂♂ [4 SA, 2 AL], St Gerard Mission [= Kapolowe], 18 Oct. 1956, 8 Sep. 1962,
leg. Félix / Michel Anciaux de Faveaux (RMCA: 26234 to 26237, SMF: 16878, 21244); 1 ♀ [UN],
Tshambi, 10 Nov. 1933, leg. unknown collector (RBINS: 4781); 1 ad ? [SA], Tumbwe, unknown date,
leg. Jacques Verschuren (INPBC: V1436); 2 ♀♀, 3 ♂♂ [5 AL], Yalosemba, 17 Jun. 1979, 22 Jun. 1979,
leg. C. Brian Robbins (USNM: 463477 to 463481).
RWANDA: 1 ad ♀, 1 ad ♂ [2 SS], Astrida [= Butare], 25 Oct. 1966, 25 Dec. 1966, leg. Félix /
Michel Anciaux de Faveaux (RBINS: 16005, 16006); 1 ? [UN], Kagera National Park [= Akagera
Park], unknown date, leg. unknown collector (INPBC: V1904); 1 ad ♂ [SA], Kakitumba, unknown
date, leg. Jacques Verschuren (INPBC: V3131); 1 sad ♀, 1 juv ♂ [2 SA], Kidaho, 10 Jun. 1982, leg.
Benny Baeten (RMCA: 87.046-M-0344, 87.046-M-0345); 2 ad ♂♂ [2 AL], Kionja, unknown date,
leg. Jacques Verschuren (INPBC: V3085A, V3085B); 1 ? [UN], Kisenyi [= Gisenyi], unknown date,
leg. unknown collector (INPBC: 1804); 3 ad ♀♀, 3 ad ♂♂, 1 ad ? [5 SA, 2 AL], Muluchu, unknown
date, leg. Jacques Verschuren (INPBC: V3146A to V3146G); 1 ad ♀ [SA], Rukira, 22 Nov. 1981, leg.
Frits De Vree, Benny Baeten and Victor Van Cakenberghe (RMCA: 82.006-M-0930); 2 ad ♂♂ [2 SA],
Shangi Schad [= Shangi], 5 Jan. 1985, leg. Benny Baeten and Luc Janssens (RMCA: 85.006-M-0455,
85.006-M-0456); 1 neonate ♂ [UN], Simbi Cave, 6 Jul. 1968, leg. unknown collector (RBINS: 15936).
199
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Molossidae Gervais, 1856
DEMOCRATIC REPUBLIC OF THE CONGO: 7 ad ♀♀, 1 sad ♂, 96 ?? [104 AL], unknown locality,
unknown date, prior to 18 Aug. 1965, between 1 Jun. 1964 and 30 Jun. 1964, leg. Antoon Emeric Marcel
De Roo, between 1 Jan. 1976 and 31 Dec. 1977, leg. M. Lejeune (RMCA: 32780, 32782, 32785, 32786,
77.029-M-0040 to 77.029-M-0135, 32784, 33569 to 33571); 1 ? [SO], Butembo, between 1 Nov. 1971
and 30 Nov. 1971, leg. M. Lejeune (RMCA: 81.049-M-0002); 1 ? [SN], Cel II, 19 May 1952, leg. H. de
Saeger (RMCA: 36936); 1 ? [AL], Epulu, unknown date, leg. Luc Bijnens (RMCA: 90.031-M-0003);
1 ♂ [AL], Farm ‘des Grands Bois’ [= Likasi], 13 Nov. 1962, leg. Félix / Michel Anciaux de Faveaux
(RMCA: 33985); 8 ♀♀, 1 ♂, 1 ? [10 AL], Ganda-Sundi, 31 Mar. 1961, 31 Mar. 1964, leg. Alexandre
Fain (RMCA: 34195, 33263, 33266, 33267, 33269, 33270, 33272 to 33275); 1 ? [AL], Imbonga,
between 1 Jan. 1987 and 31 Dec. 1987, leg. P. Lootens (RMCA: 87.043-M-0002); 1 ? [SS], Ituri ht,
prior to 28 Apr. 1914, leg. Cuthbert Christy (RMCA: 3387); 1 ? [UN], Kabongo, prior to 27 Oct. 1952,
leg. M. Dierckx (RMCA: 21481); 8 ?? [8 AL], Kamponde, between 1 Apr. 1965 and 30 Apr. 1965,
leg. Antoon Emeric Marcel De Roo (RMCA: 33372 to 33379); 2 ad+foetus ♀♀ [2 AL], Keyberg,
9 Feb. 1956, leg. Félix / Michel Anciaux de Faveaux (RMCA: 90.011-M-0001, 90.011-M-0002); 1 ♂
[AL], Kinshasa, 1 Apr. 1987, leg. WHO and Marc Colyn (RMCA: 88.011-M-0027); 2 ad+foetus ♀♀
[2 AL], Kipushi, 15 km from [= Wasseige Farm], 6 Feb. 1956, leg. Félix / Michel Anciaux de Faveaux
(RMCA: 90.011-M-0003, 90.011-M-0004); 2 ♀♀, 2 ♂♂ [4 AL], Léopoldville [= Kinshasa], prior to
15 Jul. 1964, leg. Antoon Emeric Marcel De Roo and Jan Deheegher (RMCA: 32636 to 32639); 3 ??
[3 SS], Lovanium University [= Kinshasa], 1 Dec. 1962, 11 Jan. 1963, 23 Apr. 1963, leg. De Bont
and Jo Van Orshoven (RMCA: 32633, 32635, 32634); 85 ?? [85 AL], Lulimbi, between 1 Jan. 1976
and 31 Dec. 1977, leg. M. Lejeune (RMCA: 77.029-M-0029 to 77.029-M-0039, 77.029-M-0136 to
77.029-M-0209); 5 ♀♀, 5 ♂♂ [10 AL], Luluabourg [= Kananga], unknown date, leg. Jan Deheegher,
24 Oct. 1964, 3 Jul. 1965, prior to 6 May 1964, leg. Antoon Emeric Marcel De Roo (RMCA: 31561
to 31565, 31570, 33350, 33593, 33594, 31549); 1 ? [AL], Makaw, prior to 15 Nov. 1958, leg. E. Jans
(RMCA: 27499); 2 ?? [1 SK, 1 SO], Make, 1 Aug. 1989, between 1 Jan. 1992 and 31 Dec. 1992, leg.
Tshinyenye (RMCA: 89.057-M-0003, 92.148-M-0004); 8 ?? [8 AL], Masako, unknown date, leg. Ndey
B. Ifuta (RMCA: 93.079-M-0347 to 93.079-M-0354); 8 ♀♀, 2 ♂♂ [10 AL], Musingi [= Musindji],
between 1 Jan. 1972 and 31 Dec. 1972, leg. André Guissart (RMCA: 38521 to 38530); 1 ♂ [AL],
Nord-Kivu Province, prior to 28 Apr. 1974, leg. M.J. Celis and M. Lejeune (RMCA: 36198); 1 ♂
[AL], Uvira, 5 Sep. 1956, leg. Laurent (RMCA: 35693); 1 ? [AL], Wagenia, between 1 Jan. 1987 and
31 Dec. 1988, leg. Luc De Vos, Kimbembi and Alain Vandelanoote (RMCA: 88.025-M-0008).
RWANDA: 1 ♂, 1 ? [1 AL, 1 SO], Astrida [= Butare], between 1 Apr. 1952 and 30 Apr. 1952, leg.
Laurent, between 1 Jul. 1951 and 31 Jul. 1951, leg. Aureliaan (RMCA: 79.010-M-0007, 20568); 1 ♀
[AL], Ratabo Kinazi Gitorams, 30 Apr. 1971, leg. A. Elbl (RMCA: 35915).
Chaerephon Dobson, 1874
BURUNDI: 2 ♂♂ [2 AL], Mweya, 29 Apr. 1971, leg. P.L. Niort (MHNG: 1896.019, 1896.020).
DEMOCRATIC REPUBLIC OF THE CONGO: 17 ♀♀, 2 ♂♂ [15 AL, 4 SK], Kisantu, 3 Jun. 2012,
leg. William (“Bill”) T. Stanley (FMNH: 219519 to 219532, 219537 to 219541); 1 ? [SA], Léopoldville
[= Kinshasa], unknown date, leg. Cuthbert Christy (BMNH: 1904.6.25.1); 1 ♀ [SS], Upper Congo,
unknown date, leg. Alexander Douglas Mitchell Carruthers (BMNH: 1907.6.14.6).
Chaerephon aloysiisabaudiae (Festa, 1907)
DEMOCRATIC REPUBLIC OF THE CONGO: 1 imm ♂ [SS], Avakubi, 28 Dec. 1913, leg. Herbert
Lang, James Paul Chapin and The American Museum Congo Expedition (AMNH: 48934); 1 ♂ [SS],
200
VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
Eala, 2 Jan. 1921, leg. Henri Schouteden (RMCA: 6565); 1 ♂ [UN], Iso.4 [= Iso], 4 Mar. 1952, leg.
H. de Saeger et al. (RBINS: 13826); 1 ♂ [SS], Yalosemba, 17 Jun. 1979, leg. C. Brian Robbins (USNM:
537737).
Chaerephon ansorgei (Thomas, 1913)
DEMOCRATIC REPUBLIC OF THE CONGO: 2 ♂♂ [2 SK], Biadimbi, 20 Feb. 1951, leg. H. de
Saeger et al. (RBINS: 13775.1, 13775.2); 1 ad ♀, 1 ad ♂, 6 ♀♀, 5 ♂♂, 17 ?? [9 SS, 18 AL, 2 SK,
1 SO], Faradje, 1 Apr. 1952, leg. H. de Saeger et al., 9 Mar. 1912, 20 Feb. 1913, leg. Herbert Lang,
James Paul Chapin and The American Museum Congo Expedition, 20 Feb. 1913, leg. Herbert Lang
(RBINS: 13778.1, 13778.2, AMNH: 48902 to 48907, 48909, 49231 to 49234, 49236, 49238 to 49243,
49245 to 49249, FMNH: 43865, 43866, MCZ: 17220, RMCA: 12385, AMNH: 49264); 1 ♀ [SK],
Garamba National Park, 1 Jan. 1950, leg. H. de Saeger et al. (RBINS: 13780); 1 ♀ [AL], Jadotville
[= Likasi], 25 May 1959, leg. Félix / Michel Anciaux de Faveaux (RMCA: 28210); 25 ad ♀♀, 26
ad ♂♂, 1 ad? ♂, 1 ♀, 5 ♂♂ [29 SA, 2 AL, 27 UN], Kinzambi Mission, 25 Jul. 1995, 30 Jul. 1995,
31 Jul. 1995, leg. Herwig Leirs et al. (MSB: 274752 to 274754, 274756, RMCA: 97.021-M-0581 to
97.021-M-0585, 97.021-M-1173, MSB: 274757, 274758, 274760 to 274780, RMCA: 97.021-M-0586 to
97.021-M-0610); 1 ? [SA], Lake Kivu, near, between 1 Jan. 1935 and 31 Dec. 1935, leg. Guy Babault
(MCZ: 46156); 1 ad ♀, 1 ad ♂ [2 SA], Mbalaka (Plantation), 21 Jul. 1995, between 10 Jun. 1995 and
28 Aug. 1995, leg. Herwig Leirs et al. (RMCA: 97.021-M-0580, 97.021-M-0611); 1 ♀, 1 ♂, 6 ?? [2 SS,
6 SK], Mpaza, 24 Jan. 1952, leg. H. de Saeger et al., 24 Jan. 1952, leg. H. de Saeger (RBINS: 13776.1
to 13776.6, RMCA: 36891, 36892); 2 ♀♀, 2 ♂♂ [4 AL], Muradingusha [= Mwadingusha], unknown
date, leg. the Royal Museum for Central Africa (RMCA) (BMNH: 1959.502 to 1959.505); 14 ♀♀, 8
♂♂ [8 SA, 10 AL, 4 UN], Mwadingusha, 16 Jul. 1957, 20 May 1959, leg. Félix / Michel Anciaux de
Faveaux (MHNG: 1046.025 to 1046.028, RMCA: 26757 to 26772, 28208, 28209); 31 ad ♀♀, 30 ad
♂♂ [28 SA, 2 SK, 31 UN], Sacred Heart Mission, 2 Aug. 1995, leg. Herwig Leirs et al. (MSB: 274721
to 274726, 274781 to 274805, RMCA: 97.021-M-0612 to 97.021-M-0641); 4 ?? [1 SS, 3 SK], Uduku,
24 Jul. 1952, leg. H. de Saeger et al., 24 Jul. 1952, leg. H. de Saeger (RBINS: 13779.1 to 13779.3,
RMCA: 36893).
RWANDA: 1 ad ♂ [SK], Gabiro, 19 Jan. 1938, leg. unknown collector (RBINS: 4907); 2 ♀♀, 9 ♂♂
[11 SS], Rukara, between 27 Aug. 1955 and 31 Dec. 1955, between 16 Sep. 1955 and 31 Dec. 1955,
between 15 Sep. 1955 and 31 Dec. 1955, leg. Alexandre Fain (RMCA: 81.012-M-0050 to 81.012-M0057, 81.012-M-0049, 81.012-M-0058, 81.012-M-0059).
Chaerephon bemmeleni (Jentink, 1879)
DEMOCRATIC REPUBLIC OF THE CONGO: 1 ? [UN], Lume River, near, unknown date, leg. unknown
collector (INPBC: V857); 1 ♀ [SS], Rutshuru, 16 Jan. 1937, leg. J. Ghesquière (RMCA: 13826).
Chaerephon bivittatus (Heuglin, 1861)
DEMOCRATIC REPUBLIC OF THE CONGO: 1 imm ? [SS], Bitshumbi [= Vitshumbi], prior to
22 Dec. 1943, leg. Gaston-François de Witte (RMCA: 17076).
Chaerephon chapini J.A. Allen, 1917
DEMOCRATIC REPUBLIC OF THE CONGO: 1 ♂ [UN], Banana, between 1 Jun. 1948 and
30 Jun. 1948, leg. unknown collector (RBINS: 14535); 1 ad ♂, 1 ? [1 SS, 1 SA], Faradje, 6 May 1910,
11 Nov. 1912, leg. Herbert Lang, James Paul Chapin and The American Museum Congo Expedition
(AMNH: 48841 [holotype Chærephon (Lophomops) chapini J.A. Allen, 1917], 49209); 1 ♀ [UN],
Léopoldville [= Kinshasa], 15 Mar. 1964, leg. unknown collector (RBINS: 14536).
201
European Journal of Taxonomy 382: 1–327 (2017)
RWANDA: 1 ♀ [UN], Masha, 1 Jan. 1956, leg. unknown collector (RBINS: 22466).
Chaerephon gallagheri (Harrison, 1975)
DEMOCRATIC REPUBLIC OF THE CONGO: 1 ad ♂ [UN], Scierie Forest, 14 Nov. 1974, leg. Michael
D. Gallagher (BMNH: 1976.207 [holotype Tadarida (Chaerephon) gallagheri Harrison, 1975]).
Chaerephon leucogaster (A. Grandidier, 1869)
DEMOCRATIC REPUBLIC OF THE CONGO: 6 ♀♀, 6 ♂♂ [1 SS, 11 AL], Boende, 27 Jun. 2013,
4 Jul. 2013, 6 Jul. 2013, leg. William (“Bill”) T. Stanley (FMNH: 222431 to 222434, 222440 to 222443,
222472 to 222474, 222629); 1 ad ♂, 1 ♂, 2 ?? [1 SS, 2 SA, 1 AL], Boma, 25 Jan. 1915, 26 Jan. 1915,
29 Jan. 1915, 15 Jun. 1915, leg. Herbert Lang, James Paul Chapin and The American Museum Congo
Expedition (AMNH: 49259, 49237, 48844 [holotype Chærephon (Lophomops) cristatus J.A. Allen,
1917], 49216); 1 ? [SA], Lukolela, 27 Sep. 1930, leg. James Paul Chapin (AMNH: 86949); 1 ad ♀,
2 ♀♀, 22 ?? [9 SA, 16 AL], Malela, 6 May 1910, 8 Jul. 1915, leg. Herbert Lang, James Paul Chapin
and The American Museum Congo Expedition (AMNH: 49210 to 49212, 49214, 49215, 49217, 49218,
49220, 49221, 49223, 49226, 49227, 49229, 49261, 49263, 49268 to 49271, 49273, 49274, 49275
[holotype Chærephon frater J.A. Allen, 1917], 49276, MCZ: 17219 [topotype Chærephon frater
J.A. Allen, 1917], 17221 [topotype Chærephon frater J.A. Allen, 1917]).
Chaerephon major (Trouessart, 1897)
DEMOCRATIC REPUBLIC OF THE CONGO: 1 ad ♀, 16 ♀♀, 5 ♂♂, 11 ?? [20 SS, 11 SK, 2 SO],
Aba, 12 Dec. 1910, 13 Dec. 1911, 15 Dec. 1911, 16 Dec. 1911, 17 Dec. 1911, leg. Herbert Lang, James
Paul Chapin and The American Museum Congo Expedition, 15 Dec. 1911, leg. unknown collector
(AMNH: 48867, 48869 to 48872, 51257 to 51267, MCZ: 17389, RMCA: 12365, AMNH: 48873, 48840,
48875 to 48886, 48887 [holotype Chærephon (Lophomops) abæ J.A. Allen, 1917]); 3 ♂♂ [1 AL,
2 UN], Biadimbi, 20 Feb. 1951, leg. H. de Saeger et al., 20 Feb. 1951, leg. H. de Saeger (RBINS:
13783.1, 13783.2, RMCA: 36895); 1 ad ♀, 4 ♀♀ [4 SS, 1 UN], Faradje, 25 Feb. 1911, 7 Mar. 1911,
leg. Herbert Lang, James Paul Chapin and The American Museum Congo Expedition, 7 Mar. 1911, leg.
unknown collector (AMNH: 48836, 52119, RMCA: 12364, AMNH: 48838, MCZ: 17388); 1 ♂ [UN],
Mbandaka, prior to 31 Dec. 1970, leg. unknown collector (RBINS: 22467); 4 ♀♀ [1 SS, 3 AL], Nagera
[= Nagero], 24 Mar. 1952, leg. H. de Saeger (RMCA: 36894, 36896 to 36898); 1 ♀ [SO], Nagero,
11 Mar. 1954, leg. unknown collector (RBINS: 14630); 1 ♀ [SA], Rwindi, 2 Sep. 1976, leg. Frieder
Meier (MHNG: 1874.059); 1 ♀, 11 ♂♂ [5 SS, 7 AL], Tandala, 12 Jul. 1979, 13 Jul. 1979, 21 Jul. 1979,
leg. unknown collector, 12 Jul. 1979, 21 Jul. 1979, leg. C. Brian Robbins (MHNG: 1870.098, USNM:
463518 to 463521, 537748 to 537750, MHNG: 1870.099, 1870.100, 1871.001, USNM: 537752).
Chaerephon nigeriae Thomas, 1913
DEMOCRATIC REPUBLIC OF THE CONGO: 1 imm ?, 1 ♀ [1 AL, 1 UN], Elisabethville
[= Lubumbashi], 1 Jun. 1969, leg. unknown collector, between 1 Apr. 1926 and 30 Apr. 1926, leg. Henri
Schouteden (RBINS: 18511, RMCA: 16191); 1 ♀, 2 ♂♂, 1 ? [4 SS], Gangala-na-Bodio, 8 Apr. 1945,
13 Nov. 1947, leg. unknown collector (RBINS: 12952, 12949 to 12951); 1 ♀, 2 ♂♂ [2 SA, 1 AL],
Kabongo, unknown date, leg. the Royal Museum for Central Africa (RMCA), prior to 27 Oct. 1952,
leg. M. Dierckx (BMNH: 1954.836, RMCA: 21480, 21482); 2 ad ♀♀ [1 SA, 1 AL], Katanga, unknown
date, leg. Félix / Michel Anciaux de Faveaux (MNHN: ZM-MO-1957-132 A, ZM-MO-1957-132 B);
1 imm ♀, 1 imm ?, 4 ♀♀, 3 ?? [1 SA, 8 AL], Kinke, unknown date, leg. the Royal Museum for Central
Africa (RMCA), 23 Dec. 1955, leg. Félix / Michel Anciaux de Faveaux, 25 Oct. 1955, 23 Dec. 1955,
leg. M. Lips (BMNH: 1957.434, MHNG: 923.008, RMCA: 26288, 26291 to 26295, 26289); 1 ♂ [SS],
Kisiushi, 30 Jul. 1956, leg. M. Lips (RMCA: 33279); 4 ?? [4 AL], Kitubulushi, 3 Aug. 1954, leg.
202
VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
De Smet (RMCA: 22462 to 22465); 1 ♂ [AL], Musonge, 29 Dec. 1955, leg. M. Lips (RMCA: 26290);
1 ♀ [UN], Musosa, near, between 1 Jan. 1941 and 31 Dec. 1941, leg. unknown collector (RBINS:
14534); 1 sad ♂ [SS], Niangara, 10 Dec. 1910, leg. Herbert Lang, James Paul Chapin and The American
Museum Congo Expedition (AMNH: 48842); 3 ♀♀ [3 AL], Sakania, unknown date, leg. the Royal
Museum for Central Africa (RMCA) (BMNH: 1954.837 to 1954.839); 1 juv ♂ [SA], Shaba (former)
Province [= Katanga (former) Province], 23 Dec. 1955, leg. unknown collector (SMF: 16881); 1 ♀,
1 ♂ [1 SS, 1 AL], Tandala, 12 Jul. 1979, 21 Jul. 1979, leg. C. Brian Robbins (USNM: 537751, 463522).
Chaerephon pumilus (Cretzschmar, 1826)
BURUNDI: 1 ? [AL], Bujumbura, 10 Nov. 1968, leg. Lewall (RMCA: 35491); 1 ? [AL], Bururi,
between 24 May 1956 and 30 May 1956, leg. Laurent (RMCA: 26564); 1 ad ♂ [AL], Imbo, unknown
date, leg. Jacques Vissault (MNHN: ZM-MO-1996-1539); 3 ?? [3 SA], Kinazi, between 5 Jan. 1953
and 8 Jan. 1953, leg. P. Basilewsky (RMCA: 21936 to 21938); 1 ♀ [SK], Lake Cohoha [= Cohoha,
Lake], 9 Mar. 1979, leg. Heinz Stephan (SMF: 57559); 1 ? [SA], Nyankende, between 11 Mar. 1952
and 14 Mar. 1952, leg. Laurent (RMCA: 79.010-M-0010); 2 ad ♀♀, 4 ♀♀, 1 ♂ [4 SA, 3 SK], Resha,
1 Mar. 1979, 15 Mar. 1979, 2 Apr. 1979, 3 Apr. 1979, leg. Heinz Stephan (SMF: 57560 to 57566);
1 ? [AL], Russissi, bank [= Rusizi Delta], 16 Nov. 1968, leg. Lewall (RMCA: 35492); 4 ?? [4 UN],
Usumbura [= Bujumbura], 1 Aug. 1976, leg. unknown collector (RBINS: 19926.1 to 19926.4).
DEMOCRATIC REPUBLIC OF THE CONGO: 6 juv ♂♂, 1 ♀, 2 ♂♂ [8 AL, 1 SO], unknown locality,
between 16 Jan. 1956 and 19 Jan. 1956, leg. J.J. Laarman, prior to 31 Dec. 1889, leg. P. Hesse
(RMNH: MAM.16373, SMF: 34273 to 34280); 2 ?? [2 UN], Albert National Park [= Virunga National
Park], unknown date, leg. unknown collector (INPBC: V1805, V1844); 1 ♂, 2 ?? [2 AL, 1 UN],
Albertville [= Kalemie], 19 Sep. 1947, leg. unknown collector, prior to 18 Nov. 1959, leg. H. Bomans
(RBINS: 9071, RMCA: 29305, 29306); 3 juv ??, 29 ?? [28 AL, 4 SK], Banana, 13 Mar. 1952,
12 Apr. 1952, between 1 May 1952 and 31 May 1952, leg. Is. Mesmaekers, prior to 31 Dec. 1889, leg.
P. Hesse (RMCA: 20765 to 20778, 20753 to 20760, 21378 to 21383, SMF: 2527 to 2530); 1 ? [AL], Bas
Congo (former) Province [= Kongo Central Province], unknown date, leg. Alphonse Cabra (RMCA:
66); 2 ?? [2 UN], Beno, between 1 Jan. 1949 and 31 Dec. 1949, leg. P. Bayet (RMCA: 19824, 19825);
1 ? [SO], Bokalakala, 18 Nov. 1959, leg. N’Kele (RMCA: 28543); 9 ♀♀, 4 ♂♂ [13 SS], Bolobo,
27 Mar. 1921, leg. Henri Schouteden, prior to 23 Jul. 1946, leg. N’Kele (RMCA: 6577, 6581 to 6587,
7252 to 7254, 7308, 17403); 1 juv ?, 5 ♀♀, 1 ♂, 39 ?? [9 SS, 33 AL, 2 SO, 2 UN], Boma, unknown date,
23 Nov. 1920, 24 Nov. 1920, 25 Nov. 1920, between 1 Aug. 1920 and 31 Aug. 1920, between 1 Jan. 1922
and 31 Dec. 1933, between 1 Jan. 1922 and 31 Dec. 1922, leg. Henri Schouteden, between 1 Aug. 1928
and 31 Aug. 1928, leg. Gerard, between 1 Sep. 1954 and 30 Sep. 1954, leg. Cassart, prior to 26 May
1933, leg. Colsoulle, prior to 4 Aug. 1937, leg. Dartevelle (RMCA: 16491, 6598, 6604, 6605, 7301 to
7303, 7251, 16234, 16235, 16253, 7719 to 7722, 16202, 16269, 16270, 22388, 12128a to 12128j,
12129a to 12129m, 13959 to 13962); 5 ?? [5 AL], Bondo, between 1 Nov. 1957 and 30 Nov. 1957, leg.
Roland (RMCA: 26918 to 26922); 1 ? [UN], Bongeya River, unknown date, leg. unknown collector
(INPBC: V264.4); 1 ? [AL], Boteka, 31 Mar. 1985, leg. P. Lootens (RMCA: 85.030-M-0002); 18 ??
[17 AL, 1 UN], Bukama, unknown date, prior to 8 Apr. 1939, between 1 Jun. 1939 and 30 Jun. 1939,
between 1 Jun. 1937 and 30 Jun. 1937, leg. Brien (RMCA: 16254, 15261, 15251 to 15260, 15262 to
15267); 3 ?? [3 UN], Bumali, unknown date, leg. unknown collector (INPBC: W138.01 to W138.03);
1 ? [AL], Busu Mokalu [= Busu-Mokala], between 5 Nov. 1938 and 10 Nov. 1938, leg. Baudart
(RMCA: 18097); 1 ♂, 2 ?? [1 SS, 2 AL], Eala, unknown date, leg. Mayné, 3 Jan. 1921, leg. Henri
Schouteden, between 1 Mar. 1921 and 31 Mar. 1921, leg. unknown collector (RMCA: 5392, 6603,
26285); 1 ? [AL], Elisabethville [= Lubumbashi], between 1 Jan. 1933 and 31 Dec. 1933, leg.
Kerkvoorde (RMCA: 12140); 3 foetus ??, 9 ?? [12 AL], Equateur Province, between 1 Jan. 1921 and
31 Dec. 1921, leg. Verlaine (RMCA: 88.044-M-0001 to 88.044-M-0012); 4 ?? [2 SA, 2 AL], Ganda203
European Journal of Taxonomy 382: 1–327 (2017)
Sundi, 25 Mar. 1964, 31 Mar. 1964, leg. Alexandre Fain (RMCA: 33278, 33264, 33265, 33271); 1 ?
[AL], Irangi, 8 Feb. 1984, leg. Heinz Stephan (SMF: 65039); 134 ?? [34 AL, 100 UN], Ishango River,
unknown date, leg. unknown collector, between 1 Jan. 1982 and 31 Dec. 1982, leg. Marc Colyn, between
26 Dec. 1968 and 29 Dec. 1968, leg. M. Lejeune (INPBC: V1176.043 to V1176.142, RMCA: 82.031M-0002 to 82.031-M-0004, 35460 to 35490); 1 ? [UN], Kabakuli River, unknown date, leg. unknown
collector (INPBC: W253); 2 ♀♀, 1 ♂, 7 ?? [9 SS, 1 SO], Kabalo, unknown date, leg. Henri Schouteden,
17 Nov. 1926, leg. John Todd Zimmer (RMCA: 9164, 9165, 9167 to 9172, 9904, FMNH: 29429); 9 ♀♀
[8 SO, 1 UN], Kahitumba (Mutaka), between 16 Jan. 1956 and 19 Jan. 1956, leg. J.J. Laarman
(RMNH: MAM.16374 to MAM.16382); 7 ?? [7 AL], Kalina, prior to 16 Oct. 1952, leg. Theunissen,
prior to 16 Nov. 1950, leg. Jean Jacques Deheyn (RMCA: 21416, 21417, 21418a, 21418b, 21419, 21420,
20102); 1 ? [AL], Kambukabakali, 4 Nov. 1954, leg. Gaston-François de Witte (RMCA: 37383); 1 ♀
[SA], Kapolowe, 3 Aug. 1962, leg. Félix / Michel Anciaux de Faveaux (SMF: 21313); 1 ♂ [SS],
Kapushia, 3 Dec. 1973, leg. Western New Mexico University (FMNH: 152991); 11 ♀♀, 9 ♂♂, 3 ??
[2 SS, 1 AL, 20 UN], Kasenyi [= Kasenye], unknown date, 10 Feb. 1953, 16 Feb. 1953, 6 May 1953,
15 Jun. 1953, 26 Jun. 1953, 24 Sep. 1953, 20 Nov. 1953, 23 Nov. 1953, leg. unknown collector,
19 Feb. 1954, leg. Gaston-François de Witte, 30 Jul. 1951, leg. Alexandre Fain, prior to 16 Dec. 1949,
leg. Jean Hiernaux (INPBC: W1389, RBINS: 14539, 14550, 14555, RMCA: 37382, RBINS: 14541,
14552, 14538, 14547, 14554, 14551, RMCA: 23297, RBINS: 14537, 14540, 14543 to 14546, 14549,
14553, 14556, 14557, RMCA: 19099); 5 ?? [1 SA, 4 AL], Kashiobwe, 20 Oct. 1955, leg. Van Iseghem,
30 Oct. 1955, leg. unknown collector (RMCA: 26282 to 26284, 26286, SMF: 16877); 1 juv ?, 1 ♀
[2 AL], Kashiobwe Mission [= Kashiobwe], 20 Oct. 1955, leg. Félix / Michel Anciaux de Faveaux
(RMCA: 26227, 26266); 3 ?? [3 AL], Kasongo, unknown date, leg. Van Delft, 13 Nov. 1959, leg. Segers
and Benoit Mission (RMCA: 16201, 29166, 29308); 1 ? [AL], Katanga, prior to 31 Aug. 1900, leg.
Charles Lemaire (RMCA: 424); 1 ♂, 1 ? [2 SS], Kawa (Forest), 24 Jul. 1952, between 1 Oct. 1950 and
31 Oct. 1950, leg. Alexandre Fain (RMCA: 81.012-M-0062, 23296); 1 ? [AL], Keyberg, 27 Oct. 1955,
leg. M. Lips (RMCA: 26287); 3 ♂♂ [3 SS], Kidada, 20 Feb. 1922, 21 Feb. 1922, leg. Henri Schouteden
(RMCA: 6588, 6589, 6578); 54 ad ♀♀, 28 ad ♂♂, 5 ad? ♀♀, 3 ad? ♂♂ [46 SA, 44 UN], Kikwit,
6 Jul. 1995, 12 Jul. 1995, 14 Jul. 1995, leg. Herwig Leirs et al. (MSB: 274598 to 274600, RMCA:
97.021-M-0656 to 97.021-M-0659, MSB: 274607 to 274612, 274615 to 274620, 274623 to 274634,
274639 to 274642, RMCA: 97.021-M-0660 to 97.021-M-0687, MSB: 274579 to 274583, 274589 to
274593, 274595 to 274597, RMCA: 97.021-M-0642 to 97.021-M-0655); 1 ad ♀ [UN], Kikwit ITPK
[= Kikwit], 1 Aug. 1995, leg. Herwig Leirs et al. (MSB: 274882); 16 ad ♀♀, 6 ad ♂♂, 1 ad? ♂, 4 ♀♀,
3 ♂♂ [17 SA, 13 UN], Kinzambi Mission, 24 Jul. 1995, 25 Jul. 1995, 26 Jul. 1995, 27 Jul. 1995,
28 Jul. 1995, 30 Jul. 1995, leg. Herwig Leirs et al. (MSB: 274858 to 274861, RMCA: 97.021-M-0688
to 97.021-M-0693, MSB: 274863, 274864, RMCA: 97.021-M-0694, 97.021-M-0695, MSB: 274865 to
274867, RMCA: 97.021-M-0718 to 97.021-M-0720, MSB: 274869, RMCA: 97.021-M-0721, 97.021M-0722, MSB: 274870 to 274872, RMCA: 97.021-M-0723 to 97.021-M-0726); 2 ♀♀, 8 ♂♂, 1 ? [8 SS,
1 AL, 2 SO], Kisantu, 11 Dec. 1920, 12 Dec. 1920, 15 Dec. 1920, 18 Dec. 1920, 19 Dec. 1920, leg.
Henri Schouteden, prior to 7 Nov. 1932, leg. R.P. Vanderijst (RMCA: 7300a, 6606, 6599 to 6602, 7306,
7304, 7305, 7307, 11956); 1 ? [AL], Kivu region, between 1 Jan. 1962 and 31 Dec. 1962, leg. unknown
collector (SMF: 33527); 2 ♀♀, 1 ♂ [3 AL], Ko[g]enga, unknown date, leg. the Royal Museum for
Central Africa (RMCA) (BMNH: 1954.831 to 1954.833); 2 ?? [2 AL], Kondue [River], unknown date,
leg. Luja (RMCA: 3435, 3436); 12 ♀♀, 4 ♂♂, 1 ? [14 SS, 2 SO, 1 UN], Kunungu, unknown date, leg.
unknown collector, unknown date, prior to 6 Aug. 1925, prior to 25 Mar. 1929, prior to 1 Sep. 1924,
between 1 Jan. 1922 and 31 Dec. 1922, leg. Ngwe, between 1 Jan. 1922 and 31 Dec. 1922, leg. Henri
Schouteden, prior to 10 Dec. 1932, prior to 1 Aug. 1939, leg. N’Kele (RMCA: 7437 to 7439, 7659,
7256, 7751, 15516, 7658, 12002, 9709, 9710, 8166 to 8171); 1 ? [AL], Lemba Plain, 3 Jul. 1929, leg.
Van Delft (RMCA: 16272); 1 emb ?, 1 ♀, 3 ♂♂, 17 ?? [2 SS, 16 AL, 3 SO, 1 UN], Léopoldville
[= Kinshasa], unknown date, leg. J. Camp, 15 Jan. 1919, 30 Sep. 1941, between 1 Mar. 1942 and
31 Mar. 1942, between 1 Jan. 1942 and 31 Jan. 1942, leg. R. Fiasse, 4 Apr. 1964, leg. Antoon Emeric
204
VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
Marcel De Roo, prior to 12 Mar. 1913, leg. Cuthbert Christy, prior to 23 Jul. 1946, leg. Lepersonne
(USNM: 21668 to 21675, RMCA: 18125, 18126, 17657, 17658, 32658, 17660, 17659, 1532a, 1532b,
1533a, 1533b, 1534, 1535, 18130); 1 ? [SS], Lovanium University [= Kinshasa], unknown date, leg.
De Bont (RMCA: 31202); 23 ?? [23 SA], Lulimbi, between 1 Jan. 1976 and 31 Dec. 1977, leg.
M. Lejeune (RMCA: 77.029-M-0006 to 77.029-M-0028); 3 ♂♂ [3 SS], Mabwe, 2 Dec. 1948,
31 Jan. 1949, leg. unknown collector, 21 Aug. 1947, leg. l’ Institut des Parcs nationaux du Congo belge
(RBINS: 10712, 10711, 10713); 8 ♀♀, 2 ♂♂, 8 ?? [9 SS, 8 AL, 1 SO], Makaia N’Tete, 30 Mar. 1922,
between 1 Jan. 1922 and 31 Dec. 1922, leg. Henri Schouteden (RMCA: 6579, 6580, 6590 to 6597,
16236 to 16242, 16259); 1 ? [AL], Makaw, prior to 15 Nov. 1958, leg. E. Jans (RMCA: 27500); 8 ??
[8 AL], Makengo, 11 Dec. 1946, leg. Dartevelle (RMCA: 18152 to 18159); 1 ♂ [UN], Maleli,
10 Jun. 1951, leg. H. de Saeger et al. (RBINS: 13781); 1 ? [UN], Mangbwelu River, unknown date, leg.
unknown collector (INPBC: W240c); 2 ♀♀ [2 UN], Manono, 1 Nov. 1974, leg. Michael D. Gallagher
(HZM: 127.7821, 143.7954); 2 ?? [2 AL], Manono Katanga [= Manono], prior to 11 Dec. 1953, leg.
Torfs (RMCA: 22000, 22001); 1 ad? ♀ [UN], Mbalaka (Plantation), 19 Jul. 1995, leg. Herwig Leirs
et al. (MSB: 274827); 1 ♀ [SS], Mbuji-Mayi, 3 Nov. 1973, leg. Western New Mexico University
(FMNH: 152992); 1 ad ? [UN], Mbwambala, 19 Jul. 1995, leg. Herwig Leirs et al. (MSB: 274613); 1 ?
[AL], Moliro, prior to 28 Feb. 1899, leg. Charles Lemaire (RMCA: 161); 2 ad ??, 1 juv ?, 3 ♀♀, 1 ♂,
24 ?? [25 SS, 3 AL, 3 UN], Mount Wago [= Wago, Mount], unknown date, 13 Sep. 1952, 16 Sep. 1952,
17 Sep. 1952, 18 Sep. 1952, 21 Sep. 1952, 23 Sep. 1952, prior to 28 Mar. 1952, between 1 Jul. 1952 and
31 Jul. 1952, between 1 Jan. 1951 and 31 Dec. 1951, leg. Alexandre Fain, 1 Nov. 1951, 17 Sep. 1952,
1 Nov. 1952, leg. unknown collector (BMNH: 1953.493 to 1953.495, RBINS: 21653, 21654, 21656,
RMCA: 81.012-M-0078, 81.012-M-0079, 81.012-M-0083, 81.012-M-0084, 81.012-M-0087 to 81.012M-0090, 81.012-M-0100, 81.012-M-0081, 81.012-M-0085, 81.012-M-0096, RBINS: 21655, RMCA:
81.012-M-0086, 81.012-M-0092, 81.012-M-0080, 81.012-M-0082, 21465, 21450 to 21454, 20786,
20787); 15 ?? [15 UN], Mulongo, 28 Oct. 1974, leg. Michael D. Gallagher (HZM: 128.7930 to
142.7944); 1 juv ♂ [SS], Mvuazi [= Maunzi], between 1 Sep. 1942 and 30 Sep. 1942, leg. J.M. Vrijdagh
(RMCA: 17757); 2 ?? [2 AL], Nagera [= Nagero], 4 May 1951, leg. H. de Saeger (RMCA: 36889,
36890); 4 ?? [4 UN], Nagero, 4 May 1951, leg. H. de Saeger et al. (RBINS: 13782.1 to 13782.4); 1 ♂,
1 ? [1 SS, 1 SO], Né, Mount, between 1 Jun. 1951 and 30 Jun. 1951, between 1 Jan. 1950 and
31 Dec. 1950, leg. Alexandre Fain (RMCA: 23298, 20509 [holotype Tadarida (Chaerephon) faini
Hayman, 1951]); 1 ♂ [UN], Ngeze Kidoko, 26 Sep. 1953, leg. unknown collector (RBINS: 14548); 1 ?
[AL], Ngwiro, 22 Nov. 1956, leg. Gaston-François de Witte (RMCA: 37381); 3 ?? [3 SS], Nionga,
3 May 1925, leg. Gaston-François de Witte (RMCA: 9231 to 9233); 2 ?? [2 UN], Rwindi, unknown
date, leg. unknown collector (INPBC: V1545, W231); 15 ad ♀♀, 4 ad ♂♂ [10 SA, 9 UN], Sacred Heart
Mission, 2 Aug. 1995, leg. Herwig Leirs et al. (MSB: 274873 to 274881, RMCA: 97.021-M-0727 to
97.021-M-0736); 2 ♂♂ [1 SS, 1 SO], St. Joseph de Luluabourg Mission [= Mikalaya], 6 May 1939,
leg. Jean Jacques Deheyn, 7 Jun. 1922, leg. Richard Callewaert (RMCA: 15376, 7543); 1 ? [AL],
Stanleyville [= Kisangani], prior to 31 Jul. 1899, leg. Weyns (RMCA: 283); 10 ♂♂, 66 ?? [76 AL],
Thysville [= Mbanza-Ngungu], unknown date, leg. the Royal Museum for Central Africa (RMCA),
23 Jul. 1949, 8 Aug. 1949, 18 Aug. 1949, between 1 Aug. 1949 and 31 Aug. 1949, leg. Narcisse Leleup,
between 1 Dec. 1952 and 31 Dec. 1952, leg. P. Basilewsky (BMNH: 1954.821 to 1954.830, RMCA:
22570 to 22583, 22525 to 22550, 22515 to 22524, 22551 to 22565, 21618); 43 ad ♀♀, 20 ad ♂♂, 4 ♂♂
[33 SA, 34 UN], Trappist Mission, 27 Jul. 1995, 6 Aug. 1995, leg. Herwig Leirs et al. (MSB: 274806
to 274814, 274883 to 274895, RMCA: 97.021-M-0696 to 97.021-M-0717, MSB: 274815 to 274826,
RMCA: 97.021-M-0737 to 97.021-M-0747); 4 ?? [4 AL], Tshibambo [= Shibambo], 31 Aug. 1954,
leg. De Smet (RMCA: 22466 to 22469); 1 ♀ [SS], Usambiers, unknown date, leg. Emin Pasha (BMNH:
1890.6.8.16); 2 ♂♂, 14 ?? [1 SS, 15 AL], Uvira, 2 Mar. 1955, 3 Mar. 1955, prior to 21 Jun. 1958,
between 1 Mar. 1955 and 31 Mar. 1955, leg. Narcisse Leleup, 5 Mar. 1959, 29 Apr. 1959, leg. Uwe
Rahm (RMCA: 27206, 27207, 28908, 28909, 27204, 27205, 27208, 28907, 22909, 22910, 27215 to
27219, 27221); 3 ?? [3 SS], Vaku, prior to 8 Aug. 1940, leg. C. Vleeschouwers (RMCA: 16151 to
205
European Journal of Taxonomy 382: 1–327 (2017)
16153); 1 ♂ [UN], Vitshumbi, 21 Apr. 1953, leg. unknown collector (RBINS: 14542); 1 ♀, 1 ♂, 13 ??
[13 SS, 2 SO], Wago, Mount, 7 Nov. 1951, 28 Aug. 1952, 30 Aug. 1952, 13 Sep. 1952, 17 Sep. 1952,
between 1 Nov. 1951 and 30 Nov. 1951, between 1 Jul. 1951 and 31 Jul. 1951, between 1 Jan. 1951 and
31 Dec. 1954, between 1 Jan. 1951 and 31 Dec. 1951, leg. Alexandre Fain (RMCA: 81.012-M-0094,
81.012-M-0101, 81.012-M-0093, 81.012-M-0091, 81.012-M-0098, 81.012-M-0099, 81.012-M-0095,
81.012-M-0097, 23278, 23284, 23288, 81.012-M-0102, 23289, 23279, 23281); 1 ad ♀ [SS], Wasa
Wago Forest, unknown date, leg. Alexandre Fain (BMNH: 1951.629 [paratype Tadarida (Chaerephon)
faini Hayman, 1951]); 1 ♂ [UN], Zeya, 4 Aug. 1953, leg. unknown collector (RBINS: 14558).
RWANDA: 4 ♀♀, 3 ♂♂, 22 ?? [13 SS, 13 SA, 1 AL, 2 SO], Astrida [= Butare], unknown date,
1 Jan. 1954, 29 Jul. 1954, 25 Oct. 1954, 26 Oct. 1954, 27 Oct. 1954, 28 Oct. 1954, between 1 Jan. 1954
and 31 Dec. 1954, leg. Alexandre Fain, 20 Feb. 1962, between 1 Mar. 1962 and 31 Mar. 1962, leg. Van
Parijs, prior to 9 Jul. 1952, leg. P.G. Vercammen (MHNG: 921.041, RMCA: 81.012-M-0070, 81.012M-0066, 31047, MHNG: 921.039, RMCA: 81.012-M-0063, 81.012-M-0065, 81.012-M-0077, MHNG:
921.038, RMCA: 81.012-M-0064, 81.012-M-0067 to 81.012-M-0069, MHNG: 921.040, RMCA:
81.012-M-0105, 81.012-M-0106, 31061, 31063 to 31068, 31071 to 31075, 21218); 1 ? [UN], Bilalu,
unknown date, leg. unknown collector (INPBC: V3062); 53 ad ♀♀, 33 ad ♂♂, 10 sad ♀♀, 16 sad ♂♂,
1 juv ♀, 1 juv ♂ [25 SA, 89 AL], Butare, 10 Dec. 1984, 11 Dec. 1984, 13 Dec. 1984, 20 Dec. 1984,
22 Dec. 1984, 24 Dec. 1984, 25 Dec. 1984, 31 Dec. 1984, 1 Jan. 1985, 2 Jan. 1985, 14 Jan. 1985,
15 Jan. 1985, leg. Benny Baeten and Luc Janssens, 1 Mar. 1985, leg. Théoneste (RMCA: 85.006-M0259 to 85.006-M-0316, 87.047-M-0098 to 87.047-M-0103, 85.006-M-0207 to 85.006-M-0211, 85.006M-0384 to 85.006-M-0388, 85.006-M-0439 to 85.006-M-0446, 85.006-M-0317, 85.006-M-0212 to
85.006-M-0220, 85.006-M-0237 to 85.006-M-0258); 1 ♀, 1 ♂ [2 SA], Cyangugu [= Shangugu],
16 Jun. 1969, leg. unknown collector (SMF: 52479, 52480); 28 ad ♀♀, 15 ad ♂♂, 3 sad ♀♀, 5 sad
♂♂, 1 juv ♀, 1 juv ♂ [12 SA, 41 AL], Cyili, 8 Apr. 1985, 13 Apr. 1985, 21 Apr. 1985, 2 May 1985, leg.
Théoneste, 24 Dec. 1984, 2 Jan. 1985, leg. Benny Baeten and Luc Janssens (RMCA: 87.047-M-0112
to 87.047-M-0116, 85.006-M-0318 to 85.006-M-0329, 87.047-M-0120 to 87.047-M-0128, 87.047-M0117 to 87.047-M-0119, 85.006-M-0221 to 85.006-M-0236, 87.047-M-0104 to 87.047-M-0111); 2 ??
[2 UN], Gabiro, unknown date, leg. unknown collector (INPBC: V3030, V3051); 10 ad ♀♀, 8 ad
♂♂, 1 sad ♀, 3 sad ♂♂ [2 SA, 20 AL], Gafumba, 12 Jan. 1985, leg. Benny Baeten and Luc Janssens,
7 Feb. 1985, leg. Théoneste (RMCA: 85.006-M-0427 to 85.006-M-0438, 87.047-M-0088 to 87.047-M0097); 3 ?? [3 AL], Ihema, Lake, between 1 Jun. 1969 and 30 Jun. 1969, leg. Kiss (RMCA: 74.023M-0001 to 74.023-M-0003); 42 ad ♀♀, 23 ad ♂♂ [65 AL], Karama, 12 Oct. 1981, 13 Oct. 1981, leg.
Frits De Vree, Benny Baeten and Victor Van Cakenberghe (RMCA: 82.006-M-0464 to 82.006-M-0528);
4 ♀♀, 1 ♂ [5 AL], Kibungo (former) Prefecture [= Eastern Prefecture], 26 Nov. 1985, leg. unknown
collector (SMF: 66858 to 66862); 143 ad ♀♀, 48 ad ♂♂, 1 sad ♂, 2 ♀♀ [8 SA, 186 AL], Kidaho,
6 Jun. 1982, 7 Jun. 1982, 8 Jun. 1982, 9 Jun. 1982, 10 Jun. 1982, 11 Jun. 1982, 12 Jun. 1982, 15 Jun. 1982,
16 Jun. 1982, 17 Jun. 1982, 18 Jun. 1982, 19 Jun. 1982, leg. Benny Baeten (RMCA: 87.046-M-0326
to 87.046-M-0328, 87.046-M-0330 to 87.046-M-0337, 87.046-M-0470 to 87.046-M-0491, 87.046M-0493, 87.046-M-0494, 87.046-M-0338 to 87.046-M-0342, 87.046-M-0495 to 87.046-M-0501,
87.046-M-0343, 87.046-M-0502 to 87.046-M-0506, 87.046-M-0329, 87.046-M-0492, 87.046-M-0507
to 87.046-M-0510, 87.046-M-0281 to 87.046-M-0285, 87.046-M-0380 to 87.046-M-0386, 87.046-M0286 to 87.046-M-0294, 87.046-M-0309 to 87.046-M-0313, 87.046-M-0387 to 87.046-M-0418, 87.046M-0295 to 87.046-M-0304, 87.046-M-0314 to 87.046-M-0317, 87.046-M-0419 to 87.046-M-0433,
87.046-M-0305 to 87.046-M-0308, 87.046-M-0318 to 87.046-M-0325, 87.046-M-0434 to 87.046-M0469); 2 ♂♂, 1 ? [2 SS, 1 SA], Kiziguru (Mission) [= Kiziguro (Mission)], 2 Jul. 1954, 31 Jul. 1954,
2 Aug. 1954, leg. Alexandre Fain (RMCA: 81.012-M-0061, MHNG: 921.042, RMCA: 81.012-M-0060);
3 ?? [3 SA], Mabanza, 9 Oct. 1980, leg. Danny Meirte (RMCA: 80.055-M-0162 to 80.055-M-0164);
3 ?? [3 UN], Mihindi, Lake, unknown date, leg. unknown collector (INPBC: W3111.1 to W3111.3);
11 ad ♀♀, 1 ad ♂, 5 sad ♀♀, 1 sad ♂ [1 SA, 17 AL], Mugogwe, 5 Feb. 1985, leg. Théoneste (RMCA:
206
VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
87.047-M-0069 to 87.047-M-0086); 184 ad ♀♀, 81 ad ♂♂, 1 ♀ [266 AL], Nyamata, 11 Nov. 1981,
12 Nov. 1981, 13 Nov. 1981, 14 Nov. 1981, 15 Nov. 1981, 16 Nov. 1981, leg. Frits De Vree, Benny
Baeten and Victor Van Cakenberghe (RMCA: 82.006-M-0529 to 82.006-M-0794); 15 ad ♀♀, 5 ad ♂♂,
4 sad ♀♀, 15 sad ♂♂ [5 SA, 34 AL], Rubona, 7 Jan. 1985, leg. Benny Baeten and Luc Janssens, between
1 Jan. 1985 and 31 Dec. 1985, leg. Théoneste (RMCA: 85.006-M-0389 to 85.006-M-0426, 87.047-M0087); 65 ad ♀♀, 33 ad ♂♂, 1 sad ♀, 1 juv ♀, 1 ♀ [101 AL], Rukira, 19 Nov. 1981, 20 Nov. 1981,
21 Nov. 1981, 22 Nov. 1981, 23 Nov. 1981, 24 Nov. 1981, 25 Nov. 1981, 26 Nov. 1981, 27 Nov. 1981,
leg. Frits De Vree, Benny Baeten and Victor Van Cakenberghe (RMCA: 82.006-M-0795 to 82.006-M0813, 82.006-M-0848 to 82.006-M-0880, 82.006-M-0814 to 82.006-M-0816, 82.006-M-0881, 82.006M-0817 to 82.006-M-0847, 82.006-M-0882 to 82.006-M-0895); 25 ad ♀♀, 9 ad ♂♂, 13 sad ♀♀, 8 sad
♂♂ [15 SA, 39 AL, 1 SK], Shangi, 5 Jan. 1985, 6 Jan. 1985, leg. Benny Baeten and Luc Janssens
(RMCA: 85.006-M-0330 to 85.006-M-0360, 85.006-M-0540, 85.006-M-0361 to 85.006-M-0383).
Chaerephon russatus J.A. Allen, 1917
DEMOCRATIC REPUBLIC OF THE CONGO: 1 ad ♀, 2 ad ♂♂, 1 foetus ?, 21 ♀♀, 4 ♂♂ [11 SS,
1 SK, 17 UN], Medje, 8 Sep. 1910, leg. Herbert Lang, James Paul Chapin and The American Museum
Congo Expedition, 8 Sep. 1910, leg. Herbert Lang, 8 Sep. 1910, leg. unknown collector (AMNH: 48911
to 48923, 48925 [holotype Chærephon russatus J.A. Allen, 1917], 48926 to 48929, 48931, 48935,
49219, 49265, 49266, 49278, BMNH: 1970.2507, FMNH: 43867, MCZ: 17386 [topotype Chærephon
russatus J.A. Allen, 1917], 17387 [topotype Chærephon russatus J.A. Allen, 1917], RMCA: 12389);
1 ♀ [SS], Tandala, 14 Jul. 1979, leg. C. Brian Robbins (USNM: 537762).
Mops Lesson, 1842
Mops (Mops) Lesson, 1842
Mops (Mops) condylurus (A. Smith, 1833)
BURUNDI: 1 ♀, 32 ♂♂, 145 ?? [1 SA, 62 AL, 115 UN], Bujumbura, unknown date, prior to 21 Feb. 1962,
leg. Kiss, 21 Apr. 1982, 26 Apr. 1982, 27 Apr. 1982, leg. Danny Meirte, prior to 31 Dec. 1964, leg.
unknown collector (RMCA: 31038, 31039, 31040-01 to 31040-46, 31041-01 to 31041-69, 82.016-M0001, 82.016-M-0094 to 82.016-M-0123, 82.016-M-0129, 31010 to 31037, SMF: 23789); 2 ♀♀ [1 SA,
1 SK], Bujumbura, 7 km NW, 21 Mar. 1979, leg. Heinz Stephan (SMF: 57552, 57553); 4 ♂♂ [2 SA,
2 SK], Lake Cohoha [= Cohoha, Lake], 9 Mar. 1979, 10 Mar. 1979, leg. Heinz Stephan (SMF: 57550,
57551, 57548, 57549).
DEMOCRATIC REPUBLIC OF THE CONGO: 21 ?? [1 AL, 20 UN], unknown locality, unknown
date, leg. Antoon Emeric Marcel De Roo, between 1 Apr. 1999 and 30 Apr. 1999, leg. unknown collector
(RMCA: 32781, SMF: 93323 to 93342); 3 ?? [3 UN], Aba, 12 Apr. 1948, between 12 Apr. 1948 and
31 Dec. 1948, leg. unknown collector (RBINS: 8154.1 to 8154.3); 1 ad ♀, 1 emb ?, 4 ♀♀, 3 ♂♂,
25 ?? [5 SS, 10 AL, 18 SK, 1 SO], Cel II, 1 Jan. 1950, 10 May 1951, 30 Apr. 1952, leg. H. de Saeger
et al., 2 Apr. 1951, 10 May 1951, 7 Mar. 1952, 30 Apr. 1952, 10 May 1952, leg. H. de Saeger (RBINS:
14004, 14000.01 to 14000.11, 14001.1, 14001.2, RMCA: 36908 to 36910, 36899, 36900, 36903 to
36907, RBINS: 14003.1 to 14003.4, 14005, 14006, RMCA: 36902, 36913, 36914, 36901); 1 ? [SS],
Defrenne Cave [= Mwanga Cave], 2 Apr. 1957, leg. G. Verhoustraete (RMCA: 26213); 1 ♂ [SA],
Duma, 12 Oct. 1910, leg. unknown collector (SMF: 6397); 2 ♀♀, 5 ♂♂ [6 AL, 1 UN], Elisabethville
[= Lubumbashi], 1 Mar. 1967, leg. unknown collector, 6 Feb. 1956, 15 Mar. 1956, 10 Jul. 1956, leg.
Félix / Michel Anciaux de Faveaux (RBINS: 18510, RMCA: 23916, 23917, 23914, 23915, 23906,
23907); 2 ?? [2 SK], Gangala-na-Bodio, 9 Apr. 1948, 10 Apr. 1948, leg. unknown collector (RBINS:
8156, 8155); 1 ♂ [UN], Goma, 24 Dec. 1952, leg. unknown collector (RBINS: 15461); 4 ad ♀♀, 4 ad
♂♂, 1 sad ♀, 2 sad ♂♂, 1 imm ♀, 3 imm ♂♂, 4 ♀♀, 5 ♂♂, 60 ?? [40 AL, 44 UN], Ishango River,
207
European Journal of Taxonomy 382: 1–327 (2017)
unknown date, leg. unknown collector, between 1 Jan. 1982 and 31 Dec. 1982, leg. Marc Colyn, between
26 Dec. 1968 and 29 Dec. 1968, leg. M. Lejeune (INPBC: V112, V1127, V1176.001 to V1176.042,
RMCA: 82.031-M-0001, 35421 to 35459); 10 ?? [2 AL, 8 SK], Iso, 4 Mar. 1952, leg. H. de Saeger
et al., 4 Mar. 1952, leg. H. de Saeger (RBINS: 14002.1 to 14002.8, RMCA: 36911, 36912); 12 ♀♀,
2 ♂♂ [13 AL, 1 UN], Jadotville [= Likasi], 10 Dec. 1957, 26 Mar. 1959, leg. Félix / Michel Anciaux
de Faveaux, 4 Apr. 1960, leg. unknown collector (RMCA: 27541 to 27547, 27899 to 27904, RBINS:
14559); 27 ?? [8 AL, 19 UN], Kambukabakali, unknown date, leg. unknown collector, 23 Oct. 1954,
27 Oct. 1954, 29 Oct. 1954, 5 Nov. 1954, leg. Gaston-François de Witte (INPBC: W1284, W1286,
W1288, W1290, W1292, W1294, W1296, W1298, W1302.01 to W1302.11, RMCA: 37384, 37385,
37398 to 37400, 37386, 37397, 37401); 2 ?? [2 SA], Kapolowe, between 1 Jul. 1954 and 31 Jul. 1954,
leg. De Smet (RMCA: 22438, 22439); 1 ♀, 2 ♂♂ [1 SS, 1 AL, 1 SO], Kasapa, 2 Jan. 1969, 14 Jan. 1969,
leg. unknown collector, 29 Apr. 1956, leg. Félix / Michel Anciaux de Faveaux (RBINS: 18845, 18844,
RMCA: 23920); 1 ♂ [UN], Kasenye, 10 km N, 23 Nov. 1954, leg. unknown collector (RBINS:
15463); 2 ♂♂, 1 ? [1 AL, 2 UN], Kasenyi [= Kasenye], 19 Feb. 1955, leg. Gaston-François de Witte,
21 Sep. 1953, 19 Nov. 1953, leg. unknown collector (RMCA: 37402, RBINS: 14560, 14561); 1 ♀,
3 ♂♂, 9 ?? [13 AL], Kashiobwe, 20 Oct. 1955, leg. Van Iseghem (RMCA: 26307 to 26319); 3 ??
[3 AL], Kasongo, 13 Nov. 1959, leg. Segers and Benoit Mission (RMCA: 29165, 29167, 29168); 1 ♀,
1 ♂ [2 AL], Kenya, 26 Feb. 1956, leg. Félix / Michel Anciaux de Faveaux (RMCA: 23825, 23918); 2 juv
♀♀, 1 juv ♂, 10 ♀♀, 8 ♂♂, 1 ? [22 AL], Keyberg, 6 Feb. 1956, 17 Feb. 1956, 17 Mar. 1956, leg. Félix /
Michel Anciaux de Faveaux, 19 Dec. 1955, leg. M. Lips, 6 Feb. 1956, leg. service de l’hygiene publique
(RMCA: 23909 to 23913, 23908, 26297 to 26302, 26304, 26305, 23822, 23901 to 23905, 26296,
26320); 7 ad ♀♀, 3 ad ♂♂ [5 SA, 5 UN], Kikwit, 15 Jul. 1995, leg. Herwig Leirs et al. (MSB: 274584 to
274588, RMCA: 97.021-M-0811 to 97.021-M-0815); 1 ad ♀, 1 ad ♂ [2 SA], Kinshasa, 22 Dec. 1914,
leg. Herbert Lang, James Paul Chapin and The American Museum Congo Expedition (AMNH: 49230
[holotype Mops (Allomops) osborni J.A. Allen, 1917], 49244 [paratype Mops (Allomops) osborni
J.A. Allen, 1917]); 1 ♂ [AL], Kipushi, road to, 6 Feb. 1956, leg. M. Lips (RMCA: 26306); 54 ??
[54 UN], Kongolo, between 8 Nov. 1974 and 9 Nov. 1974, leg. Michael D. Gallagher (HZM: 156.7822
to 209.7875); 4 ?? [4 SS], Léopoldville [= Kinshasa], 4 Apr. 1964, leg. Antoon Emeric Marcel De Roo
(RMCA: 32390 to 32392, 32659); 1 ♂, 1 ? [2 AL], Lovanium University [= Kinshasa], 23 Apr. 1963,
8 Jul. 1964, leg. Jo Van Orshoven (ZMA: MAM.11165, MAM.11166); 1 ? [AL], Lubondai-Tshimbulu
[= Lubondaie-Tshimbulu], between 1 Jan. 1953 and 31 Dec. 1953, leg. S. Marks (RMCA: 26303);
1 ♀ [UN], Luluabourg [= Kananga], unknown date, leg. Richard Callewaert (BMNH: 1926.7.6.107);
23 ♀♀, 24 ♂♂ [1 SS, 27 SK, 19 UN], Luvungi, 15 Dec. 1932, leg. Burgeon, 6 May 2015, 7 May
2015, 17 May 2015, leg. Prescott Musaba, Moïse Bipoo, Charle Andabhati, Benjamin Kirongozi and
Kambale Karupao (RMCA: 12045, UNIKIS: BUKCH17, BUKCH01, BUKCH03 to BUKCH06,
BUKCH08 to BUKCH15, BUKCH18 to BUKCH20, BUKCH22, BUKCH24 to BUKCH27, BUKCH07,
BUKCH28, BUKCH30 to BUKCH32, BUKCH34 to BUKCH38, BUKCH40, BUKCH42, BUKCH44
to BUKCH46, BUKCH48, BUKCH50 to BUKCH56, BUKCH58); 1 ? [SS], Nionga, 3 May 1925,
leg. Gaston-François de Witte (RMCA: 9230); 48 ?? [48 UN], Rwindi, unknown date, leg. unknown
collector (INPBC: V1051, V1542, V1543, V1546.01 to V1546.45); 10 ?? [1 SA, 9 AL], Sakania, prior
to 1 Sep. 1954, leg. De Smet (RMCA: 22442 to 22451); 16 ♀♀, 12 ♂♂ [10 SK, 18 UN], Sange, 13 May
2015, 14 May 2015, 15 May 2015, leg. Prescott Musaba, Moïse Bipoo, Charle Andabhati, Benjamin
Kirongozi and Kambale Karupao (UNIKIS: BUKCH60, BUKCH61, BUKCH65, BUKCH68, BUKCH70
to BUKCH84, BUKCH86 to BUKCH88, BUKCH91 to BUKCH94, BUKCH96, BUKCH99); 3 ♀♀,
17 ♂♂, 1 ? [6 SS, 1 SA, 13 AL, 1 UN], Tandala, 12 Jul. 1979, 13 Jul. 1979, 16 Jul. 1979, leg. unknown
collector, 12 Jul. 1979, 13 Jul. 1979, 16 Jul. 1979, 21 Jul. 1979, leg. C. Brian Robbins, 13 Jul. 1979,
leg. Lynn W. Robbins, 16 Jul. 1979, leg. Frieder Meier (MHNG: 1870.051, 1870.052, USNM: 463507
to 463509, 537738, 537739, 537741, MHNG: 1870.046, USNM: 463510 to 463512, 537740, MHNG:
1743.060, 1870.047 to 1870.050, USNM: 463513 to 463515); 3 ?? [3 AL], Tshibambo [= Shibambo],
31 Aug. 1954, leg. De Smet (RMCA: 22459 to 22461); 7 ♀♀, 10 ♂♂ [2 SS, 15 AL], Uvira, 2 May 1959,
208
VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
leg. Uwe Rahm, 27 Jul. 1956, leg. Alvin Novick (RMCA: 28905, 28906, MCZ: 48200 to 48214); 1 ♂
[UN], Vitshumbi, 12 Jan. 1953, leg. unknown collector (RBINS: 14562).
RWANDA: 2 ♂♂ [2 SA], unknown locality, 12 Jul. 1964, leg. unknown collector (SMF: 34954,
35792); 2 ♀♀, 1 ♂, 1 ? [2 SS, 1 SA, 1 UN], Astrida [= Butare], unknown date, 25 Oct. 1954, leg.
Alexandre Fain, 26 Feb. 1966, leg. unknown collector (BMNH: 1953.681, MHNG: 921.043, RMCA:
81.012-M-0048, RBINS: 15496); 2 ?? [2 SA], Bugesera, between 1 Sep. 1957 and 30 Sep. 1957,
leg. Narcisse Leleup (RMCA: 26574, 26575); 2 ad ♀♀, 3 sad ♂♂ [5 AL], Butare, 20 Dec. 1984,
21 Dec. 1984, 22 Dec. 1984, 15 Jan. 1985, leg. Benny Baeten and Luc Janssens (RMCA: 85.006M-0462, 85.006-M-0458 to 85.006-M-0461); 121 ?? [52 AL, 9 SK, 60 UN], Gabiro, unknown date,
23 Sep. 1948, 27 Sep. 1948, leg. unknown collector, 9 Feb. 1960, leg. I.P.N.C. (INPBC: V3128.01 to
V3128.60, RBINS: 8157, 8158.1 to 8158.8, RMCA: 78.038-M-0002 to 78.038-M-0053); 14 ad ♀♀,
24 ad ♂♂, 3 sad ♀♀, 6 sad ♂♂ [47 AL], Gafumba, 7 Feb. 1985, leg. Théoneste (RMCA: 87.047M-0022 to 87.047-M-0068); 3 ♀♀, 2 ♂♂, 1 ? [3 SK, 3 UN], Gahinga (Vulcano), 18 Feb. 1966, leg.
unknown collector (RBINS: 15498.1 to 15498.3, 15935.1 to 15935.3); 2 ♀♀, 2 ♂♂ [4 SK], Gihinda
[= Gihinga], 3 Mar. 1966, leg. unknown collector (RBINS: 15497.1 to 15497.4); 15 ad ♀♀, 20 ad ♂♂,
1 sad ♀, 1 juv ♀, 1 juv ♂ [34 SA, 4 AL], Kidaho, 9 Jun. 1982, 10 Jun. 1982, 11 Jun. 1982, 12 Jun. 1982,
14 Jun. 1982, leg. Benny Baeten (RMCA: 87.046-M-0273 to 87.046-M-0280, 87.046-M-0511 to 87.046M-0513, 87.046-M-0247 to 87.046-M-0272, 87.046-M-0379); 12 ♀♀, 5 ♂♂, 18 ?? [4 SS, 12 SA, 6 SK,
6 SO, 7 UN], Kigali, 19 Feb. 1967, leg. unknown collector, 30 Sep. 1980, leg. Danny Meirte (RBINS:
15929.1 to 15929.7, 16007 to 16022, RMCA: 80.055-M-0113 to 80.055-M-0124); 37 ?? [35 SA, 2 AL],
Mabanza, 9 Oct. 1980, leg. Danny Meirte (RMCA: 80.055-M-0125 to 80.055-M-0161); 1 ♂ [UN],
Masha, 1 Jan. 1956, leg. unknown collector (RBINS: 22465); 7 ad ♀♀, 12 ad ♂♂ [1 SS, 17 SA, 1 AL],
Nyamata, 13 Nov. 1981, 14 Nov. 1981, 15 Nov. 1981, leg. Frits De Vree, Benny Baeten and Victor
Van Cakenberghe (RMCA: 82.006-M-0339 to 82.006-M-0357); 1 ♂ [UN], Rubona, 27 Jan. 1966, leg.
Félix / Michel Anciaux de Faveaux (RBINS: 15495); 28 ad ♀♀, 53 ad ♂♂ [77 SA, 4 AL], Rukira,
19 Nov. 1981, 20 Nov. 1981, 21 Nov. 1981, 23 Nov. 1981, 25 Nov. 1981, 26 Nov. 1981, 27 Nov. 1981,
leg. Frits De Vree, Benny Baeten and Victor Van Cakenberghe (RMCA: 82.006-M-0358 to 82.006-M0438); 1 ♂ [SA], Rwamagana, between 1 Jan. 1954 and 31 Dec. 1955, leg. Alexandre Fain (MHNG:
921.044).
Mops (Mops) congicus J.A. Allen, 1917
DEMOCRATIC REPUBLIC OF THE CONGO: 2 ad ♀♀, 1 foetus ?, 14 ♀♀, 1 ? [9 SS, 2 AL, 7 UN],
Medje, 8 Sep. 1910, leg. Herbert Lang, James Paul Chapin and The American Museum Congo
Expedition, 8 Sep. 1910, leg. Herbert Lang, 8 Sep. 1910, leg. unknown collector (AMNH: 48888 to
48892, 48893 [holotype Mops congicus J.A. Allen, 1917], 48894 to 48897, 48900, 49213, 49225,
49277, 49285, FMNH: 43864, MCZ: 17385, RMCA: 12380); 4 ♀♀, 5 ♂♂ [6 SS, 1 SA, 2 AL], Tandala,
12 Jul. 1979, leg. O.W.H., 12 Jul. 1979, 16 Jul. 1979, 21 Jul. 1979, leg. C. Brian Robbins, 21 Jul. 1979,
leg. unknown collector (TTU: 36358, USNM: 537742, 537743, 537745, MHNG: 1870.043, 1870.044,
USNM: 463516, 463517, 537746); 1 ? [SO], Tshopo, between 1 Sep. 1985 and 30 Sep. 1985, leg. Marc
Colyn (RMCA: 86.024-M-0006); 1 ♂ [SS], Yalosemba, 28 Jun. 1979, leg. C. Brian Robbins (USNM:
537744).
Mops (Mops) demonstrator (Thomas, 1903)
DEMOCRATIC REPUBLIC OF THE CONGO: 4 ♀♀ [4 UN], Bagunda, 19 Jan. 1951, leg. H. de
Saeger et al. (RBINS: 13806.1 to 13806.4); 3 ♀♀, 1 ♂, 16 ?? [5 SS, 15 UN], Cel II, unknown date, leg.
unknown collector, 2 Apr. 1951, 12 May 1951, 5 Oct. 1951, 26 Nov. 1951, 7 Apr. 1952, 28 May 1952,
3 Sep. 1952, leg. H. de Saeger et al., 30 Apr. 1952, 3 Sep. 1952, leg. H. de Saeger (INPBC: 3514, 4735,
RBINS: 15805, 13801, 13800, 13804.1, 13804.2, 13809, 13811.1 to 13811.5, 13813, RMCA: 36933 to
209
European Journal of Taxonomy 382: 1–327 (2017)
36935, 36932, RBINS: 13805, 13808.1); 1 ad ♂, 1 ♀ [1 SS, 1 SA], Faradje, 8 Aug. 1911, 12 Jan. 1913,
leg. Herbert Lang, James Paul Chapin and The American Museum Congo Expedition (AMNH: 49222
[holotype Mops (Allomops) faradjius J.A. Allen, 1917], 48910); 1 ♂ [UN], Garamba National Park,
between 1 Jan. 1950 and 31 Dec. 1952, leg. H. de Saeger et al. (RBINS: 13812); 2 juv ♀♀, 1 juv ?,
1 foetus ?, 1 ♀ [1 SA, 4 AL], Keyberg, 19 Dec. 1955, 4 Feb. 1956, 9 Feb. 1956, leg. Félix / Michel
Anciaux de Faveaux (RMCA: 26218, 26219, 26222, 26223, 26228); 2 ♀♀, 2 ?? [4 SS], Parc National
de Garamba [= Garamba National Park], 30 Jun. 1951, 7 Apr. 1952, 28 May 1952, leg. H. de Saeger
et al. (RBINS: 13824, 13825, 13807, 13810); 1 ? [SO], PFNK, 7 Feb. 1952, leg. H. de Saeger (RMCA:
36931).
RWANDA: 1 ♂ [SS], Musha, 14 Nov. 1955, leg. Alexandre Fain (RMCA: 37983).
Mops (Mops) midas (Sundevall, 1843)
DEMOCRATIC REPUBLIC OF THE CONGO: 2 ♀♀, 1 ♂ [3 UN], Bagunda, 19 Jan. 1951, leg. H. de
Saeger et al. (RBINS: 13815.1 to 13815.3); 2 ?? [2 SS], Bwamanda [Mission], between 1 Jul. 1961 and
31 Jul. 1961, leg. Vedast (Gaston Alfons) Maes (RMCA: 30949, 30950); 1 ad ♀, 1 emb ?, 4 ♀♀, 1 ♂,
12 ?? [6 SS, 1 SO, 12 UN], Cel II, unknown date, leg. unknown collector, 8 Mar. 1951, 21 Feb. 1952,
8 Apr. 1952, 30 Apr. 1952, 3 Oct. 1952, leg. H. de Saeger et al., 8 Apr. 1952, 3 Oct. 1952, 22 Oct. 1952, leg.
H. de Saeger (INPBC: 5159.4.1 to 5159.4.3, RBINS: 13819, RMCA: 36929, 36930, RBINS: 13822.1,
13822.2, RMCA: 36927, 36928, RBINS: 13821.1, 13821.2, 13814.1, 13814.2, 13820.1, 13820.2,
RMCA: 36924 to 36926); 1 ad ♀, 1 ♀, 4 ?? [6 SA], Faradje, 7 Mar. 1912, 9 Mar. 1912, leg. Herbert
Lang, James Paul Chapin and The American Museum Congo Expedition (AMNH: 49254 to 49257,
49262, 49272); 3 ?? [3 UN], Kamohohora, unknown date, leg. unknown collector (INPBC: V183.1 to
V183.3); 3 ?? [3 UN], Mwiga, unknown date, leg. unknown collector (INPBC: V500.1 to V500.3); 1 ♂,
17 ?? [7 SS, 1 AL, 9 UN, 1 SN], Ndelele, 8 Feb. 1952, leg. H. de Saeger et al., 8 Feb. 1952, leg. H. de
Saeger (RBINS: 13816.1 to 13816.4, 13817.1, 13817.2, 13818.1 to 13818.3, RMCA: 36915 to 36923);
1 ♂ [UN], Parc National de Garamba [= Garamba National Park], 3 Oct. 1952, leg. H. de Saeger
et al. (RBINS: 13823); 6 ♀♀, 4 ?? [2 SS, 8 AL], Uvira, 2 Mar. 1959, leg. Uwe Rahm, 23 Sep. 1954,
between 1 Mar. 1955 and 31 Mar. 1955, leg. Narcisse Leleup, 25 May 1955, leg. Waldo LaSalle Schmitt
and E. Baker, 27 Jul. 1956, leg. Alvin Novick, 5 Sep. 1956, leg. Laurent, prior to 31 Oct. 1953, leg.
Georges Marlier (RMCA: 33530, 27214, USNM: 301727, 301728, MCZ: 48197 to 48199, RMCA:
35694, 22911, 38152).
Mops (Mops) niangarae J.A. Allen, 1917
DEMOCRATIC REPUBLIC OF THE CONGO: 1 ad ♂ [SS], Niangara, 12 Dec. 1910, leg. Herbert
Lang, James Paul Chapin and The American Museum Congo Expedition (AMNH: 48901 [holotype
Mops niangaræ J.A. Allen, 1917]).
Mops (Mops) niveiventer Cabrera & Ruxton, 1926
BURUNDI: 4 ad ♀♀ [2 SA, 2 SK], Kayanza, 12 Mar. 1979, leg. Heinz Stephan (SMF: 57555 to
57558); 1 ♂ [SK], Rutana, 4 Mar. 1979, leg. Heinz Stephan (SMF: 57554).
DEMOCRATIC REPUBLIC OF THE CONGO: 1 sad ♂, 1 ♀, 1 ♂ [2 SA, 1 AL], Elisabethville
[= Lubumbashi], 19 Dec. 1955, 14 Nov. 1956, leg. unknown collector, 2 Nov. 1956, leg. Félix /
Michel Anciaux de Faveaux (SMF: 16867, 16876, RMCA: 26245); 1 ♂ [SA], Jadotville [= Likasi],
23 Sep. 1957, leg. Félix / Michel Anciaux de Faveaux (MHNG: 1046.029); 2 ♀♀ [1 SA, 1 AL],
Jadoville [= Likasi], 10 Dec. 1957, leg. Félix / Michel Anciaux de Faveaux (SMF: 18942, 18943);
1 ? [AL], Kasongo Samba, 22 Sep. 1959, leg. Segers and Benoit Mission (RMCA: 29206); 1 juv ♂,
6 ♀♀, 2 ?? [1 SA, 8 AL], Keyberg, 4 Feb. 1956, 6 Feb. 1956, 9 Feb. 1956, 17 Feb. 1956, leg. Félix /
210
VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
Michel Anciaux de Faveaux (MHNG: 923.003, 923.004, 923.007, 923.002, 923.005, 923.001, 923.006,
RMCA: 23823, 26229); 1 ♀ [SA], Kikole-Pecheur, 17 May 1980, leg. M. Mutashia (SMF: 58073);
3 ad ♂♂ [1 SA, 2 UN], Kimbinga Market, 20 Jul. 1995, leg. Herwig Leirs et al. (MSB: 274601,
274602, RMCA: 97.021-M-0823); 1 ♀, 1 ♂ [2 SS], Luluabourg [= Kananga], unknown date, leg.
Richard Callewaert (BMNH: 1926.7.6.104, 1926.7.6.106); 5 ♀♀, 1 ♂ [6 SS], Matamba, 20 Dec. 1983,
leg. Betu Nsankulu (RMCA: 84.035-M-0014, 84.035-M-0016 to 84.035-M-0018, 84.035-M-0020,
84.035-M-0021); 1 ♀ [SS], Mikalaya, 22 Dec. 1983, leg. Betu Nsankulu (RMCA: 84.035-M-0022);
1 ♀, 1 ♂ [2 SS], Mikalaya Mission [= Mikalaya], 22 Dec. 1983, leg. Betu Nsankulu (RMCA: 84.035M-0015, 84.035-M-0019); 1 ♀ [UN], St. Joseph de Luluabourg Mission [= Mikalaya], 16 Apr. 1924,
leg. Richard Callewaert (BMNH: 1926.7.6.109 [holotype Mops angolensis niveiventer Cabrera and
Ruxton, 1926]); 1 juv ♀, 1 foetus ? [2 AL], Wasseige Farm, 6 Feb. 1956, leg. Félix / Michel Anciaux
de Faveaux (RMCA: 26225, 26226).
RWANDA: 1 ad ♀, 1 ♂, 8 ?? [3 SS, 7 SA], Astrida [= Butare], 13 May 1952, leg. P.G. Vercammen,
20 Feb. 1962, between 1 Mar. 1962 and 31 Mar. 1962, leg. Van Parijs, 26 Dec. 1950, leg. Aureliaan,
between 1 Jan. 1948 and 31 Dec. 1948, leg. Alexandre Fain (RMCA: 23171, 31045, 31046, 20489,
18643, 18991, 31060, 31062, 31069, 31070); 3 ad ♀♀ [3 AL], Butare, 1 Mar. 1985, leg. Théoneste,
15 Jan. 1985, leg. Benny Baeten and Luc Janssens (RMCA: 87.047-M-0009, 87.047-M-0010, 85.006-M0206); 15 ad ♀♀, 8 ad ♂♂ [22 SA, 1 AL], Butare INRS [= Butare], 8 Oct. 1981, 9 Oct. 1981, 8 Nov. 1981,
9 Nov. 1981, leg. Frits De Vree, Benny Baeten and Victor Van Cakenberghe (RMCA: 82.006-M-0439 to
82.006-M-0454, 82.006-M-0460, 82.006-M-0455 to 82.006-M-0459, 82.006-M-0461); 18 ad ♀♀, 9 ad
♂♂, 14 sad ♀♀, 7 sad ♂♂ [48 AL], Cyili, 8 Apr. 1985, 13 Apr. 1985, 21 Apr. 1985, 2 May 1985, leg.
Théoneste, 24 Dec. 1984, 2 Jan. 1985, leg. Benny Baeten and Luc Janssens (RMCA: 87.047-M-0013
to 87.047-M-0016, 85.006-M-0192 to 85.006-M-0195, 87.047-M-0020, 87.047-M-0021, 87.047-M0017 to 87.047-M-0019, 85.006-M-0159 to 85.006-M-0191, 87.047-M-0011, 87.047-M-0012); 1 ad ♀,
2 ad ♂♂, 1 sad ♀ [4 AL], Gafumba, 12 Jan. 1985, leg. Benny Baeten and Luc Janssens, 7 Feb. 1985,
leg. Théoneste (RMCA: 85.006-M-0205, 87.047-M-0006 to 87.047-M-0008); 1 ad ♀, 3 ad ♂♂ [1 SA,
3 AL], Mugogwe, 5 Feb. 1985, leg. Théoneste (RMCA: 87.047-M-0002 to 87.047-M-0005); 1 ad ♂
[SA], Nyamata, 13 Nov. 1981, leg. Frits De Vree, Benny Baeten and Victor Van Cakenberghe (RMCA:
82.006-M-0462); 5 ad ♀♀, 4 ad ♂♂ [9 AL], Rubona, 2 Jan. 1985, 7 Jan. 1985, leg. Benny Baeten and
Luc Janssens (RMCA: 85.006-M-0196 to 85.006-M-0204); 1 ad ♀ [SA], Rukira, 21 Nov. 1981, leg.
Frits De Vree, Benny Baeten and Victor Van Cakenberghe (RMCA: 82.006-M-0463).
Mops (Mops) trevori J.A. Allen, 1917
DEMOCRATIC REPUBLIC OF THE CONGO: 1 ad ♀ [SA], Faradje, 29 Sep. 1912, leg. Herbert
Lang, James Paul Chapin and The American Museum Congo Expedition (AMNH: 49250 [holotype
Mops trevori J.A. Allen, 1917]).
Mops (Xiphonycteris) Dollman, 1911
Mops (Xiphonycteris) brachypterus (Peters, 1852)
DEMOCRATIC REPUBLIC OF THE CONGO: 2 ?? [2 UN], Abatupi River, unknown date, leg.
unknown collector (INPBC: V1285.4.1, V1285.4.2); 1 ad ♀ [AL], Léopoldville [= Kinshasa],
unknown date, leg. Guy Babault (MNHN: ZM-MO-1984-1212); 1 ad ♀, 17 ♀♀, 3 ♂♂, 6 ?? [19 SS,
2 SA, 6 UN], Medje, 25 Jan. 1910, 6 Mar. 1910, 16 Mar. 1910, leg. Herbert Lang, James Paul Chapin
and The American Museum Congo Expedition, 16 Mar. 1910, leg. Herbert Lang, 16 Mar. 1910, leg.
unknown collector (AMNH: 48819, 48820, 48821 [holotype Nyctinomus ochraceus J.A. Allen, 1917],
48822 to 48826, 48828 to 48835, 49252, 49253, 49258, 49260, 49267, FMNH: 43876, 43877, MCZ:
17390, 17391, AMNH: 49224, RMCA: 12391); 1 ad ♀ [SS], Panga, 18 Sep. 1914, leg. Herbert Lang,
211
European Journal of Taxonomy 382: 1–327 (2017)
James Paul Chapin and The American Museum Congo Expedition (AMNH: 48843); 1 ♀ [SS], Tandala,
16 Jul. 1979, leg. C. Brian Robbins (USNM: 537769); 1 ? [UN], Thysville [= Mbanza-Ngungu],
11 Feb. 1987, leg. unknown collector (RBINS: 22432); 2 ♀♀ [2 SS], Yalosemba, 17 Jun. 1979, leg.
C. Brian Robbins (USNM: 537747, 537768).
RWANDA: 4 ?? [2 UN, 2 SN], unknown locality, unknown date, leg. Anthony N. Start, unknown date,
leg. unknown collector (BMNH: 1975.2809, 1975.2809_1, 1975.2809_2, 1975.2810).
Mops (Xiphonycteris) nanulus J.A. Allen, 1917
DEMOCRATIC REPUBLIC OF THE CONGO: 3 ?? [3 AL], unknown locality, unknown date, leg.
Anthony N. Start, unknown date, leg. unknown collector, between 1 Jan. 1976 and 31 Dec. 1977, leg.
M. Lejeune (RMCA: 77.029-M-0003 to 77.029-M-0005); 1 ♀ [SS], Aba, 17 Dec. 1911, leg. Herbert
Lang, James Paul Chapin and The American Museum Congo Expedition (AMNH: 48874); 1 ? [AL],
Albertville [= Kalemie], prior to 18 Nov. 1959, leg. H. Bomans (RMCA: 29304); 4 ♀♀, 6 ♂♂, 4 ??
[6 SA, 8 AL], Beno, unknown date, leg. the Royal Museum for Central Africa (RMCA), between
1 Jan. 1949 and 31 Dec. 1949, leg. P. Bayet (BMNH: 1954.840 to 1954.843, RMCA: 19838 to 19847);
3 ?? [3 UN], Ihunga-Semliki conluence, unknown date, leg. unknown collector (INPBC: V794.1 to
V794.3); 2 ♀♀ [2 UN], Isai River, unknown date, between 18 Nov. 1974 and 19 Nov. 1974, leg.
Michael D. Gallagher (HZM: 7.7818, 8.7819); 1 ? [AL], Kasai, prior to 11 Jul. 1905, leg. La Compagnie
du Kassai (RMCA: 740a); 6 ad ♀♀ [6 UN], Kikwit, 3 Aug. 1995, leg. Herwig Leirs et al. (MSB: 274898
to 274900, 274902, 274903, 274907); 3 ad ♀♀, 4 ad ♂♂ [5 SA, 1 AL, 1 SK], Kikwit ITPK [= Kikwit],
3 Aug. 1995, leg. Herwig Leirs et al. (RMCA: 97.021-M-0817 to 97.021-M-0822, 97.021-M-0850);
1 juv ♂ [SA], Kinzambi Mission, 24 Jul. 1995, leg. Herwig Leirs et al. (RMCA: 97.021-M-0816);
1 juv ♂ [AL], Leverville [= Lusanga], between 1 Feb. 1929 and 28 Feb. 1929, leg. Tinant (RMCA:
9988b); 1 ? [AL], Lulimbi, between 1 Jan. 1976 and 31 Dec. 1977, leg. M. Lejeune (RMCA: 77.029M-0002); 6 ad ♀♀, 2 ad ♂♂, 1 ad? ♂, 1 sad ♀, 1 juv ♀, 2 ♀♀, 1 ? [10 SS, 1 SA, 3 AL], Luluabourg
[= Kananga], unknown date, prior to 8 Jul. 1913, leg. Richard Callewaert, prior to 17 Apr. 1913, leg.
Cambier (BMNH: 1926.11.1.37 to 1926.11.1.43, 1927.12.21.19, 1927.5.9.1, 1927.5.9.2, RMCA: 1548,
1954a to 1954c); 1 ♀, 1 ? [1 SS, 1 SA], N’gombe [= Ngombe], unknown date, between 1 Jan. 1921 and
31 Dec. 1921, leg. Henri Schouteden (RMCA: 16451, 16608); 2 ad ♂♂, 3 ♀♀, 1 ♂ [6 SS], Niangara,
12 Dec. 1910, leg. Herbert Lang, James Paul Chapin and The American Museum Congo Expedition
(AMNH: 48861 [topotype Mops (Allomops) nanulus J.A. Allen, 1917], 48862 [topotype Mops
(Allomops) nanulus J.A. Allen, 1917], 48863 [topotype Mops (Allomops) nanulus J.A. Allen, 1917],
48864 [holotype Mops (Allomops) nanulus J.A. Allen, 1917], 48865 [topotype Mops (Allomops)
nanulus J.A. Allen, 1917], FMNH: 43868); 4 ♀♀, 1 ♂, 2 ?? [7 SS], Panga, between 1 Sep. 1925 and
30 Sep. 1925, leg. Henri Schouteden (RMCA: 9132 to 9135, 9141 to 9143); 4 ♀♀ [4 SS], Tandala,
12 Jul. 1979, 16 Jul. 1979, 21 Jul. 1979, leg. C. Brian Robbins (USNM: 537758 to 537761); 2 ♀♀
[2 SS], Yalosemba, 27 Jun. 1979, 28 Jun. 1979, leg. C. Brian Robbins (USNM: 537754, 537756).
Mops (Xiphonycteris) spurrelli (Dollman, 1911)
DEMOCRATIC REPUBLIC OF THE CONGO: 8 ♀♀, 1 ♂ [9 SA], Butembo, 16 Mar. 1959, prior to
26 May 1959, leg. P. Dyleff (RMCA: 27876, 27868 to 27875); 1 ♂ [UN], St. Jos. (Mission) [= Mikalaya],
12 Oct. 1924, leg. Richard Callewaert (AMNH: 86145); 1 ♀ [SS], Tandala, 16 Jul. 1979, leg. C. Brian
Robbins (USNM: 537757); 2 ♀♀ [2 SS], Yalosemba, 17 Jun. 1979, 27 Jun. 1979, leg. C. Brian Robbins
(USNM: 537753, 537755).
212
VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
Mops (Xiphonycteris) thersites (Thomas, 1903)
DEMOCRATIC REPUBLIC OF THE CONGO: 1 ♂ [SA], unknown locality, between 1 Jan. 1911
and 31 Dec. 1911, leg. unknown collector (SMF: 6396); 1 ad ♂ [SS], Avakubi, 13 Feb. 1914, leg.
Herbert Lang, James Paul Chapin and The American Museum Congo Expedition (AMNH: 48851
[holotype Mops (Allomops) occipitalis J.A. Allen, 1917]); 1 ♂ [UN], Banana, 29 Sep. 1955, leg. l’
Expédition Atlantique Sud 1955 (RBINS: 15457); 1 ♀ [AL], Bombanga - Bumba, between 1 Jan. 1986
and 31 Dec. 1986, leg. WHO and Marc Colyn (RMCA: 88.011-M-0026); 1 ♀ [UN], Isai River,
19 Dec. 1974, leg. Michael D. Gallagher (HZM: 3.7820); 17 ♀♀, 5 ♂♂ [22 UN], Kasenye, 10 km N,
17 Feb. 1954, 10 Mar. 1954, 17 Mar. 1954, 20 Mar. 1954, 30 Mar. 1954, 22 Nov. 1954, 23 Nov. 1954,
24 Dec. 1954, leg. unknown collector (RBINS: 15470, 15472, 15469, 15471, 15460.01 to 15460.13,
15468, 15474.1, 15474.2, 15466, 15467); 1 ad ♂ [SK], Kikwit ITPK [= Kikwit], 3 Aug. 1995,
leg. Herwig Leirs et al. (RMCA: 97.021-M-0824); 2 ♀♀, 2 ♂♂ [4 SS], Luebo, 19 Aug. 1921, leg.
Henri Schouteden (RMCA: 6607, 6608, 7298, 7299); 2 ♀♀, 2 ♂♂, 2 ?? [6 SS], Lulua [= Kananga],
25 Feb. 1981, 27 Feb. 1981, leg. Betu Nsankulu (RMCA: 81.050-M-0002, 81.050-M-0005, 81.050M-0006, 81.050-M-0003, 81.050-M-0004, 81.050-M-0007); 10 ♀♀, 1 ♂ [9 SS, 2 SK], Luluabourg
[= Kananga], unknown date, leg. Richard Callewaert, 15 Jan. 1964, leg. Antoon Emeric Marcel De Roo
(BMNH: 1926.11.1.28 to 1926.11.1.31, 1926.11.1.33, 1926.11.1.34, 1926.11.1.36, 1927.12.21.15,
1927.12.21.17, 1927.12.21.18, AMNH: 206845); 1 ad ♀ [UN], Lusano and KKK Zone, unknown date,
leg. unknown collector (DNSM: 14073); 3 ♀♀, 3 ♂♂ [2 SS, 4 UN], Medje, 19 Jan. 1910, 15 Sep. 1910,
16 Sep. 1910, between 16 Sep. 1910 and 19 Sep. 1910, leg. Herbert Lang, James Paul Chapin and The
American Museum Congo Expedition (AMNH: 48846 to 48848, 48850, 48845, RMCA: 12362); 2 ♂♂
[2 AL], Mosenge, 11 Aug. 1986, 14 Aug. 1986, leg. WHO and Marc Colyn (RMCA: 88.011-M-0024,
88.011-M-0025); 1 ? [SS], Ndwa, prior to 27 Mar. 1952, leg. Ngwe (RMCA: 20837); 1 ♂ [SS], St. Jos.
(Mission) [= Mikalaya], 14 Nov. 1924, leg. Richard Callewaert (AMNH: 86101); 3 ♀♀, 1 ♂, 1 ? [4 SS,
1 AL], St. Joseph de Luluabourg Mission [= Mikalaya], 19 May 1922, 3 Jun. 1922, 19 Jul. 1922,
27 Jul. 1922, leg. Richard Callewaert, 24 May 1939, leg. Jean Jacques Deheyn (RMCA: 7550, 7542,
15398, 7544, 7549); 1 ♀, 3 ♂♂ [4 AL], Tandala, 12 Jul. 1979, leg. C. Brian Robbins, 7 Jul. 1979, leg.
unknown collector (USNM: 463530, MHNG: 1871.007 to 1871.009); 4 ♀♀, 13 ♂♂ [5 SS, 2 SA, 9 AL,
1 SO], Yalosemba, 17 Jun. 1979, 27 Jun. 1979, leg. unknown collector, 17 Jun. 1979, 27 Jun. 1979,
leg. C. Brian Robbins (MHNG: 1871.003 to 1871.005, USNM: 463523 to 463528, 537763 to 537767,
MHNG: 1871.002, 1871.006, USNM: 463529).
RWANDA: 15 ♀♀, 3 ♂♂, 2 ?? [20 UN], Astrida [= Butare], 26 Feb. 1966, leg. unknown collector
(RBINS: 15475.1, 15475.2, 15478.1, 15478.2, 15482.01 to 15482.10, 15932.1 to 15932.6); 1 ? [SK],
Byumba, 18 Apr. 1990, leg. Manuel Ruedi and Raphaël Arlettaz (MHNG: 1919.023); 3 ♀♀ [3 UN],
Gihinda [= Gihinga], 13 Feb. 1966, leg. unknown collector (RBINS: 15476.1 to 15476.3); 1 ♀ [UN],
Nyumba Cave, 13 Feb. 1966, leg. unknown collector (RBINS: 15477); 1 ♀ [SS], Rubona, 8 Jan. 1967,
leg. Félix / Michel Anciaux de Faveaux (RBINS: 16036).
Myopterus E. Geoffroy Saint-Hilaire, 1818
Myopterus daubentonii Desmarest, 1820
DEMOCRATIC REPUBLIC OF THE CONGO: 1 ? [AL], Bunia, prior to 20 Feb. 1939, leg. Maristes
(RMCA: 15224); 1 ad ♀, 6 ♀♀, 1 ♂ [7 SS, 1 AL], Niangara, 26 Dec. 1910, 27 Dec. 1910, leg. Herbert
Lang, James Paul Chapin and The American Museum Congo Expedition, 26 Dec. 1910, leg. unknown
collector (AMNH: 48852 to 48856, MCZ: 17384, AMNH: 48857, 49228); 2 ?? [1 SO, 1 UN], Uele
[River], 28 Apr. 1914, prior to 28 Apr. 1914, leg. M. Hutereau (BMNH: 1919.5.9.1 [holotype Myopterus
albatus Thomas, 1915], RMCA: 2911).
213
European Journal of Taxonomy 382: 1–327 (2017)
Myopterus whitleyi (Scharff, 1900)
DEMOCRATIC REPUBLIC OF THE CONGO: 1 ♂ [UN], unknown locality, 5 May 1940, leg.
NATURE ??? (TM: 9766); 1 ♂ [UN], Banana, 29 Sep. 1955, leg. l’ Expédition Atlantique Sud 1955
(RBINS: 15459); 3 ?? [3 AL], Beno, prior to 3 Jan. 1950, between 1 Jan. 1949 and 31 Dec. 1949, leg.
P. Bayet (RMCA: 19826, 19827, 19476); 1 ♀ [UN], Kasenye, 10 km N, 6 Mar. 1954, leg. l’ Expédition
Atlantique Sud 1955 (RBINS: 15473); 1 ad ♀ [UN], Kikwit ITPK [= Kikwit], 3 Aug. 1995, leg.
Herwig Leirs et al. (MSB: 274906); 1 ad ♀ [SA], Kinzambi Mission, 30 Jul. 1995, leg. Herwig Leirs
et al. (RMCA: 97.021-M-0825); 1 ♀ [UN], Léopoldville [= Kinshasa], 7 Oct. 1950, leg. unknown
collector (RBINS: 7478); 11 ♀♀, 1 ♂, 5 ?? [10 SS, 5 AL, 2 SK], Luluabourg [= Kananga], unknown
date, 22 May 1922, 13 Jun. 1922, prior to 8 Jul. 1913, between 1 Jan. 1913 and 31 Dec. 1913, leg.
Richard Callewaert (BMNH: 1926.11.1.45, 1926.11.1.46, 1926.7.6.100 to 1926.7.6.103, 1926.7.6.99,
1934.10.24.27, 1934.10.24.28, RMCA: 7589, 7590, 7588, 16196, 16197, 1954h to 1954j); 4 ♂♂ [2 SS,
2 SO], St. Jos. (Mission) [= Mikalaya], 7 Aug. 1924, 26 Sep. 1924, 10 Oct. 1924, 23 Oct. 1924, leg.
Richard Callewaert (AMNH: 86144, 86148, 86140, 86176).
Otomops Thomas, 1913
Otomops martiensseni (Matschie, 1897)
BURUNDI: 1 ♂ [UN], Kibira National Park, 27 Mar. 1990, leg. unknown collector (FMNH: 137633);
1 ♂ [AL], Rwegura, 25 Apr. 1969, leg. P.L. Niort (MHNG: 1896.021).
DEMOCRATIC REPUBLIC OF THE CONGO: 1 ad ♂ [SS], Cel II, 8 Sep. 1952, leg. H. de Saeger
et al. (RBINS: 13827); 1 ♂ [UN], Marungu, unknown date, leg. unknown collector (NMZL: 3395);
1 ad ♂ [SS], Mbuye-Bala River, 26 Mar. 1948, leg. unknown collector (RBINS: 10704); 1 ♀ [SS],
Poko, 16 Jul. 1914, leg. Cuthbert Christy (BMNH: 1919.5.9.2).
RWANDA: 1 sad ? [SN], unknown locality, 30 Nov. 1985, leg. unknown collector (SMF: 66857); 1 ♂
[SA], Kigali, between 1 Jan. 1933 and 31 Dec. 1933, leg. Becquet (RMCA: 12166).
Tadarida Rainesque, 1814
DEMOCRATIC REPUBLIC OF THE CONGO: 1 ? [UN], unknown locality, unknown date, leg.
unknown collector (BMNH: 1990.234); 1 ? [AL], Léopoldville [= Kinshasa], unknown date, leg.
Cuthbert Christy (BMNH: 1904.6.25.2).
Tadarida fulminans (Thomas, 1903)
DEMOCRATIC REPUBLIC OF THE CONGO: 1 ? [UN], Rumangabo, unknown date, leg. unknown
collector (INPBC: V1234).
RWANDA: 1 ? [SS], Nyakibanda, between 1 Apr. 1955 and 30 Apr. 1955, leg. Alexandre Fain (RMCA:
23748); 1 ad ♀ [UN], Nyanza, 6 Feb. 1966, leg. unknown collector (RBINS: 15494); 1 ad ♀, 1 ad
♂, 5 ♀♀, 9 ♂♂ [16 SS], Rukara, 27 Aug. 1955, 12 Sep. 1955, 13 Sep. 1955, 15 Sep. 1955, between
12 Sep. 1955 and 30 Sep. 1955, leg. Alexandre Fain (RMCA: 37972 to 37978, 81.012-M-0104, 37979
to 37982, 81.012-M-0103, 37970, 37971, 81.012-M-0047).
Tadarida ventralis (Heuglin, 1861)
DEMOCRATIC REPUBLIC OF THE CONGO: 1 ♀ [SS], Alima, 2 Aug. 1960, leg. P. Dyleff (RMCA:
29241).
214
VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
Vespertilionoidea Gray, 1821
DEMOCRATIC REPUBLIC OF THE CONGO: 1 ad ?, 3 ?? [4 AL], unknown locality, 20 Nov. 1988,
between 1 Jan. 1921 and 31 Dec. 1921, leg. Verlaine, 4 Jul. 1958, leg. E. Jans (RMCA: 88.044-M-0014,
88.044-M-0015, 27368, 88.044-M-0013); 1 imm ♀ [SO], Bambesa, 18 Mar. 1937, leg. J.M. Vrijdagh
(RMCA: 14073); 1 juv ?, 2 ♀♀, 2 ?? [5 AL], Boende, between 1 Jan. 1967 and 31 Dec. 1967, between
1 Jan. 1966 and 31 Jan. 1966, leg. P. Lootens (RMCA: 34077, 34093, 34156, 38732, 38733); 1 imm ?
[AL], Bokuma, 25 Jul. 1952, leg. P. Lootens (RMCA: 21672); 1 juv ? [AL], Bunkeya, 26 Jan. 1958, leg.
Van Oost (RMCA: 28802); 2 ?? [1 AL, 1 UN], Elisabethville [= Lubumbashi], between 1 Jan. 1930
and 31 Dec. 1930, leg. Lamoral, prior to 26 Jan. 1934, leg. Quarré (RMCA: 11708, 12236); 1 ? [SS],
Grotte de Baya [= Baya Cave], 14 May 1956, leg. M. Lips (RMCA: 23799); 1 ? [AL], Ibembo,
25 Jan. 1951, leg. Jozef Hutsebaut (RMCA: 89.004-M-0001); 1 ♀ [SS], Kakontwe Cave, 8 Feb. 1957,
leg. G. Verhoustraete (RMCA: 26212); 1 ad ♀ [AL], Kalutenge, between 19 May 1919 and 19 May
1924, leg. Seydel (RMCA: 90.006-M-0001); 1 ♀, 1 ? [1 SS, 1 UN], Léopoldville [= Kinshasa],
14 Feb. 1945, leg. Lepersonne, 20 Jan. 1947, leg. Dartevelle (RMCA: 17338, 18164); 1 ? [AL], Lula,
prior to 4 Jul. 1958, leg. A.J. Jobaert (RMCA: 27201); 1 ad ♀, 1 neonate ?, 1 ♀, 2 ?? [5 AL], Luluabourg
[= Kananga], unknown date, 3 Jul. 1965, leg. Jan Deheegher, 16 Feb. 1965, between 30 Dec. 1964
and 25 Feb. 1965, leg. Antoon Emeric Marcel De Roo (RMCA: 31573, 33351, 33961, 33962, 33349);
1 ♀ [AL], Mosenge, 15 Aug. 1986, leg. WHO and Marc Colyn (RMCA: 88.011-M-0022); 2 juv ??
[2 AL], Rumanura River, 3 Dec. 1954, leg. Gaston-François de Witte (RMCA: 37376, 37377); 1 ♀
[AL], Uvira, 5 Sep. 1956, leg. Laurent (RMCA: 35692); 5 ?? [5 AL], Wafanya [= Wafania], unknown
date, 26 May 1992, 29 May 1992, between 1 Apr. 1992 and 30 Apr. 1992, between 1 Mar. 1992 and
31 Mar. 1992, leg. P. Lootens (RMCA: 92.079-M-0150, 92.079-M-0042, 92.079-M-0043, 92.079-M0103, 92.079-M-0104); 8 juv ?? [8 AL], Yalusaka, 16 Mar. 1958, leg. F. Carbone (RMCA: 27166
to 27173); 1 ? [SO], Yolohotiri Cave, between 1 Sep. 1951 and 30 Sep. 1951, leg. Alexandre Fain
(RMCA: 23299).
Miniopteridae Dobson, 1875
Miniopterus Bonaparte, 1837
Miniopterus fraterculus Thomas & Schwann, 1906
BURUNDI: 1 ♀ [SB], Giserama, Colline, 2 Aug. 1991, leg. Julian C. Kerbis Peterhans (FMNH: 148984);
5 ♂♂ [1 SB, 4 NS], Mpinga Cave [= Mpinga Mission], 31 Dec. 1968, 1 Jan. 1969, 25 Mar. 1970, leg.
P.L. Niort (MHNG: 1607.077 to 1607.079, 1607.098, 1607.082); 6 ♂♂ [5 SB, 1 SN], Nyamugari Hill,
10 Sep. 1990, 11 Sep. 1990, 12 Sep. 1990, leg. L. Davenport (FMNH: 148369, 148365, 148371 to
148373, 148370).
DEMOCRATIC REPUBLIC OF THE CONGO: 1 ad ♂, 9 ♀♀, 8 ♂♂, 2 ?? [19 SS, 1 SO], Aba,
unknown date, 16 Dec. 1911, leg. Herbert Lang, James Paul Chapin and The American Museum Congo
Expedition, 16 Dec. 1911, leg. unknown collector (AMNH: 51271, 51272, 49017, 49018, 49019
[holotype Miniopterus breyeri vicinior J.A. Allen, 1917], 49020 to 49029, 49031, 49032, MCZ:
17380, 17381, RMCA: 12397); 1 ? [UN], Albert National Park [= Virunga National Park], unknown
date, leg. unknown collector (INPBC: V1766.1); 1 ♂ [AL], Bushokwe Caves, 1 Aug. 1956, leg. Alvin
Novick (MCZ: 48263); 4 ♀♀ [4 SS], Butembo, 10 Aug. 1947, 14 Aug. 1947, leg. unknown collector
(RBINS: 7081 to 7084); 2 ?? [2 UN], Gangala-na-Bodio, 1 May 1948, leg. unknown collector (RBINS:
8124.1, 8124.2); 1 ♀ [SS], Hoyo, Mount, 10 Aug. 1947, leg. unknown collector (RBINS: 7085); 1 ♀,
1 ♂ [2 UN], Ituri Forest, between 1 Jan. 1947 and 31 Dec. 1948, leg. unknown collector (RBINS:
7475, 7476); 2 ?? [2 AL], Kabatibi Cave [= Mount Hoyo Caves], unknown date, leg. The Cambridge
East Africa Expedition 1959 (BMNH: 1960.108, 1960.109); 1 ? [AL], Lwiro [= Luiro], 4 Jun. 1956,
leg. Uwe Rahm (RMCA: 33533); 3 ?? [1 SS, 2 AL], Matupi Cave, unknown date, leg. The Cambridge
215
European Journal of Taxonomy 382: 1–327 (2017)
East Africa Expedition 1959, between 1 Sep. 1951 and 30 Sep. 1951, leg. Alexandre Fain (BMNH:
1960.112, RMCA: 21464, 23300); 4 ?? [2 SS, 2 AL], Mount Hoyo [= Hoyo, Mount], unknown date,
leg. The Cambridge East Africa Expedition 1959 (BMNH: 1960.106, 1960.107, 1960.110, 1960.111);
2 ?? [2 UN], Mugongo, unknown date, leg. unknown collector (INPBC: V1766.2, V1766.3); 1 ? [AL],
Mulungu, prior to 1 Feb. 1940, leg. F.L. Hendricks (RMCA: 15869); 4 ?? [4 UN], Muvo, Mount,
unknown date, leg. unknown collector (INPBC: V1228.1 to V1228.4); 1 ? [UN], Talatala, unknown
date, leg. unknown collector (RBINS: 7096).
RWANDA: 18 ad ♂♂ [10 SA, 8 AL], Mutura, 15 Dec. 1984, 13 Jan. 1985, leg. Benny Baeten and Luc
Janssens, 27 Oct. 1981, leg. Frits De Vree, Benny Baeten and Victor Van Cakenberghe (RMCA: 85.006M-0158, 85.006-M-0152 to 85.006-M-0157, 82.006-M-0328 to 82.006-M-0338).
Miniopterus inlatus Thomas, 1903
BURUNDI: 4 ♂♂ [4 NS], Mpinga Cave [= Mpinga Mission], 1 Jan. 1969, 25 Mar. 1970, leg. P.L. Niort
(MHNG: 1607.080, 1607.081, 1608.001, 1608.002).
DEMOCRATIC REPUBLIC OF THE CONGO: 2 ?? [2 AL], Baya Carrière [= Baya Quarry],
26 Jun. 1962, 26 Jun. 1963, leg. Parent (RMCA: 31507, 31508); 5 ?? [5 UN], Gangala-na-Bodio,
9 Apr. 1948, leg. H. de Saeger et al. (RBINS: 13577.1 to 13577.5); 5 ♀♀, 33 ♂♂ [38 UN], Goroumbwa
Mine, 17 May 1999, 18 May 1999, 19 May 1999, 19 Oct. 1999, 22 Oct. 1999, 23 Oct. 1999, 24 Oct. 1999,
25 Oct. 1999, leg. Robert (Bob) Swanepoel (TM: DRC0033, DRC0107, DRC0122, DRC0150,
DRC0156, DBA010, DBA011, DBA015, DBA113, DBA115, DBA164 to DBA177, DBA198,
DBA202, DBA239 to DBA244, DBA255, DBA277, DBA300, DBA317, DBA321, DBA327); 1 ?
[UN], Issehe River, unknown date, leg. unknown collector (INPBC: W1457.3); 2 ♂♂, 1 ? [3 UN],
Jadotville [= Likasi], 28 Jun. 1960, leg. unknown collector, 7 Apr. 1957, leg. Félix / Michel Anciaux
de Faveaux (RBINS: 1696, 1697, RMCA: 26265); 3 ?? [3 AL], Kabambi, 13 Jul. 1955, leg. GastonFrançois de Witte (RMCA: 37170 to 37172); 1 ♂ [SS], Kafwe, 5 Jul. 1948, leg. unknown collector
(RBINS: 10705); 2 ♀♀, 4 ♂♂ [3 AL, 3 UN], Kakontwe Cave, unknown date, leg. the Royal Museum
for Central Africa (RMCA), 13 Feb. 1957, 12 Jul. 1957, leg. Félix / Michel Anciaux de Faveaux (BMNH:
1959.513 to 1959.515, RMCA: 26740, 26248, 26249); 3 ♀♀, 2 ♂♂ [4 AL, 1 UN], Kalomoni Cave
[= Kakontwe], 3 Jan. 1964, 8 May 1965, leg. Félix / Michel Anciaux de Faveaux (RMCA: 33993 to
33996, 34017); 11 ♀♀, 2 ♂♂ [13 AL], Kalumbu Cave, 1 Dec. 1957, leg. Félix / Michel Anciaux de
Faveaux (RMCA: 27548 to 27552, 27554, 27555, 27559 to 27564); 1 ♀, 1 ♂ [2 SA], Kasoma Cave,
13 Aug. 1962, leg. Félix / Michel Anciaux de Faveaux (MHNG: 1046.094, 1046.095); 3 ♀♀ [3 AL],
Kasowena Cave, 28 Jun. 1964, leg. Félix / Michel Anciaux de Faveaux (RMCA: 34007 to 34009); 1 ad
♀, 2 sad ♂♂, 3 ♀♀ [1 SA, 5 AL], Katanga (former) Province, 1 Dec. 1957, 13 Aug. 1962, leg. Félix
/ Michel Anciaux de Faveaux (SMF: 18926, 18927, 18931, 21310 to 21312); 1 ad ♂ [SS], Katobwe,
28 Nov. 1926, leg. John Todd Zimmer (FMNH: 29416 [holotype Miniopterus rufus Sanborn, 1936]);
4 ♂♂ [4 AL], Komanda, 21 Aug. 1976, leg. Frieder Meier (MHNG: 1874.006 to 1874.009); 1 ♀ [SS],
Masombwe, 20 Sep. 1948, leg. unknown collector (RBINS: 10706); 1 ♀, 1 ♂ [2 SS], Mbuji-Mayi,
32 km SE, 25 Feb. 1974, leg. Western New Mexico University (FMNH: 152883, 152884); 5 ♂♂, 1 ?
[4 AL, 2 SK], Mount Hoyo Caves, 5 Oct. 1957, leg. Robin E. Leech and Edward S. Ross (CAS:
12500 to 12503, 12531, 12532); 1 juv ♂, 1 ♀ [2 AL], Mura Pumping Station, 20 Oct. 1963, leg.
Félix / Michel Anciaux de Faveaux (RMCA: 33989, 33990); 1 ♀, 6 ♂♂ [6 AL, 1 UN], Mwanga Cave,
30 Jun. 1964, leg. Félix / Michel Anciaux de Faveaux (RMCA: 34010 to 34016); 1 ♀ [AL], Pierkat
Quarry [= Baya Quarry], 12 Mar. 1956, leg. Félix / Michel Anciaux de Faveaux (RMCA: 23878); 1 ♂
[SK], Talatala, 11 Aug. 1947, leg. unknown collector (RBINS: 7086); 1 ? [AL], Tsebahu, 12 Jul. 1955,
leg. Gaston-François de Witte (RMCA: 37169); 2 ?? [2 UN], Tshebahu River, unknown date, leg.
unknown collector (INPBC: W1456a, W1456b).
216
VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
RWANDA: 7 ?? [7 SK], Kibumbu Crater, 9 Dec. 1991, leg. J.R. Wilson and Anthony Michael
Hutson (BMNH: 1992.391 to 1992.397); 5 ad ♀♀, 69 ad ♂♂ [40 SA, 34 AL], Mutura, 15 Dec. 1984,
16 Dec. 1984, 18 Dec. 1984, 12 Jan. 1985, 13 Jan. 1985, leg. Benny Baeten and Luc Janssens,
22 Oct. 1981, 23 Oct. 1981, 25 Oct. 1981, 26 Oct. 1981, 27 Oct. 1981, leg. Frits De Vree, Benny Baeten
and Victor Van Cakenberghe (RMCA: 85.006-M-0136 to 85.006-M-0151, 85.006-M-0120 to 85.006M-0135, 82.006-M-0286 to 82.006-M-0327); 2 ♂♂ [2 UN], Ruhengeri, 7 Jun. 1966, leg. unknown
collector (RBINS: 15490.1, 15490.2).
Miniopterus minor Peters, 1867
DEMOCRATIC REPUBLIC OF THE CONGO: 3 ?? [1 SA, 2 AL], Kinshasa, between 1 Oct. 1987
and 31 Oct. 1987, leg. WHO and Marc Colyn (RMCA: 88.011-M-0029 to 88.011-M-0031); 1 ? [UN],
Kisala [= Kisala-Kiama], unknown date, leg. unknown collector (INPBC: W1462); 2 ♂♂ [1 SA, 1 UN],
Thysville [= Mbanza-Ngungu], unknown date, leg. the Royal Museum for Central Africa (RMCA),
between 1 Nov. 1935 and 14 Jan. 1936, leg. unknown collector (BMNH: 1954.866, RBINS: 1456); 1 ?
[SA], Thysville, Cave [= Banjan Caves], between 1 Apr. 1947 and 30 Apr. 1947, leg. Alexandre Fain
(RMCA: 18016); 1 ? [SA], Thysville, Cave b 12 a, 30 May 1949, leg. Narcisse Leleup (RMCA: 22502).
Miniopterus natalensis (A. Smith, 1833)
DEMOCRATIC REPUBLIC OF THE CONGO: 2 ♀♀, 8 ♂♂ [6 SA, 4 AL], unknown locality,
24 Apr. 1962, 20 May 1962, 30 Jun. 1962, 18 Jul. 1962, 13 Aug. 1962, leg. Félix / Michel Anciaux
de Faveaux (SMF: 21309, 21300 to 21302, 21308, 21304, 21305, 21303, 21306, 21307); 1 ? [AL],
Albertville [= Kalemie], prior to 18 Nov. 1959, leg. H. Bomans (RMCA: 29303); 1 ad ♂, 44 ?? [1 SA,
44 AL], Baya Carrière [= Baya Quarry], 17 May 1965, leg. Walter N. Verheyen (RMCA: 34018 to
34043, 35020, 35021, 35025, 35027, 35029 to 35041, 35044, 35045); 1 ♀, 1 ♂ [2 AL], Baya Cave
[= Baya Quarry], 12 Mar. 1956, leg. Félix / Michel Anciaux de Faveaux (RMCA: 23821, 23921);
52 ♀♀, 51 ♂♂ [103 AL], Defrenne Cave [= Mwanga Cave], 21 Jun. 1959, leg. Félix / Michel Anciaux de
Faveaux (RMCA: 28215 to 28317); 1 ♂ [AL], Dethioux Farm, Cave II [= Shinkolobwe], 15 Jun. 1958,
leg. Félix / Michel Anciaux de Faveaux (RMCA: 27914); 3 ad ♂♂, 1 ♂ [4 AL], Dethioux Hoeve, Grot I
[= Shinkolobwe], 29 Sep. 1957, leg. Félix / Michel Anciaux de Faveaux (MHNG: 1046.088 to 1046.091);
2 ad ♂♂, 14 ♀♀, 6 ♂♂ [2 SA, 20 UN], Elisabethville [= Lubumbashi], unknown date, leg. Félix /
Michel Anciaux de Faveaux, 25 Mar. 1966, leg. unknown collector (MNHN: ZM-MO-1957-127 A, ZMMO-1957-127 B, RBINS: 17204.01 to 17204.20); 11 ♀♀ [11 SA], Grotte de Baya [= Baya Cave],
12 Mar. 1956, 14 May 1956, 22 May 1956, leg. Félix / Michel Anciaux de Faveaux (MHNG: 922.071
to 922.073, 922.075 to 922.081, 922.074); 1 ♂ [UN], Grottes Kiwakishi, 18 Oct. 1974, leg. Michael
D. Gallagher (HZM: 214.7800); 4 ♀♀, 5 ♂♂ [7 AL, 2 UN], Jadotville [= Likasi], 3 Jun. 1958, leg.
Félix / Michel Anciaux de Faveaux, 5 Apr. 1961, leg. unknown collector (RMCA: 27574 to 27580,
RBINS: 14618, 14619); 3 ♂♂ [2 AL, 1 UN], Kaboyaboya Cave, 24 Apr. 1962, 23 Jul. 1962, leg. Félix
/ Michel Anciaux de Faveaux, 23 Jul. 1962, leg. unknown collector (MHNG: 1046.093, RBINS: 14617,
MHNG: 1046.092); 1 ♂ [AL], Kakanda Cave, 22 Apr. 1959, leg. Félix / Michel Anciaux de Faveaux
(RMCA: 28166); 19 ♀♀, 13 ♂♂, 8 ?? [1 SA, 37 AL, 1 SK, 1 UN], Kakontwe Cave, 26 Jun. 1956,
28 Apr. 1957, 8 Jul. 1957, 12 Jul. 1957, 15 Jan. 1958, 6 Nov. 1958, leg. Félix / Michel Anciaux de
Faveaux, 6 Feb. 1958, leg. Robin E. Leech and Edward S. Ross, prior to 24 Sep. 1962, leg. L. Poelman
(RMCA: 26729 to 26739, 26741, 26745, 26754, 26755, 26790, 26805, 27565, 23900, MHNG: 1046.072
to 1046.078, CAS: 12510 to 12513, 12535, RMCA: 27566, 27567, 26727, 26728, 26809, 26879, 31218
to 31220); 2 ♀♀, 2 ♂♂ [4 AL], Kalumbu Cave, 1 Dec. 1957, leg. Félix / Michel Anciaux de Faveaux
(RMCA: 27553, 27556 to 27558); 13 ♀♀, 19 ♂♂ [30 AL, 2 UN], Kandu (Cave), 25 Jul. 1957, leg.
Félix / Michel Anciaux de Faveaux (MHNG: 1046.079 to 1046.083, RMCA: 26782, 26787, 26789,
26803, 26813, 26816, 26818, 26820, 26832, 26834, 26845 to 26850, 26854 to 26864); 2 ♀♀ [2 AL],
Kapolowe Mission [= Kapolowe], 16 Feb. 1956, leg. Félix / Michel Anciaux de Faveaux (RMCA:
217
European Journal of Taxonomy 382: 1–327 (2017)
23820, 23897); 1 ♀ [AL], Kasapa, 22 Apr. 1956, leg. Félix / Michel Anciaux de Faveaux (RMCA:
23899); 2 ♀♀, 1 ♂ [3 UN], Kasoma Cave, 13 Aug. 1962, leg. unknown collector (RBINS: 14622 to
14624); 7 ♀♀, 9 ♂♂ [12 SA, 4 AL], Katanga (former) Province, 18 Dec. 1957, 6 Feb. 1958, 1 May
1958, 3 Jun. 1958, 11 Jun. 1960, 28 Jun. 1960, 3 Apr. 1961, leg. Félix / Michel Anciaux de Faveaux
(SMF: 18928, 18935, 19074, 19076, 19077, 18932, 19072, 19073, 19075, 19078, 19573, 18929, 18930,
18936, 18933, 18934); 1 ? [AL], Kiwakishi Cave [= Kiamokoto], between 1 Jul. 1958 and 31 Jul. 1958,
leg. Narcisse Leleup (RMCA: 27293); 7 ♂♂ [7 UN], Kyamakonde Cave, 18 Jul. 1962, leg. unknown
collector (RBINS: 14610 to 14616); 1 ♀, 4 ♂♂ [5 AL], Kyantapo Cave, 29 Jul. 1957, leg. Félix / Michel
Anciaux de Faveaux (MHNG: 1046.086, RMCA: 26797, 26799, 26871, 26872); 1 ♀, 1 ♂, 1 ? [3 AL],
Kyasala Cave [= Kiasala Cave], 31 Mar. 1956, 28 Jul. 1957, leg. Félix / Michel Anciaux de Faveaux
(RMCA: 26796, 26801, 23896); 1 ♀ [AL], Kyasala Grande Grotte [= Kiasala Cave], 28 Jul. 1957,
leg. Félix / Michel Anciaux de Faveaux (MHNG: 1046.084); 5 ?? [1 SA, 4 AL], Lukatu Cave [= Ntoto
Cave], prior to 2 Feb. 2007, leg. A. Kimbembi-ma-Ibaka (RMCA: a7.006-M-0007 to a7.006-M-0011);
2 ad ♀♀, 1 juv ♀, 1 ♀, 1 ♂, 9 ?? [12 SS, 2 AL], Luluabourg [= Kananga], 13 Dec. 1963, 11 Jan. 1964,
1 Feb. 1964, 11 Feb. 1964, 12 Feb. 1964, 14 Feb. 1964, 21 Feb. 1964, 23 Feb. 1964, leg. Antoon Emeric
Marcel De Roo (AMNH: 207743 to 207745, 207740 to 207742, 207746 to 207749, 207753, 207750
to 207752); 3 ♀♀, 3 ♂♂ [6 AL], Lusolo Cave [= Lubudi], 28 Jul. 1957, leg. Félix / Michel Anciaux
de Faveaux (MHNG: 1046.085, RMCA: 26804, 26810, 26841, 26870, 26873); 93 ?? [93 AL], M’baya
Cave [= Baya Cave], 25 Mar. 1966, leg. Xavier Misonne (RMCA: 34619 to 34711); 1 ♀, 1 ♂ [2 AL],
Mpopola Cave, 2 May 1959, leg. Félix / Michel Anciaux de Faveaux (RMCA: 28191, 28192); 1 ♀,
5 ♂♂ [6 AL], Mulonga Cave, 30 Jul. 1957, 30 Aug. 1957, 1 May 1959, leg. Félix / Michel Anciaux
de Faveaux (RMCA: 28205 to 28207, MHNG: 1046.087, RMCA: 26842, 26788); 1 ? [AL], Ntoto
Cave, between 1 Jan. 2002 and 13 Jul. 2002, leg. A. Kimbembi-ma-Ibaka (RMCA: a2.040-M-0001);
10 ♀♀, 16 ♂♂ [26 AL], Pierkat Quarry [= Baya Quarry], 12 Mar. 1956, leg. Félix / Michel Anciaux
de Faveaux (RMCA: 23818, 23870 to 23877, 23879 to 23895); 5 ♀♀, 5 ♂♂ [10 UN], Salomoni Cave,
11 Jun. 1960, 28 Jun. 1960, 20 May 1961, leg. unknown collector (RBINS: 14602 to 14605, 14620,
14621, 14606 to 14609); 6 ♂♂ [6 AL], Swanepoel II Abyss, 1 May 1958, leg. Félix / Michel Anciaux de
Faveaux (RMCA: 27568 to 27573); 3 ♀♀, 2 ♂♂ [5 AL], Tantara Cave [= Shinkolobwe], 29 Sep. 1957,
29 Sep. 1967, leg. Félix / Michel Anciaux de Faveaux (RMCA: 26807, 26808, 26874, 26875, 26826);
1 ♂ [UN], Tenke, 6 Dec. 1959, leg. unknown collector (RBINS: 14625); 4 ad ♂♂, 7 ad ??, 2 ♀♀, 1 ♂,
30 ?? [6 SA, 7 AL, 31 UN], Thysville [= Mbanza-Ngungu], 1 Nov. 1935, 11 Feb. 1987, leg. unknown
collector, 2 Jul. 1905, 30 Mar. 1911, 2 Jun. 1915, 21 Jun. 1915, leg. Herbert Lang, James Paul Chapin
and The American Museum Congo Expedition (RBINS: 1456B, 22425.01 to 22425.30, AMNH: 49292,
49309, 49321, 49324, 49331, 49333 to 49335, MCZ: 17222, AMNH: 49288, 49289, 49318, 49328); 2 ??
[1 SS, 1 AL], Thysville, Cave [= Banjan Caves], unknown date, leg. Schwetz, prior to 6 Feb. 1930,
leg. Golenvaux (RMCA: 16450a, 10061); 1 ? [AL], Thysville, Cave b 12 a, 30 May 1949, leg. Narcisse
Leleup (RMCA: 22501).
Vespertilionidae Gray, 1821
DEMOCRATIC REPUBLIC OF THE CONGO: 1 juv ? [UN], Kantundwe, unknown date, leg. unknown
collector (INPBC: W616); 1 ♂ [AL], Karambi, 3 Apr. 1992, leg. Ngenge Masumbuko Kamitongo
(ZMA: MAM.24549); 1 juv ? [UN], Kasindi, unknown date, leg. unknown collector (INPBC: W979.2);
1 ? [AL], Kikondja, 6 Feb. 2010, leg. Michel Louette (RMCA: b0.004-M-0001).
Kerivoulinae Miller, 1907
Kerivoula Gray, 1842
DEMOCRATIC REPUBLIC OF THE CONGO: 1 ? [AL], Avakubi, prior to 12 Mar. 1913, leg. Cuthbert
Christy (RMCA: 1522a); 1 ? [SK], Gumba, 10 Oct. 1937, leg. Jean Jacques Deheyn (RMCA: 14263a).
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VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
Kerivoula argentata Tomes, 1861
DEMOCRATIC REPUBLIC OF THE CONGO: 1 ♀ [SK], Baliko, 7 Jun. 2012, leg. William (“Bill”)
T. Stanley (FMNH: 219604); 4 ♀♀ [2 SA, 2 UN], Kamatanda, 16 Jun. 1961, leg. unknown collector,
16 Jun. 1961, leg. Félix / Michel Anciaux de Faveaux (RBINS: 14470, 14471, SMF: 19574, 19575);
1 ♀ [SS], Kilwezi, 23 Jul. 1948, leg. unknown collector (RBINS: 10708); 5 ♀♀, 1 ? [3 SS, 1 SK, 2
UN], Luluabourg [= Kananga], unknown date, leg. Richard Callewaert, unknown date, leg. unknown
collector (BMNH: 1926.7.6.92, 1926.7.6.94, 1927.12.21.13, 1927.12.21.14, ZMB: 35910, 35911).
Kerivoula cuprosa Thomas, 1912
DEMOCRATIC REPUBLIC OF THE CONGO: 1 ? [SA], Eala, between 1 Jan. 1921 and 31 Jan. 1921, leg.
Henri Schouteden (RMCA: 16189); 1 ? [SA], Koteli [= Kotili], between 1 Dec. 1924 and 31 Dec. 1924,
leg. Henri Schouteden (RMCA: 16190); 1 ? [SA], Wafanya [= Wafania], between 1 Jan. 1986 and
31 Dec. 1987, leg. P. Lootens (RMCA: 87.043-M-0006).
Kerivoula lanosa (A. Smith, 1847)
DEMOCRATIC REPUBLIC OF THE CONGO: 1 ? [SA], Funda Biabo, 1 Apr. 1914, leg. Charliers
(RMCA: 5383); 1 ♂ [SA], Kamituga and Kampene, road between, 29 Oct. 1951, leg. P.C. Percy
(BMNH: 1952.1613); 1 ? [SA], Lukulu, between 1 Jun. 1932 and 30 Jun. 1932, leg. Remaele (RMCA:
12006); 1 ? [SA], Pawa, unknown date, leg. Reitter (RMCA: 15729).
Kerivoula phalaena Thomas, 1912
DEMOCRATIC REPUBLIC OF THE CONGO: 1 ♀ [AC], Irangi, 2 Feb. 1984, leg. Heinz Stephan
(SMF: 65005); 1 ♀ [AL], Lukolela, 30 Dec. 1930, leg. James Paul Chapin (AMNH: 86932); 1 ? [SA],
Wafanya [= Wafania], 28 May 1992, leg. P. Lootens (RMCA: 92.079-M-0152).
Kerivoula smithii Thomas, 1880
DEMOCRATIC REPUBLIC OF THE CONGO: 1 ad ♀, 1 juv ♂ [2 UN], Akenge [= Akengei],
7 Oct. 1913, leg. Herbert Lang, James Paul Chapin and The American Museum Congo Expedition
(AMNH: 49306, 49336); 4 ♀♀, 2 ♂♂ [6 AL], Aketi, 22 Jun. 2015, 26 Jun. 2015, 26 Aug. 2015, leg.
Vlir/Aketi (UNIKIS: AKETI605, AKETI606, AKETI623 to AKETI622); 3 ♀♀, 1 ? [1 SS, 2 SO,
1 UN], Avakubi, unknown date, 1 Oct. 1912, leg. Cuthbert Christy (BMNH: 1914.4.8.6, RMCA: 1425,
1427, 1428); 1 ? [SA], Bambesa, prior to 13 Sep. 1934, leg. H.J. Bredo (RMCA: 12530); 1 ? [SS],
Bukavu, prior to 14 Apr. 1955, leg. P.G. Vercammen (RMCA: 23172); 1 ? [SN], Kalehe Territory,
unknown date, leg. unknown collector (BMNH: 1962.1845); 1 ad ♂ [UN], Medje, 17 Jun. 1914, leg.
Herbert Lang, James Paul Chapin and The American Museum Congo Expedition (AMNH: 49016);
1 ♂ [SS], Stanleyville [= Kisangani], 26 Apr. 1912, leg. Cuthbert Christy (RMCA: 1422); 2 ♀♀, 1 ♂
[3 UN], Ukaika, unknown date, leg. Rudolf Grauer (NMW: 10298, 10299, 16728); 1 ? [SA], Wafanya
[= Wafania], between 1 May 1991 and 31 May 1991, leg. P. Lootens (RMCA: 91.076-M-0041).
Myotinae Tate, 1942
Myotis Kaup, 1829
Myotis bocagii (Peters, 1870)
BURUNDI: 1 ♂ [SB], Kanyinya, 6 Jun. 1969, leg. P.L. Niort (MHNG: 1607.056); 1 ♂ [SB], Mpinga
Mission, 26 Mar. 1970, leg. P.L. Niort (MHNG: 1608.003); 1 ♀ [SK], Resha, 3 Apr. 1979, leg. Heinz
Stephan (SMF: 57584).
219
European Journal of Taxonomy 382: 1–327 (2017)
DEMOCRATIC REPUBLIC OF THE CONGO: 1 ♀, 8 ?? [5 SS, 2 AL, 2 UN], unknown locality,
unknown date, leg. Herbert Lang, James Paul Chapin and The American Museum Congo Expedition,
unknown date, leg. unknown collector, between 1 Jan. 1985 and 31 Dec. 1986, leg. WHO and Marc
Colyn, prior to 28 Dec. 1971, leg. Gaston-François de Witte (AMNH: 51273 to 51277, BMNH: 1959.508,
1959.509, RMCA: 88.011-M-0016, 37418); 6 ♀♀, 6 ♂♂ [4 SS, 5 SO, 3 UN], Aba, 16 Dec. 1911,
17 Dec. 1911, 18 Dec. 1911, leg. Herbert Lang, James Paul Chapin and The American Museum Congo
Expedition, 17 Dec. 1911, leg. unknown collector (AMNH: 49001 to 49007, 49009, 49010, MCZ:
17377, RMCA: 12392, AMNH: 49011); 2 ♀♀, 2 ♂♂ [4 SS], Abimva, 21 Jun. 1925, 22 Jun. 1925,
23 Jun. 1925, leg. Henri Schouteden (RMCA: 9024, 9026, 9027, 9029); 2 ?? [2 AL], Albertville
[= Kalemie], prior to 30 Jan. 1900, leg. Célestin Hecq (RMCA: 422, 423); 6 ad ♀♀, 1 sad ♂ [4 SA,
3 UN], Athens Mission, 23 Jul. 1995, leg. Herwig Leirs et al. (MSB: 274603, 274604, 274614, RMCA:
97.021-M-0829 to 97.021-M-0832); 3 ♀♀, 5 ♂♂ [7 SS, 1 SO], Bafuaboka [= Bafwabaka], 4 Jan. 1910,
5 Jan. 1910, 6 Jan. 1910, 7 Jan. 1910, leg. Herbert Lang, James Paul Chapin and The American Museum
Congo Expedition (AMNH: 48989 to 48996); 1 ♂ [SS], Bafwabaka, between 6 Jan. 1910 and
6 Dec. 1910, leg. unknown collector (MCZ: 17376); 2 ♀♀, 1 ♂ [2 SA, 1 UN], Bafwaboka [= Bafwabaka],
6 Jan. 1910, 7 Jan. 1910, leg. Herbert Lang, James Paul Chapin and The American Museum Congo
Expedition (AMNH: 49193, 49300, 49301); 1 ♀, 1 ♂ [2 SS], Banalia, between 28 Jan. 1939 and
28 Dec. 1939, leg. Duncan M. Hodgson and William F. Coultas (MCZ: 38030, 38031); 2 ?? [2 AL], Bas
Congo (former) Province [= Kongo Central Province], unknown date, leg. Alphonse Cabra (RMCA:
65a, 68); 1 ♀, 5 ♂♂, 8 ?? [14 AL], Boende, between 1 Jan. 1967 and 31 Dec. 1967, between 1 Jan. 1966
and 31 Jan. 1966, leg. P. Lootens (RMCA: 34078, 34082, 34095, 34100, 34109, 34114, 34120, 34125,
34128, 34146, 34157 to 34159, 38731); 1 ? [AL], Bokoro, prior to 19 Nov. 1952, leg. E. Jans (RMCA:
21522); 9 ?? [9 AL], Bokuma, 7 Feb. 1952, prior to 6 May 1954, between 1 Jun. 1953 and 30 Jun. 1953,
between 1 Feb. 1954 and 28 Feb. 1954, between 1 Jan. 1954 and 31 Jan. 1954, leg. P. Lootens (RMCA:
21411, 22136, 22137, 22139, 22138, 22037, 22041, 22044, 22140); 6 ♀♀ [1 SS, 3 SO, 2 UN], Bolobo,
prior to 20 Aug. 1958, leg. N’Kele, prior to 27 Sep. 1951, leg. Ngwe (RMCA: 27279 to 27282, 20537,
20538); 1 ? [AL], Boma, 6 Apr. 1913, leg. Styczynski (RMCA: 2102); 1 ? [AL], Bomane [= Bomaneh],
23 May 2010, leg. Guy-Crispin Gembu Tungaluna (RMCA: a1.097-M-3219); 1 ? [AL], Congo da
Lemba, between 1 Jan. 1912 and 31 Dec. 1912, leg. Mayné (RMCA: 3434); 2 ♂♂ [2 UN], Djabir,
10 May 2011, 13 May 2011, leg. Prescott Musaba and Jonathan Kosele (UNIKIS: DJB173, DJB174);
1 ♂ [UN], Elisabethville [= Lubumbashi], 25 Apr. 1968, leg. unknown collector (RBINS: 18514); 1 ♂,
1 ? [1 SS, 1 SA], Faradje, 25 Mar. 1911, 9 Mar. 1912, leg. Herbert Lang, James Paul Chapin and The
American Museum Congo Expedition (AMNH: 49000, 49425); 1 ? [AL], Ganda-Sundi, 31 Mar. 1964,
leg. Alexandre Fain (RMCA: 33277); 1 ♀, 1 ♂, 1 ? [3 SO], Gangala [= Gangala-na-Bodio],
23 Mar. 1957, 24 Mar. 1957, leg. Heinz Stephan (RMCA: 27464 to 27466); 1 ♀ [SS], Gangala-naBodio, 26 Nov. 1947, leg. unknown collector (RBINS: 13331); 1 ad+foetus ♀ [AL], Ibembo, between
1 Oct. 1952 and 31 Oct. 1952, leg. Jozef Hutsebaut (RMCA: 21627); 1 ♀, 1 ♂ [1 AL, 1 UN], Île Mbiye
[= Mbiye Island], 15 Apr. 2012, leg. Prescott Musaba, Tembele and Marc Boketshu, between 31 May
2012 and 27 Jul. 2012, leg. Prescott Musaba (UNIKIS: CHIM53, RMCA: a1.097-M-3220); 1 ♀, 1 ♂
[2 SK], Irangi, 31 Jan. 1984, leg. Heinz Stephan (SMF: 65032, 65033); 1 ? [AL], Iringi, prior to
31 Aug. 1899, leg. A. or G. Lindeman (RMCA: 406); 1 ♀ [UN], Isai River, 19 Dec. 1974, leg. Michael
D. Gallagher (HZM: 7.7870); 1 ♀, 3 ♂♂ [4 SS], Kabengere, 22 Dec. 1926, 24 Dec. 1926, leg. John
Todd Zimmer (FMNH: 29425 to 29428); 2 ♀♀, 4 ♂♂ [2 SA, 2 AL, 2 UN], Kakonde, 20 Oct. 1956, leg.
Félix / Michel Anciaux de Faveaux (RMCA: 26238, 26241 to 26243, SMF: 16879, 16880); 1 ♀, 1 ?
[1 SS, 1 SO], Kalanda, 1 Aug. 1934, 8 Aug. 1934, leg. B.L. Bennett (MCZ: 31981, 31980); 1 ♂ [SS],
Kamituga, 31 Jan. 1964, leg. Alexandre Prigogine (RMCA: 32398); 1 ? [AL], Katanga, prior to
31 Aug. 1900, leg. Charles Lemaire (RMCA: 425a); 1 ♂ [AL], Katapena, 27 Dec. 1926, leg. John Todd
Zimmer (FMNH: 30699); 2 ?? [2 SS], Kawa (Forest), 19 Jul. 1952, 24 Jul. 1952, leg. Alexandre Fain
(RMCA: 81.012-M-0072, 81.012-M-0071); 2 ♀♀, 1 ♂ [3 UN], Kipangaribwe River, 1 Oct. 1947, leg.
unknown collector (RBINS: 10807.1 to 10807.3); 1 ? [SA], Kiseny [= Kasenye], 16 Sep. 1927, leg.
220
VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
Douce (RMCA: 9905); 1 ♂ [AL], Kiswishi [River], 11 Oct. 1955, leg. Félix / Michel Anciaux de
Faveaux (RMCA: 26216); 1 ad ♀, 1 juv ♀, 1 ♀, 1 ♂ [4 SS], Kivulo, 24 Aug. 1947, leg. unknown
collector (RBINS: 10723 to 10726); 1 ♀, 4 ♂♂ [5 SS], Kunungu, prior to 4 Jul. 1938, prior to
19 Nov. 1937, prior to 1 Aug. 1939, leg. N’Kele (RMCA: 15512 to 15514, 14167, 14660); 1 ? [AL],
Lake Moero [= Moero, Lake], prior to 23 Dec. 1947, leg. unknown collector (RMCA: 18173); 1 sad
♂, 2 ♀♀, 1 ♂ [1 SS, 3 SA], Léopoldville [= Kinshasa], unknown date, 8 Nov. 1893, leg. J. Camp,
8 Jul. 1909, leg. Herbert Lang, James Paul Chapin and The American Museum Congo Expedition
(USNM: 21666, 21667, AMNH: 48936, USNM: 21665); 1 ? [SS], Leverville [= Lusanga], between
1 Feb. 1929 and 28 Feb. 1929, leg. Tinant (RMCA: 9987b); 1 ad ♀, 1 ad ♂, 4 ♀♀ [4 SA, 2 UN], Luanu,
26 Jul. 1995, leg. Herwig Leirs et al. (MSB: 274911, 274914, RMCA: 97.021-M-0833 to 97.021-M0836); 2 ♂♂ [2 SS], Lukafu, 20 Dec. 1930, leg. Gaston-François de Witte and de Crawhez (RMCA:
11131, 11132); 1 ♂, 1 ? [1 SA, 1 AL], Mabali, 27 Jun. 1956, between 1 Jun. 1956 and 30 Jun. 1956, leg.
Alvin Novick (MCZ: 48261, 48262); 4 ad ♀♀, 1 ad ♂ [3 SA, 2 UN], Mbalaka (Plantation), 19 Jul. 1995,
20 Jul. 1995, 21 Jul. 1995, leg. Herwig Leirs et al. (MSB: 274917, RMCA: 97.021-M-0826, 97.021-M0827, MSB: 274918, RMCA: 97.021-M-0828); 1 juv ♂, 2 ♀♀ [1 SS, 2 UN], Medje, 27 Jun. 1910, leg.
Herbert Lang, James Paul Chapin and The American Museum Congo Expedition (AMNH: 48997,
48999, RMCA: 12393); 1 ? [SN], Moba, unknown date, leg. unknown collector (BMNH: 1954.854);
1 ♀ [AL], Mosenge, 15 Aug. 1986, leg. WHO and Marc Colyn (RMCA: 88.011-M-0017); 11 ?? [6 SS,
5 AL], Mount Wago [= Wago, Mount], unknown date, 30 Aug. 1953, prior to 28 Mar. 1952, between
1 Jan. 1951 and 31 Dec. 1951, leg. Alexandre Fain, prior to 31 Dec. 1990, leg. unknown collector
(BMNH: 1953.486, MHNG: 921.033, RMCA: 21455 to 21459, 20788 to 20790, RBINS: 21657); 1 ♀
[SS], Muhulu, 16 Jan. 1957, leg. Heinz Stephan (RMCA: 27467); 1 ? [SS], Ndwa, between 1 Oct. 1950
and 30 Nov. 1950, leg. Ngwe (RMCA: 20254); 4 ?? [4 SS], Né, Mount, unknown date, 29 Aug. 1952,
30 Aug. 1952, leg. Alexandre Fain (BMNH: 1953.498, RMCA: 21513, 21511, 21512); 5 ♀♀, 7 ♂♂
[12 UN], Obenge, 6 Feb. 2013, 7 Feb. 2013, 8 Feb. 2013, leg. Prescott Musaba and André Malekani
(UNIKIS: TLL367 to TLL371, TLL391, TLL395 to TLL400); 1 ? [AL], Pweto, unknown date, leg.
Gerard (RMCA: 5441a); 1 ? [UN], Rodahira River, unknown date, leg. unknown collector
(INPBC: W875.1); 2 ♀♀ [2 UN], Rubi-Tele, 24 Aug. 2013, leg. Guy-Crispin Gembu Tungaluna,
Prescott Musaba and André Malekani (UNIKIS: RBTL557, RBTL558); 1 ? [SS], Saliboko, 5 Aug. 1952,
leg. Alexandre Fain (RMCA: 21438); 1 ? [UN], Sombe, unknown date, leg. Lademann (ZMB: T10164);
2 ♂♂ [1 SS, 1 AL], Tandala, 16 Jul. 1979, leg. unknown collector, 16 Jul. 1979, leg. C. Brian Robbins
(MHNG: 1870.089, USNM: 537721); 1 ? [AL], Tanganyika Province, prior to 30 Jan. 1900, leg.
Célestin Hecq (RMCA: 420); 1 ? [AL], Tshakala, between 1 Jan. 1982 and 31 Dec. 1982, leg. Marc
Colyn (RMCA: 82.019-M-0008); 31 ?? [29 AL, 2 UN], Wafanya [= Wafania], unknown date,
10 Aug. 1991, 7 Oct. 1991, 11 Oct. 1991, 12 Oct. 1991, 16 Oct. 1991, 5 Mar. 1992, 20 Apr. 1992, 18 May
1992, 26 May 1992, 29 May 1992, between 1 Dec. 1991 and 31 Dec. 1991, between 1 Apr. 1991 and
30 Apr. 1991, between 1 Mar. 1992 and 31 Mar. 1992, leg. P. Lootens (RMCA: 92.079-M-0140 to
92.079-M-0148, 91.076-M-0055, 91.076-M-0107, 91.076-M-0099, 91.076-M-0111, 91.076-M-0083,
91.076-M-0113, 92.079-M-0027, 92.079-M-0029, 92.079-M-0028, 92.079-M-0032, 92.079-M-0030,
92.079-M-0031, 92.079-M-0058, 91.076-M-0104, 92.079-M-0026, 92.079-M-0059, 91.076-M-0028,
92.079-M-0057, 92.079-M-0060 to 92.079-M-0062, 92.079-M-0116); 1 juv ?, 13 ?? [13 SS, 1 SO],
Wago, Mount, 30 Aug. 1952, between 1 Jul. 1951 and 31 Jul. 1951, between 1 Jan. 1951 and
31 Dec. 1951, leg. Alexandre Fain (RMCA: 21504 to 21510, 81.012-M-0074, 81.012-M-0075, 23285 to
23287, 81.012-M-0073, 23290); 2 ♀♀, 4 ♂♂ [2 SS, 4 AL], Yalosemba, 17 Jun. 1979, between
1 Jun. 1979 and 30 Jun. 1979, leg. C. Brian Robbins (USNM: 463485 to 463487, 537719, 537720,
MHNG: 1870.090); 1 ♂ [UN], Yoko, 5 Sep. 2012, leg. Prescott Musaba, Tembele and Marc Boketshu
(UNIKIS: YK017CH); 1 ? [AL], Zambi, prior to 31 Jul. 1899, leg. Lepez (RMCA: 284).
RWANDA: 3 ad ♀♀, 1 ad ♂, 1 ? [5 SA], Butare, 21 Dec. 1984, 22 Dec. 1984, 23 Dec. 1984, 24 Dec. 1984,
leg. Benny Baeten and Luc Janssens (RMCA: 85.006-M-0450 to 85.006-M-0454); 1 ♂, 1 ? [2 SA],
221
European Journal of Taxonomy 382: 1–327 (2017)
Gisenyi, 21 Oct. 1980, leg. Danny Meirte (RMCA: 80.055-M-0167, 80.055-M-0168); 1 ♂, 1 ? [1 SA,
1 AL], Kibuye, 7 Oct. 1980, leg. Danny Meirte (RMCA: 80.055-M-0165, 80.055-M-0166); 15 ??
[15 UN], Mihindi, Lake, unknown date, leg. unknown collector (INPBC: W3052.01 to W3052.15);
1 ♂ [UN], Rubona, 1 Jan. 1966, leg. Félix / Michel Anciaux de Faveaux (RBINS: 15492).
Myotis tricolor (Temminck, 1832)
DEMOCRATIC REPUBLIC OF THE CONGO: 1 ♂ [SA], Dethiox Caves [= Kambove], 4 Jun. 1960,
leg. Félix / Michel Anciaux de Faveaux (SMF: 18919); 1 ? [UN], Djomba [= Djemba], unknown
date, leg. unknown collector (INPBC: V1846.9); 2 ♂♂ [2 AL], Jomba, 5 Sep. 1976, leg. Frieder
Meier (MHNG: 1873.072, 1873.073); 1 ♀ [SA], Kakontwe Cave, 22 Sep. 1957, leg. Félix / Michel
Anciaux de Faveaux (MHNG: 1046.024); 1 ? [UN], Kamikoni, unknown date, leg. unknown collector
(INPBC: V1694.1); 1 ♀ [SA], Katanga (former) Province, 18 Jul. 1962, leg. Félix / Michel Anciaux
de Faveaux (SMF: 21299); 1 ♀ [AL], Kyamakonde Cave, 18 Jul. 1962, leg. Félix / Michel Anciaux de
Faveaux (MHNG: 1046.097); 1 ♂ [AL], Mulonga Cave, 30 Jul. 1957, leg. Félix / Michel Anciaux de
Faveaux (RMCA: 26792); 1 ? [SO], Thysville, Cave [= Banjan Caves], unknown date, leg. Schwetz
(RMCA: 16450b).
RWANDA: 1 ad ♀, 5 ad ♂♂ [2 SS, 4 SA], Mutura, 18 Dec. 1984, leg. Benny Baeten and Luc Janssens,
23 Oct. 1981, 25 Oct. 1981, 27 Oct. 1981, leg. Frits De Vree, Benny Baeten and Victor Van Cakenberghe
(RMCA: 85.006-M-0449, 82.006-M-0920 to 82.006-M-0924).
Myotis welwitschii (Gray, 1866)
BURUNDI: 1 ? [SS], Rwegura, between 1 Jun. 1969 and 31 Jul. 1969, leg. unknown collector (MHNG:
1607.095).
DEMOCRATIC REPUBLIC OF THE CONGO: 1 ? [SO], Dilolo Station [= Dilolo], between 1 Mar. 1934
and 31 Mar. 1934, leg. Richard (RMCA: 12661); 1 ♀, 1 ? [1 UN, 1 SB], Elisabethville [= Lubumbashi],
18 Oct. 1966, leg. unknown collector, prior to 11 Feb. 1956, leg. Seydel (RBINS: 15428, RMCA:
23462); 1 ♀ [UN], Kasapa, between 1 Jul. 1967 and 31 Jul. 1967, leg. unknown collector (RBINS:
18513); 1 ♀ [SO], Lusinga, 17 Jul. 1947, leg. unknown collector (RBINS: 10848); 1 ? [UN], Mutsora,
unknown date, leg. unknown collector (INPBC: W916); 3 ?? [3 UN], Ondo, Lake, unknown date,
leg. unknown collector (INPBC: V1240.2, V1819.3.1, V1819.3.2); 1 ♂ [SA], Pande, unknown date,
leg. Jean Romieux (MHNG: 1891.002); 1 ♂ [SS], Rutshuru, 10 Mar. 1938, leg. unknown collector
(RBINS: 4789).
RWANDA: 1 ♂ [SA], Butare, 28 Sep. 1980, leg. Danny Meirte (RMCA: 80.055-M-0169).
Scotophilinae Gray, 1866
Scotophilus Leach, 1821
BURUNDI: 7 ♂♂ [2 SS, 5 SK], Bujumbura, 6 Nov. 1967, 9 Nov. 1967, 1 Dec. 1967, 15 Dec. 1967,
23 Oct. 1968, 13 Dec. 1968, 26 Dec. 1968, leg. P.L. Niort (MHNG: 1607.045, 1607.049, 1607.046,
1607.047, 1607.050, 1607.042, 1607.043); 2 ♂♂ [2 SO], Gitega, between 1 Nov. 1968 and 30 Nov. 1968,
leg. P.L. Niort (MHNG: 1607.053, 1607.054); 1 ♀ [AN], Kanyosha, 10 Oct. 1969, leg. P.L. Niort
(MHNG: 1607.084).
DEMOCRATIC REPUBLIC OF THE CONGO: 1 ad ♂ [AL], Kivu [= Kivu region], unknown date,
leg. Guy Babault (MNHN: ZM-MO-1985-1067).
222
VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
Scotophilus dinganii (A. Smith, 1833)
BURUNDI: 2 ad ♀♀, 1 ad? ♂, 5 ♀♀, 8 ♂♂ [3 SA, 8 AL, 1 SK, 4 SB], Bujumbura, 29 Nov. 1967,
12 Nov. 1968, 20 May 1969, 5 Jan. 1970, 9 Jan. 1970, 16 Nov. 1972, leg. P.L. Niort, 21 Apr. 1982, leg.
Danny Meirte, between 1 May 1977 and 31 May 1977, leg. Jacques Verschuren, between 1 Jul. 1968 and
31 Jul. 1968, leg. Lewall (MHNG: 1607.048, 1608.012, 1607.055, RMCA: 82.016-M-0002, 82.016M-0005 to 82.016-M-0008, 82.016-M-0072, 82.016-M-0124, MHNG: 1607.044, 1607.093, 1607.094,
RBINS: 19915.1, 19915.2, RMCA: 35156); 4 ♀♀ [4 SB], Kanyosha, 10 Oct. 1969, 21 Oct. 1969, leg.
P.L. Niort (MHNG: 1607.085 to 1607.088); 1 ♀ [SB], Muyebe, between 1 Jan. 1900 and 23 Dec. 1969,
leg. P.L. Niort (MHNG: 1607.092); 5 ♂♂ [5 AL], Mweya, 19 Dec. 1972, leg. P.L. Niort (MHNG:
1608.013 to 1608.017); 1 ♂ [AL], Nyanza [= Nyanza-Lac], between 29 Jan. 1952 and 1 Feb. 1952,
leg. Laurent (RMCA: 90.008-M-0001); 3 ♂♂ [2 SA, 1 SK], Resha, 5 Apr. 1979, between 4 Apr. 1979
and 5 Apr. 1979, leg. Heinz Stephan (SMF: 57567, 57569, 57568); 3 ♂♂ [3 SB], Rwintare, between
1 Feb. 1970 and 28 Feb. 1970, leg. P.L. Niort (MHNG: 1608.008 to 1608.010).
DEMOCRATIC REPUBLIC OF THE CONGO: 2 ad ♂♂ [2 SS], Albert, Lake, 23 Oct. 1906, leg.
Alexander Douglas Mitchell Carruthers (BMNH: 1907.6.14.11, 1907.6.14.12); 1 ♂ [AL], Albertville
[= Kalemie], 12 Nov. 1946, leg. La Mission Hydrobiol. Lac Tanganiyka (RBINS: 9069); 2 ad ♂♂,
1 ad ?, 1 juv ♂ [2 SA, 1 AL, 1 SK], Banana, between 1 Jan. 1884 and 31 Dec. 1884, leg. L. Anema,
between 1 Jan. 1889 and 31 Dec. 1889, leg. P. Hesse, between 1 May 1952 and 31 May 1952, between
1 Jan. 1952 and 31 Jan. 1952, leg. Is. Mesmaekers (ZMA: MAM.09008, SMF: 2515, RMCA: 20748,
21374); 1 ad ♂, 1 juv ♀, 2 juv ♂♂ [4 AL], Beno, between 1 Jan. 1949 and 31 Dec. 1949, leg. P. Bayet
(RMCA: 19831 to 19833, 19848); 1 juv ♂ [AL], Boende, between 1 Jan. 1931 and 31 Dec. 1931, leg.
R.P. Wynants (RMCA: 18141); 1 ad ♀, 1 ad ♂ [2 SA], Boma, between 1 Feb. 1955 and 28 Feb. 1955,
leg. Is. Mesmaekers (RMCA: 26321, 26322); 7 ad ♂♂, 1 juv ♀, 1 juv ♂, 1 ? [7 SA, 2 AL, 1 SK],
Elisabethville [= Lubumbashi], unknown date, leg. J. Bequaert, 6 Apr. 1956, 22 Apr. 1956, leg. Félix /
Michel Anciaux de Faveaux, 5 Aug. 1952, leg. A. Smitz, between 1 Apr. 1926 and 30 Apr. 1926, between
1 Jan. 1931 and 31 Dec. 1931, leg. Henri Schouteden, prior to 10 Aug. 1933, between 1 Jan. 1934
and 31 Dec. 1934, leg. Quarré (RMCA: 16255, 23898, RBINS: 14454, RMCA: 23819, 9179, 12850,
16212, 16233, 12149, 12150); 1 ad ♂ [SA], Funda Biabo, between 1 Jan. 1914 and 31 Dec. 1914,
leg. de Baillet Latour (RMCA: 16232); 1 ad ♀, 1 ♀, 1 ♂ [1 SA, 1 SK, 1 UN], Gangala-na-Bodio,
10 Apr. 1948, leg. H. de Saeger et al., 9 Apr. 1948, leg. unknown collector (RBINS: 8126, 8125, 8127.1);
1 ad ♂ [SA], Haute Congo, 4 Jun. 1992, leg. The Rotterdamse Diergaarde (RMNH: MAM.28010);
2 juv ♂♂, 2 ♀♀ [2 AL, 2 UN], Jadotville [= Likasi], 1 Dec. 1957, 15 Nov. 1962, leg. Félix / Michel
Anciaux de Faveaux, 16 Dec. 1959, leg. unknown collector (RMCA: 27585, 33986, RBINS: 14452,
14453); 1 ad ♂ [SA], Kabalo, between 1 Mar. 1926 and 31 Mar. 1926, leg. Henri Schouteden (RMCA:
16249); 1 ♀, 1 ♂ [2 SS], Kabwe, 12 Nov. 1973, leg. Western New Mexico University (FMNH: 152836,
152837); 1 ? [UN], Kadia, between 1 Mar. 1926 and 31 Mar. 1926, leg. Henri Schouteden (RMCA:
16252); 1 ad? ♂ [SA], Kakontwe Cave, prior to 24 Sep. 1962, leg. L. Poelman (RMCA: 31217); 1 ad
♂ [SA], Kalenda [Kasai], 11 Jul. 1953, leg. A. Bouillon (RBINS: 7792); 5 ad ♀♀, 1 ad ♂ [5 SA, 1 AS],
Kamponde, between 1 Apr. 1965 and 30 Apr. 1965, leg. Antoon Emeric Marcel De Roo (RMCA: 33383
to 33388); 1 ♀, 1 ♂ [2 AL], Kananga, 21 Sep. 1963, leg. Antoon Emeric Marcel De Roo (AMNH:
212597, 212598); 1 ? [UN], Katanda, unknown date, leg. unknown collector (INPBC: W1369);
2 ♀♀ [2 SA], Kikole-Pecheur, 17 May 1980, leg. M. Mutashia (SMF: 58065, 58066); 1 ad ♀, 1 ad ♂
[1 SA, 1 UN], Kikwit, 6 Jul. 1995, leg. Herwig Leirs et al. (MSB: 274622, RMCA: 97.021-M-0840);
2 ad ♀♀, 3 ad ♂♂ [2 SA, 3 UN], Kikwit ITPK [= Kikwit], 1 Aug. 1995, leg. Herwig Leirs et al.
(MSB: 274926, 274986, 274987, RMCA: 97.021-M-0847, 97.021-M-0848); 4 ♀♀ [4 AL], Kinkole,
15 Aug. 1980, leg. unknown collector (ZMA: MAM.21149 to MAM.21152); 6 ad ♀♀, 1 ad ♂, 1 ad? ♀
[4 SA, 4 UN], Kinzambi Mission, 24 Jul. 1995, 25 Jul. 1995, 26 Jul. 1995, 28 Jul. 1995, 30 Jul. 1995,
31 Jul. 1995, leg. Herwig Leirs et al. (MSB: 274922, RMCA: 97.021-M-0843, MSB: 274923, 274924,
223
European Journal of Taxonomy 382: 1–327 (2017)
RMCA: 97.021-M-0844, 97.021-M-0845, MSB: 274925, RMCA: 97.021-M-0846); 2 ad ♀♀, 1 ad ♂,
1 juv ♀, 1 ♀, 1 ♂ [5 SA, 1 AL], Kipopo, 30 Nov. 1955, leg. Félix / Michel Anciaux de Faveaux,
30 Nov. 1955, leg. De Bont, 30 Nov. 1955, leg. Michel Anciaux de Faveaux (MHNG: 922.082, RMCA:
26323 to 26326, SMF: 16866); 1 ad ♀ [SA], Kipushi, 12 Aug. 1954, leg. De Smet (RMCA: 22458);
1 ad ? [SK], Kisantu, 15 Dec. 1920, leg. Henri Schouteden (RMCA: 6564); 1 ? [SK], Kyasala Cave
[= Kiasala Cave], 31 Mar. 1956, leg. Félix / Michel Anciaux de Faveaux (RMCA: 23851 b); 1 ad ♂
[SA], Losho [= Loso], 23 Nov. 1954, leg. Gaston-François de Witte (RMCA: 37380); 3 ad ♀♀, 1 ad ♂,
3 juv ?? [4 SS, 3 SO], Lovanium University [= Kinshasa], 10 Dec. 1962, 12 Mar. 1963, 6 Nov. 1963,
between 8 Dec. 1962 and 27 Nov. 1963, between 7 Nov. 1963 and 27 Nov. 1963, between 27 Nov. 1962
and 27 Nov. 1963, leg. De Bont and Jo Van Orshoven (RMCA: 32628, 32626, 32625, 32629, 32624,
32627, 32630); 2 ad ♀♀, 2 ad ♂♂ [4 SA], Lubumbashi, 6 Jul. 1970, leg. Peelman (RBINS: 18512.1
to 18512.4); 8 ad ♀♀, 7 ad ♂♂, 5 ad ??, 1 ad? ♀, 1 sad ♀, 3 ?? [3 SS, 14 SA, 2 AL, 4 SK, 2 SO],
Luluabourg [= Kananga], 8 Sep. 1963, 5 Oct. 1963, 16 Feb. 1965, 2 Mar. 1965, 19 Mar. 1965, 17 May
1965, 3 Jul. 1965, 30 Apr. 1966, between 3 Mar. 1965 and 17 May 1965, between 29 Aug. 1963 and
5 Oct. 1963, between 2 Mar. 1965 and 17 May 1965, between 17 May 1965 and 8 May 1966, between
17 May 1965 and 20 May 1965, between 17 May 1965 and 19 May 1965, between 14 Sep. 1963 and
14 Oct. 1964, leg. Antoon Emeric Marcel De Roo, between 1 Jan. 1931 and 31 Dec. 1931, leg. Jan
Deheegher (RMCA: 33314, 33650, 33318, 33319, 33315 to 33317, 33380, 33381, 33595, 35287, 32669,
32671, 32672, 33310 to 33312, 31566, 33313, 33651, 33652, 35288, 33382, 32670, 33649); 4 ad ♀♀
[4 SA], Luluabourg Ravine, 29 Aug. 1963, between 29 Jun. 1963 and 29 Aug. 1963, leg. Antoon
Emeric Marcel De Roo (RMCA: 32770 to 32772, 32769); 2 ad ♀♀, 2 ad ♂♂ [2 SA, 2 UN], Mbalaka
(Plantation), 21 Jul. 1995, 23 Jul. 1995, 24 Jul. 1995, leg. Herwig Leirs et al. (MSB: 274989, RMCA:
97.021-M-0841, MSB: 274988, RMCA: 97.021-M-0842); 1 ad ♂ [SA], Mbwambala, 20 Jun. 1995,
leg. Herwig Leirs et al. (RMCA: 97.021-M-0839); 1 juv ♂ [SO], Nagero, 22 Jul. 1954, leg. unknown
collector (RBINS: 14629); 1 juv ♀, 1 juv ♂ [2 AL], Panda [= Pande], 15 Nov. 1962, 18 Nov. 1962,
leg. Félix / Michel Anciaux de Faveaux (RMCA: 33987, 33988); 1 ad ?, 2 ?? [1 SK, 2 SO], Paulis
[= Isiro], 28 Apr. 1947, 1 Nov. 1947, between 1 Jan. 1947 and 31 Dec. 1947, leg. Abbeloos (RMCA:
18569, 18943, 17931); 3 ad ♀♀, 1 juv ♀, 1 imm ♂, 5 ?? [1 SA, 2 AL, 2 SK, 5 UN], Rutshuru, unknown
date, leg. Jacques Verschuren, unknown date, leg. unknown collector, 27 Mar. 1954, leg. GastonFrançois de Witte, between 24 Mar. 1932 and 24 Mar. 1937, between 1 May 1937 and 31 May 1937, leg.
J. Ghesquière (INPBC: V1362, V1420.2, V1426, V1522.2, V415, W334, W917, RMCA: 37379, 14065,
13868); 1 ad ♀ [AL], Rwindi, unknown date, leg. Jacques Verschuren (INPBC: V1623); 1 ad ♀, 2 ad
♂♂ [3 SK], St. Joseph de Luluabourg Mission [= Mikalaya], 19 Feb. 1923, between 1 Jan. 1931 and
31 Dec. 1931, leg. Richard Callewaert (RMCA: 7110 to 7112); 1 ♀, 1 ♂ [2 SS], Tandala, 10 Jul. 1979,
19 Jul. 1979, leg. C. Brian Robbins (USNM: 537728, 537729); 1 sad ♂ [AL], Tungudu, 3 Jun. 1957,
leg. Gaston-François de Witte (RMCA: 37378); 1 ad ♂, 1 ad? ♂, 1 juv ♀ [2 SA, 1 AL], Uvira, between
3 Mar. 1955 and 21 Jun. 1958, between 12 Sep. 1951 and 12 Sep. 1957, between 1 Apr. 1955 and
30 Apr. 1955, leg. Narcisse Leleup (RMCA: 27203, 22757, 27220); 1 ad ♀, 1 ad ♂ [1 SA, 1 AL],
Virunga National Park, unknown date, leg. Jacques Verschuren (INPBC: V1807, V1957); 1 ad ♀
[SS], Vitshumbi, 28 Sep. 1933, leg. Gaston-François de Witte et al. (RBINS: 4772); 2 ♂♂ [2 SS],
Yalosemba, 28 Jun. 1979, 30 Jun. 1979, leg. C. Brian Robbins (USNM: 537727, 537726).
RWANDA: 5 ?? [1 SA, 4 AL], Astrida [= Butare], 12 Oct. 1976, 13 Oct. 1976, leg. Frieder Meier
(MHNG: 1873.090 to 1873.094); 9 ad ♀♀, 4 ad ♂♂, 6 sad ♀♀, 5 sad ♂♂ [1 SA, 23 AL], Butare,
10 Dec. 1984, 11 Dec. 1984, 20 Dec. 1984, 21 Dec. 1984, 22 Dec. 1984, 23 Dec. 1984, 24 Dec. 1984,
15 Jan. 1985, leg. Benny Baeten and Luc Janssens, 3 Oct. 1981, leg. Frits De Vree, Benny Baeten
and Victor Van Cakenberghe (RMCA: 85.006-M-0097 to 85.006-M-0101, 85.006-M-0118, 85.006M-0119, 85.006-M-0102 to 85.006-M-0117, 82.006-M-0915); 1 ad ♀ [SA], Cyili Ntyano [= Cyili],
21 Apr. 1985, leg. Théoneste (RMCA: 87.047-M-0131); 1 juv ♀ [AL], Gabiro, prior to 27 Apr. 1936, leg.
Verhulst (RMCA: 13281); 1 ad ♀ [SA], Kigali, between 1 Jul. 1982 and 31 Jul. 1982, leg. Benny Baeten
224
VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
(RMCA: 87.046-M-0246); 1 ad ♀ [SA], Ruboma Ruhashya, between 1 Jan. 1985 and 31 Dec. 1985,
leg. Théoneste (RMCA: 87.047-M-0130); 2 sad ♂♂ [2 AL], Rukira, 20 Nov. 1981, 24 Nov. 1981, leg.
Frits De Vree, Benny Baeten and Victor Van Cakenberghe (RMCA: 82.006-M-0916, 82.006-M-0917);
1 juv ♀ [AL], Rwankwi, between 1 Apr. 1948 and 30 Apr. 1948, leg. J.V. Leroy (RMCA: 19068).
Scotophilus leucogaster (Cretzschmar, 1826)
BURUNDI: 2 ♂♂ [2 AL], Mweya, 29 Apr. 1971, leg. P.L. Niort (MHNG: 1896.016, 1896.017).
DEMOCRATIC REPUBLIC OF THE CONGO: 1 foetus ?, 1 ♀, 1 ♂ [2 AL, 1 SO], Faradje, 6 Mar. 1912,
7 Mar. 1912, leg. Herbert Lang, James Paul Chapin and The American Museum Congo Expedition
(AMNH: 49359, 49015, 49283).
Scotophilus nigrita (Schreber, 1774)
BURUNDI: 1 ♂ [SK], Bujumbura, 31 Jan. 1969, leg. P.L. Niort (MHNG: 1607.052).
DEMOCRATIC REPUBLIC OF THE CONGO: 4 ad ♀♀, 5 ad ♂♂, 3 ♂♂, 5 ?? [9 SS, 1 SA, 5 AL,
1 SO, 1 UN], Albertville [= Kalemie], 5 Jan. 1964, prior to 18 Aug. 1965, leg. Antoon Emeric Marcel
De Roo, prior to 12 Jul. 1955, leg. service hygiene publique albertville, prior to 18 Nov. 1959, leg.
H. Bomans, prior to 5 Dec. 1951, leg. unknown collector (AMNH: 206844, 212591 to 212596, RMCA:
31718, 22948, 33585, 33586, 29293 to 29297, RBINS: 14598).
Scotophilus nux Thomas, 1904
DEMOCRATIC REPUBLIC OF THE CONGO: 1 ad ? [SS], Ibembo, between 6 Jun. 1951 and
21 Dec. 1951, leg. Jozef Hutsebaut (RMCA: 20661); 1 ♂ [UN], Kongolo, 9 Nov. 1974, leg. Michael
D. Gallagher (HZM: 64.7806); 2 ad ♀♀, 2 ad ?? [2 SS, 2 UN], Medje, unknown date, 26 Mar. 1914, leg.
Cuthbert Christy, between 1 Mar. 1914 and 31 Mar. 1914, leg. Herbert Lang, James Paul Chapin and The
American Museum Congo Expedition (BMNH: 1919.5.8.13, RMCA: 4344, AMNH: 49286, 49287);
2 ad ♂♂ [2 SS], Panga, between 1 Sep. 1925 and 30 Sep. 1925, leg. Henri Schouteden (RMCA: 9125,
9126); 1 ? [UN], Poko, unknown date, leg. unknown collector (RMCA: 4346); 3 ad ♂♂ [3 SS], Teturi,
23 Mar. 1947, 24 Mar. 1947, leg. Jean Hiernaux (RBINS: 7041 to 7043); 3 ♀♀, 3 ♂♂ [6 SS], Yalosemba,
17 Jun. 1979, 26 Jun. 1979, 28 Jun. 1979, leg. C. Brian Robbins (USNM: 537730 to 537735).
Vespertilioninae Gray, 1821
Glauconycteris Dobson, 1875
DEMOCRATIC REPUBLIC OF THE CONGO: 1 ♂ [AL], Île Mbiye [= Mbiye Island],
23 Jun. 2014, leg. Prescott Musaba, Moïse Bipoo, Richard Tamaru, Pauline Isude and Joseph Kussa
(UNIKIS: CHIM020/14); 1 ♀ [SS], Luluabourg [= Kananga], unknown date, leg. Richard Callewaert
(BMNH: 1926.7.6.95).
Glauconycteris alboguttata J.A. Allen, 1917
DEMOCRATIC REPUBLIC OF THE CONGO: 1 ♀ [SO], Lukolela, 14 Mar. 1933, leg. Lecocq (RMCA:
12175); 1 ad ♀ [UN], Medje, 14 Mar. 1914, leg. Herbert Lang, James Paul Chapin and The American
Museum Congo Expedition (AMNH: 49317 [holotype Glauconycteris alboguttatus J.A. Allen, 1917]);
1 ? [UN], Yalosemba, unknown date, leg. unknown collector (CM: 86658); 1 ♀ [UN], Yangambi,
4 Mar. 1947, leg. unknown collector (RBINS: 14516).
225
European Journal of Taxonomy 382: 1–327 (2017)
Glauconycteris argentata (Dobson, 1875)
BURUNDI: 2 ♀♀ [2 SS], Rumonge, between 1 Aug. 1969 and 31 Aug. 1969, leg. P.L. Niort (MHNG:
1607.090, 1607.091).
DEMOCRATIC REPUBLIC OF THE CONGO: 4 ♀♀, 1 ♂, 2 ?? [7 AL], unknown locality, between
1 Apr. 1800 and 31 May 1899, leg. P. Hesse, prior to 28 Jun. 1960, leg. Uwe Rahm (SMF: 11850
[paralectotype Chalinolobus congicus Noack, 1889], 11851 [paralectotype Chalinolobus congicus
Noack, 1889], 11852 [paralectotype Chalinolobus congicus Noack, 1889], 11853 [paralectotype
Chalinolobus congicus Noack, 1889], 11854 [paralectotype Chalinolobus congicus Noack, 1889],
11855 [paralectotype Chalinolobus congicus Noack, 1889], RMCA: 28921); 1 ad ♂, 3 ?? [3 AL,
1 UN], Banana, unknown date, leg. unknown collector, 13 Mar. 1952, between 1 May 1952 and 31 May
1952, leg. Is. Mesmaekers (BMNH: 1954.863, RMCA: 20751, 20752, 21376); 1 ♀ [AL], Boende,
between 1 Jan. 1966 and 31 Jan. 1966, leg. P. Lootens (RMCA: 34160); 1 ? [AL], Boga, prior to
13 May 1913, leg. André Pilette (RMCA: 1862); 5 ?? [4 AL, 1 UN], Bokuma, 7 Feb. 1952, 7 Mar. 1952,
between 1 Jan. 1954 and 31 Jan. 1954, leg. P. Lootens (RMCA: 21412, 21414, 21415, 21413, 22135);
2 ♂♂ [2 SO], Bokungu, 25 Dec. 1948, leg. L. Dupuis (RMCA: 18713, 18714); 1 juv ?, 1 ♀, 5 ♂♂ [5 SS,
2 SO], Boma, 23 Nov. 1920, 28 Nov. 1920, leg. Henri Schouteden (RMCA: 6571 to 6576, 7250); 1 ?
[SS], Bushushu, 26 Jun. 1959, leg. Uwe Rahm (RMCA: 33532); 1 ♀, 2 ♂♂, 1 ? [1 SS, 3 SO], Butembo,
25 Apr. 1954, prior to 26 Jan. 1955, leg. P. Dyleff (RMCA: 22723, 22816 to 22818); 5 ?? [5 AL],
Djeka, 29 May 1955, prior to 15 Jun. 1956, leg. Roiseux (RMCA: 23635 to 23639); 21 ?? [21 UN],
Fataki, between 1 Jan. 1948 and 6 Apr. 1964, leg. unknown collector (RBINS: 18526.01 to 18526.17,
18527.1 to 18527.4); 1 ad ♂, 5 ?? [3 AL, 3 UN], Ikela, unknown date, leg. unknown collector, prior to
4 Jan. 1958, prior to 3 Nov. 1956, leg. P. Lootens (BMNH: 1959.510, RMCA: 25827, 25832 to 25834,
27022); 1 ? [SO], Inkongo [= Inkongo-Kakese], 10 Nov. 1932, leg. Wilson (RMCA: 12060); 3 ??
[3 SS], Jiba, 1 Aug. 1952, leg. Alexandre Fain (RMCA: 21444, 21448, 81.012-M-0041); 1 ? [SS], Jiba
Risasi, unknown date, leg. Alexandre Fain (BMNH: 1953.491); 1 ? [SS], Jiba Trisasi [= Jiba Risasi],
1 Aug. 1952, leg. Alexandre Fain (RMCA: 21441); 2 ?? [2 AL], Kamituga, 13 Jul. 1966, leg. Alexandre
Prigogine (RMCA: 32862a, 32862b); 1 ad ♂ [SA], Kivu region, 10 Jul. 1964, leg. P. Kunkel (SMF:
31836); 4 ?? [3 SS, 1 SO], Korovi, Mount, 25 Oct. 1951, 25 Nov. 1951, leg. Alexandre Fain (RMCA:
23274 to 23277); 1 ? [AL], Lemera, between 1 Nov. 1956 and 30 Nov. 1956, leg. Laurent (RMCA:
35690); 1 ♂ [AL], Lodja, 20 Jul. 1986, leg. WHO and Marc Colyn (RMCA: 88.011-M-0015); 1 ♀ [SS],
Luluabourg [= Kananga], 26 Sep. 1922, leg. Richard Callewaert (RMCA: 7570); 1 ♂ [SS], Lwiro
[= Luiro], 16 Jun. 1966, leg. A. Elbl (USNM: 535398); 3 ♀♀ [3 SS], Mboga [= Boga], unknown date,
leg. Howard de Walden (BMNH: 1930.11.11.178 to 1930.11.11.180); 3 ?? [3 SS], Misu, 3 Aug. 1952,
leg. Alexandre Fain (RMCA: 21440, 21443, 21447); 1 juv ?, 6 ?? [3 SS, 3 AL, 1 SK], Mount Wago
[= Wago, Mount], unknown date, between 1 Jul. 1951 and 31 Jul. 1951, between 1 Jan. 1951 and
31 Dec. 1951, leg. Alexandre Fain, 29 Jul. 1952, leg. unknown collector (BMNH: 1953.487, 1953.488,
RBINS: 21669, RMCA: 20802, 21467, 21476, 21477); 1 ♀, 1 ♂, 1 ? [2 SS, 1 AL], Mwanda, 12 Mar. 1959,
leg. Uwe Rahm (RMCA: 28027, 28910, 28911); 1 ♂, 6 ?? [7 SK], Netonna, between 1 Apr. 1800
and 31 May 1899, leg. P. Hesse (SMF: 2516 [lectotype Chalinolobus congicus Noack, 1889], 2517
[paralectotype Chalinolobus congicus Noack, 1889], 2518 [paralectotype Chalinolobus congicus
Noack, 1889], 2519 [paralectotype Chalinolobus congicus Noack, 1889], 2520 [paralectotype
Chalinolobus congicus Noack, 1889], 2521 [paralectotype Chalinolobus congicus Noack, 1889],
2522 [paralectotype Chalinolobus congicus Noack, 1889]); 1 ? [UN], Rutshuru, unknown date, leg.
unknown collector (INPBC: V1599); 1 ♀ [SO], St. Joseph de Luluabourg Mission [= Mikalaya],
6 May 1939, leg. Jean Jacques Deheyn (RMCA: 15375); 1 ♂ [AL], Tandala, 11 Jul. 1979, leg. Frieder
Meier (MHNG: 1870.074); 1 ♂ [AL], Uvira, 5 Sep. 1956, leg. Laurent (RMCA: 35691); 1 ? [SA],
Wafanya [= Wafania], 3 Dec. 1988, leg. P. Lootens (RMCA: 89.020-M-0048); 1 ? [SS], Wago, Mount,
prior to 29 Nov. 1955, leg. Alexandre Fain (RMCA: 23273); 4 ?? [3 SS, 1 SO], Wasa [= Wasa Wago
226
VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
Forest], 1 Aug. 1952, 3 Aug. 1952, prior to 16 Oct. 1952, leg. Alexandre Fain (RMCA: 21446, 21445,
21449, 21442); 3 ?? [3 SS], Wasa Wago Forest, unknown date, 1 Aug. 1952, leg. Alexandre Fain
(BMNH: 1953.489, 1953.490, RMCA: 21439).
RWANDA: 1 ? [SA], Astrida [= Butare], between 1 Jan. 1948 and 31 Dec. 1948, leg. Alexandre
Fain (RMCA: 18993); 1 ad ♀, 1 ad ♂ [2 SA], Kidaho, 10 Jun. 1982, 15 Jun. 1982, leg. Benny Baeten
(RMCA: 87.046-M-0347, 87.046-M-0348); 1 ? [UN], Kissenji [= Gisenyi], unknown date, leg. Köhl
(ZMB: 37016).
Glauconycteris beatrix Thomas, 1901
DEMOCRATIC REPUBLIC OF THE CONGO: 1 ♂ [AL], Aketi, 22 Jun. 2015, leg. Vlir/Aketi
(UNIKIS: AKETI604); 1 ? [AL], Ibembo, 1 Jul. 1950, leg. Jozef Hutsebaut (RMCA: 20065);
1 ♀, 5 ♂♂, 1 ? [1 SS, 2 SA, 1 AL, 2 SO, 1 UN], Irangi, unknown date, between 16 Sep. 1959 and
19 Sep. 1959, leg. Uwe Rahm, unknown date, leg. unknown collector, 3 Oct. 1955, leg. J.J. Laarman,
between 11 Mar. 1990 and 21 Mar. 1990, leg. Klaus-Gerhard Heller and Marianne Volleth (BMNH:
1962.1847, SMF: 7940, RMNH: MAM.16362, MAM.16363, SMF: 79439, 79440, BMNH: 1963.1172);
1 ad ♂, 1 ♂ [1 SA, 1 UN], Kimbinga Market, 20 Jul. 1995, leg. Herwig Leirs et al. (MSB: 274578,
RMCA: 97.021-M-0810); 1 ? [SS], Paulis [= Isiro], 1 Jan. 1950, leg. Abbeloos (RMCA: 19820); 6 ??
[6 AL], Wafanya [= Wafania], 20 Nov. 1988, 17 Oct. 1991, between 1 Jan. 1991 and 18 Oct. 1991,
leg. P. Lootens (RMCA: 91.076-M-0076, 89.020-M-0044, 91.076-M-0052, 91.076-M-0057, 91.076-M0058, 91.076-M-0074).
Glauconycteris curryae Eger & Schlitter, 2001
DEMOCRATIC REPUBLIC OF THE CONGO: 1 ? [UN], Yalosemba, unknown date, leg. unknown
collector (CM: 86663).
Glauconycteris humeralis J.A. Allen, 1917
DEMOCRATIC REPUBLIC OF THE CONGO: 1 ♀ [UN], unknown locality, unknown date, leg.
unknown collector (BMNH: 1990.0233); 1 ♀ [SS], Avakubi, 21 Feb. 1914, leg. Herbert Lang, James
Paul Chapin and The American Museum Congo Expedition (AMNH: 49014); 1 ♂ [UN], Avakubi,
near, 9 Dec. 1974, leg. Michael D. Gallagher (HZM: 1.8023); 2 ♀♀, 1 ♂ [1 AL, 1 UN, 1 SB], Epulu,
2 km W, 27 Jun. 1991, 28 Jun. 1991, leg. Julian C. Kerbis Peterhans (FMNH: 149417 to 149419); 1 ♀,
2 ♂♂ [3 SK], Irangi, 17 Jan. 1984, 20 Jan. 1984, leg. Heinz Stephan (SMF: 64984 to 64986); 1 ad ♀,
1 ♀, 1 ♂, 1 ? [4 UN], Medje, 7 Sep. 1910, leg. Herbert Lang, James Paul Chapin and The American
Museum Congo Expedition (AMNH: 49012, 49013 [holotype Glauconycteris humeralis J.A. Allen,
1917], 49312, 49315); 1 ♂ [SO], St. Jos. (Mission) [= Mikalaya], 30 Oct. 1924, leg. Richard Callewaert
(AMNH: 86152); 1 ? [UN], Yalosemba, unknown date, leg. unknown collector (CM: 110209).
Glauconycteris poensis (Gray, 1842)
DEMOCRATIC REPUBLIC OF THE CONGO: 1 ? [AL], Bokuma, prior to 5 Mar. 1953, leg. P. Lootens
(RMCA: 21684); 1 ? [AL], Busu Mokalu [= Busu-Mokala], between 5 Nov. 1938 and 10 Nov. 1938,
leg. Baudart (RMCA: 18092); 1 ♂ [SO], Faradje, 15 Mar. 1911, leg. Herbert Lang, James Paul Chapin
and The American Museum Congo Expedition (AMNH: 49066); 1 ad ♂ [SA], Irangi, 21 Jan. 1984, leg.
Heinz Stephan (SMF: 64987).
227
European Journal of Taxonomy 382: 1–327 (2017)
Glauconycteris superba Hayman, 1939
DEMOCRATIC REPUBLIC OF THE CONGO: 1 ad ♂, 1 ♂ [1 SA, 1 AL], Île Mbiye [= Mbiye
Island], 11 Feb. 2012, leg. Guy-Crispin Gembu Tungaluna, 24 Dec. 2015, leg. Prescott Musaba,
Moïse Bipoo, Charle Andabhati, Benjamin Kirongozi and Kambale Karupao (RMCA: a1.097-M-3153,
UNIKIS: ABCHIM01); 1 ♂ [SA], Pawa, prior to 8 Aug. 1938, leg. Albert Dubois (RMCA: 14765
[holotype Glauconycteris superba Hayman, 1939]).
Glauconycteris variegata (Tomes, 1861)
DEMOCRATIC REPUBLIC OF THE CONGO: 1 ♀, 2 ♂♂ [1 SA, 2 SO], Aba, 13 Dec. 1911, leg.
Herbert Lang, James Paul Chapin and The American Museum Congo Expedition (AMNH: 49069,
49070, 49347); 1 ♀, 3 ♂♂, 7 ?? [8 SS, 1 SA, 2 AL], Boma, unknown date, prior to 17 Jul. 1924, leg.
J. Rodhain, 23 Nov. 1920, leg. Henri Schouteden, between 1 Aug. 1928 and 31 Aug. 1928, leg. Gerard
(RMCA: 7636 to 7639, 6566 to 6569, 16229, 16230, 7635); 1 ? [AL], Boteka, 20 Oct. 1985, leg.
P. Lootens (RMCA: 85.052-M-0003); 1 ? [SO], Elisabethville [= Lubumbashi], between 1 Aug. 1934
and 31 Aug. 1934, leg. Richard (RMCA: 12662); 11 ♀♀, 6 ♂♂, 1 ? [11 SS, 4 SO, 3 UN], Faradje,
14 Mar. 1911, 15 Mar. 1911, leg. Herbert Lang, James Paul Chapin and The American Museum Congo
Expedition, 14 Mar. 1911, leg. Herbert Lang, 14 Mar. 1911, leg. unknown collector (AMNH: 49058 to
49065, FMNH: 43872, MCZ: 17382, 17383, AMNH: 49067, 49068, 49195, 52116 to 52118, RMCA:
12381); 3 ?? [2 AL, 1 UN], Kashwa, unknown date, leg. unknown collector, 24 Aug. 1955, leg. GastonFrançois de Witte (INPBC: W1519, RMCA: 37167, 37168); 2 ?? [2 UN], Katanda, unknown date, leg.
unknown collector (INPBC: W548, W549); 1 ♂ [SO], Kwamouth, 2 Jun. 1921, leg. Henri Schouteden
(RMCA: 6570); 1 ? [UN], Lake Tanganyika, NW bank, unknown date, leg. Rudolf Grauer (ZMB:
72234); 1 juv ♂ [AL], Moenda, between 1 Oct. 1920 and 31 Oct. 1920, leg. Henri Schouteden (RMCA:
16489); 1 ♀, 1 ♂, 2 ?? [4 UN], Mpala, unknown date, prior to 30 Apr. 1948, prior to 13 Oct. 1948,
leg. unknown collector (INPBC: 14517, 14518, RBINS: 14517, 14518); 1 ? [AL], Mukishi [Station],
1 Dec. 1927, leg. Becquet (RMCA: 16213); 3 ♂♂, 2 ?? [3 SS, 2 SA], Niangara, 14 Nov. 1910,
16 Nov. 1910, 6 Dec. 1910, leg. Herbert Lang, James Paul Chapin and The American Museum Congo
Expedition, 16 Nov. 1910, leg. Herbert Lang (RMCA: 12382, AMNH: 49056, FMNH: 43871, AMNH:
49313, 49341).
Hypsugo Kolenati, 1856
Hypsugo anchietae (Seabra, 1900)
DEMOCRATIC REPUBLIC OF THE CONGO: 2 ad ♂♂ [2 AL], Lubenge, 10 Mar. 1929, 13 Mar. 1929,
leg. J. Sterling Rockefeller (AMNH: 55773, 55774).
Hypsugo crassulus (Thomas, 1904)
DEMOCRATIC REPUBLIC OF THE CONGO: 1 ad ♂ [AS], Bokuma, 20 Jul. 1952, leg. P. Lootens
(RMCA: 21669); 2 ad ♀♀, 3 ad ♂♂ [2 SS, 1 AL, 2 SK], Irangi, 17 Jan. 1984, 20 Jan. 1984, leg.
Heinz Stephan, between 13 Mar. 1990 and 25 Mar. 1990, leg. Klaus-Gerhard Heller and Marianne
Volleth (SMF: 65011, 65012, 79441 to 79443); 1 ad ♀ [SA], Mulolo-Kisanga [= Lulola-Kasanga],
18 Mar. 1960, leg. Uwe Rahm (BMNH: 1963.1174).
Hypsugo cf. eisentrauti Hill, 1968
DEMOCRATIC REPUBLIC OF THE CONGO: 1 ad ♀ [SB], Kabobo Forest, 17 Feb. 2007, leg.
C. Kahindo (FMNH: 195086); 1 ad ♀, 1 ♀ [2 AL], Lukolela, 10 Oct. 1930, leg. James Paul Chapin
(AMNH: 86916, 86917).
228
VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
RWANDA: 1 ad ♂ [SS], Nyungwe Forest Reserve, 28 Mar. 1990, leg. Klaus-Gerhard Heller and
Marianne Volleth (SMF: 79444).
Hypsugo musciculus (Thomas, 1913)
DEMOCRATIC REPUBLIC OF THE CONGO: 1 ad ♀ [SA], Bata, unknown date, leg. Charles-Henri
Pobéguin (MNHN: ZM-MO-1892-223); 1 ad ♂ [AL], Bokuma, between 1 Jun. 1953 and 30 Jun. 1953,
leg. P. Lootens (RMCA: 22049).
Laephotis Thomas, 1901
Laephotis angolensis Monard, 1935
DEMOCRATIC REPUBLIC OF THE CONGO: 1 ♀ [SS], Mumene [= Mumena], 28 Dec. 1955, leg.
unknown collector (BMNH: 1957.437); 1 ♂ [SS], Musonge, 27 Dec. 1955, leg. unknown collector
(BMNH: 1957.435).
Laephotis botswanae Setzer, 1971
DEMOCRATIC REPUBLIC OF THE CONGO: 8 ♀♀, 1 ? [2 SS, 6 SA, 1 AL], Mumene [= Mumena],
27 Dec. 1955, 28 Dec. 1955, leg. unknown collector, 28 Dec. 1955, leg. The Hygiene Services of the
Congo, 28 Dec. 1955, leg. M. Lips (SMF: 16869, BMNH: 1957.436, 1957.438, RMCA: 26402 to 26406,
SMF: 16868); 2 ?? [1 AL, 1 UN], Musonge, unknown date, leg. unknown collector, 27 Dec. 1955, leg.
M. Lips (SMF: 16869a, RMCA: 26407).
Mimetillus Thomas, 1904
Mimetillus moloneyi (Thomas, 1891)
DEMOCRATIC REPUBLIC OF THE CONGO: 2 ?? [2 AL], Arebi Bondo Mabe, unknown date,
leg. Henri Schouteden (RMCA: 16264, 16265); 1 ? [SA], Avakubi, 15 Dec. 1913, leg. Herbert Lang,
James Paul Chapin and The American Museum Congo Expedition (AMNH: 49305); 3 ?? [3 AL], Buta,
unknown date, prior to 31 May 1939, prior to 23 Jun. 1949, leg. Jozef Hutsebaut (RMCA: 16279,
19081, 15326); 1 ad ♂, 1 ? [1 AL, 1 UN], Irumu, unknown date, leg. unknown collector, between
1 Nov. 1951 and 30 Nov. 1951, leg. Alexandre Fain (BMNH: 1954.862, RMCA: 21462); 1 ad ♂, 1 ♀,
1 ♂ [3 UN], Isai River, between 19 Dec. 1974 and 21 Dec. 1974, leg. Michael D. Gallagher (HZM:
2.7802 to 4.7804); 6 ?? [5 AL, 1 UN], Katanga, prior to 31 Aug. 1899, leg. Charles Lemaire (RMCA:
386a to 386f); 1 ? [AL], Koteli [= Kotili], between 1 Jan. 1925 and 31 Jan. 1925, leg. Henri Schouteden
(RMCA: 16266); 3 ♀♀, 3 ♂♂ [6 AL], Luluabourg [= Kananga], 3 Jul. 1965, leg. Antoon Emeric
Marcel De Roo (RMCA: 33587 to 33592); 2 ♂♂ [2 UN], Medje, 26 Jun. 1910, 7 Sep. 1910, leg.
Herbert Lang, James Paul Chapin and The American Museum Congo Expedition (AMNH: 48859,
49198); 1 ♂ [AL], Mosenge, 11 Aug. 1986, leg. WHO and Marc Colyn (RMCA: 88.011-M-0023);
1 juv ? [AL], Poko, between 1 Jul. 1914 and 31 Jul. 1914, leg. Cuthbert Christy (RMCA: 16514);
1 ♂ [SS], Stanleyville [= Kisangani], 25 Aug. 1909, leg. Herbert Lang, James Paul Chapin and The
American Museum Congo Expedition (AMNH: 48858); 1 ♀, 4 ♂♂ [5 UN], Université de Kisangani,
Faculté des Sciences [= Kisangani], 10 Aug. 2015, 11 Aug. 2015, leg. Prescott Musaba, Moïse Bipoo,
Charle Andabhati, Benjamin Kirongozi and Kambale Karupao (UNIKIS: CHFS01 to CHFS05); 3 ♀♀,
9 ?? [1 SA, 11 AL], Wafanya [= Wafania], unknown date, 3 Jan. 1988, 5 Dec. 1988, 26 May 1992,
29 May 1992, between 25 Nov. 1987 and 3 Jan. 1988, between 1 Nov. 1991 and 30 Nov. 1991, between
1 Jan. 1992 and 31 Jan. 1992, leg. P. Lootens (RMCA: 92.079-M-0123, 92.079-M-0124, 92.079-M0038, 92.079-M-0037, 92.079-M-0039, 92.079-M-0040, 88.005-M-0001, 88.005-M-0002, 89.020-M0045, 92.079-M-0041, 92.079-M-0036, 88.005-M-0007); 1 ♂ [UN], Yaekela, 7 May 2010, leg. Guy229
European Journal of Taxonomy 382: 1–327 (2017)
Crispin Gembu Tungaluna, Prescott Musaba and José Akaibe (UNIKIS: CRT1888); 2 ♀♀ [1 SS, 1 AL],
Yalosemba, 17 Jun. 1979, between 1 Jun. 1979 and 30 Jun. 1979, leg. C. Brian Robbins (USNM:
537736, MHNG: 1870.091); 1 ♀ [SS], Yangambi, 22 Aug. 1947, leg. unknown collector (RBINS:
6988); 3 ♂♂ [3 UN], Zobia, prior to 18 Apr. 1946, leg. unknown collector (RBINS: 14519 to 14521).
Mimetillus thomasi Hinton, 1920
DEMOCRATIC REPUBLIC OF THE CONGO: 1 ? [AL], Elisabethville [= Lubumbashi], between 1
Mar. 1926 and 31 Mar. 1926, leg. Henri Schouteden (RMCA: 16496).
Neoromicia Roberts, 1926
BURUNDI: 1 ♀ [AB], Kanyosha, 14 Apr. 1970, leg. P.L. Niort (MHNG: 1608.004).
DEMOCRATIC REPUBLIC OF THE CONGO: 1 ad ♀, 4 ad ♂♂ [5 AL], Cel II, 22 Apr. 1951,
1 Jun. 1951, 23 Jul. 1951, 10 Nov. 1951, 26 May 1952, leg. H. de Saeger et al. (RBINS: 13831, 13837,
13830, 13833, 13839); 1 ♀ [AL], Kinkole, 15 Aug. 1980, leg. unknown collector (ZMA: MAM.21154);
1 ad ♂ [AL], Ndelele, 3 Dec. 1951, leg. H. de Saeger et al. (RBINS: 13836); 1 juv ? [UN], Rutshuru,
unknown date, leg. unknown collector (INPBC: V1397.2); 1 ♂ [SS], Stanleyville [= Kisangani],
13 Jan. 1927, leg. Harold Jefferson Coolidge and L. Whitman (MCZ: 23382); 1 ad ♀ [SA], Zobia,
18 Apr. 1946, leg. R.P. Van Woensel (RBINS: 14508).
RWANDA: 2 ♀♀ [2 AL], Gisenyi, 5 Oct. 1980, leg. Hans H. de Iongh (ZMA: MAM.21237, MAM.21238).
Neoromicia brunnea (Thomas, 1880)
DEMOCRATIC REPUBLIC OF THE CONGO: 1 ad ♀ [AL], Lukolela, 24 Dec. 1930, leg. James Paul
Chapin (AMNH: 86933).
Neoromicia capensis (A. Smith, 1829)
BURUNDI: 1 ad ♀ [SB], Makamba, 23 Mar. 1970, leg. P.L. Niort (MHNG: 1607.097).
DEMOCRATIC REPUBLIC OF THE CONGO: 1 ? [UN], Ango, unknown date, leg. unknown collector
(RMCA: 12598); 3 ad ♀♀ [3 AL], Bagbele Moko, 3 Apr. 1950, leg. H. de Saeger et al. (RBINS:
13840.1 to 13840.3); 1 sad ♂, 1 juv ♂ [2 AL], Beno, between 1 Jan. 1949 and 31 Dec. 1949, leg.
P. Bayet (RMCA: 19851, 19852); 2 juv ♀♀, 1 imm ♂, 1 ♀, 2 ♂♂ [6 AL], Boende, between 1 Aug. 1972
and 31 Aug. 1972, between 1 Jan. 1969 and 31 Dec. 1974, between 1 Jan. 1967 and 31 Dec. 1967,
leg. P. Lootens (RMCA: 38717, 38718, 38725, 75.035-M-0024, 75.035-M-0025, 75.035-M-0057); 3 ??
[1 SA, 2 UN], Buta, between 1 Jun. 1934 and 30 Jun. 1934, leg. Jozef Hutsebaut (RMCA: 12590a to
12590c); 1 ad ♀ [AL], Cel I [= Bagbele], 3 Apr. 1950, leg. H. de Saeger (RMCA: 36852); 8 ad ♀♀,
3 ad ♂♂, 2 imm ♀♀, 5 imm ♂♂, 1 ♂, 2 ?? [1 SS, 17 AL, 3 UN], Cel II, unknown date, leg. unknown
collector, 16 Mar. 1951, 24 Aug. 1951, leg. H. de Saeger et al., 16 Mar. 1951, 25 Feb. 1952, between
8 Jun. 1951 and 26 Nov. 1951, between 8 Apr. 1951 and 8 Jun. 1951, leg. H. de Saeger (INPBC:
3559, 4360, RBINS: 13841.1 to 13841.5, RMCA: 36853 to 36862, RBINS: 13834, RMCA: 36866,
36864, 36865); 1 ad ♂, 1 ad ? [1 SS, 1 SA], Elisabethville [= Lubumbashi], unknown date, leg.
Guarre, between 1 Mar. 1926 and 31 Mar. 1926, leg. Henri Schouteden (BMNH: 1954.859, RMCA:
8872); 1 juv ♀, 1 juv ♂ [2 UN], Gangala-na-Bodio, 1 May 1948, leg. unknown collector (RBINS:
11656.1, 11656.2); 1 ad ♂ [SA], Garamba National Park, 14 Mar. 1912, leg. Herbert Lang, James Paul
Chapin and The American Museum Congo Expedition (AMNH: 49340 [holotype Eptesicus garambæ
J.A. Allen, 1917]); 1 ad ♂ [SC], Kipangaribwe River, 12 Jan. 1948, leg. l’ Institut des Parcs nationaux
du Congo belge (RBINS: 10707); 1 ad ♂ [SS], Kisantu, 27 Dec. 1920, leg. Henri Schouteden (RMCA:
230
VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
6560); 1 ad ♀ [SA], Léopoldville [= Kinshasa], between 1 Jan. 1961 and 31 Dec. 1961, leg. Xavier
Misonne (RBINS: 14507); 1 ad ♂ [SS], Luebo, 10 Sep. 1921, leg. Henri Schouteden (RMCA: 7309);
2 ad ♀♀, 1 ad ? [1 SS, 2 AL], Mabanga, 1 Aug. 1952, leg. H. de Saeger et al., 1 Aug. 1952, leg.
H. de Saeger (RBINS: 13848.1, 13848.2, RMCA: 36863); 1 ad ♂ [AL], Mayumbe, unknown date, leg.
unknown collector (RMCA: 90.012-M-0001); 2 ad ♀♀ [2 AL], Moba, 6 Feb. 1954, 26 Feb. 1954, leg.
H. Bomans (RMCA: 26327, 22126); 1 ? [SS], PFSK.20, 16 Jun. 1952, leg. H. de Saeger et al. (RBINS:
13847); 2 juv ?? [2 UN], PPK.51, 2 Apr. 1952, leg. H. de Saeger et al. (RBINS: 13846.1, 13846.2); 1 ad
♂, 1 ad? ♂, 1 sad ♀ [3 AL], Wafanya [= Wafania], 5 Dec. 1987, 10 Dec. 1988, between 21 Nov. 1987
and 28 Nov. 1987, leg. P. Lootens (RMCA: 89.020-M-0046, 88.005-M-0014, 88.005-M-0003).
Neoromicia guineensis (Bocage, 1889)
DEMOCRATIC REPUBLIC OF THE CONGO: 1 ad ♂ [SA], Isiro, 3 Jul. 1913, leg. Herbert Lang,
James Paul Chapin and The American Museum Congo Expedition (AMNH: 49339); 1 ad? ♂ [SA],
Niangara, 27 Mar. 1913, leg. Herbert Lang, James Paul Chapin and The American Museum Congo
Expedition (AMNH: 49290).
Neoromicia nana (Peters, 1852)
BURUNDI: 1 ♀ [SK], unknown locality, 15 Mar. 1979, leg. Heinz Stephan (SMF: 57574); 2 ad ♀♀
[2 AL], Bujumbura, 22 Apr. 1982, leg. Danny Meirte, between 1 Jul. 1968 and 31 Jul. 1968, leg. Lewall
(RMCA: 82.016-M-0009, 35155); 1 ad ♀, 5 ad ♂♂, 1 ♂ [7 NS], Busiga, 5 Jun. 1969, leg. P.L. Niort
(MHNG: 1607.057 to 1607.063); 1 ad ♂ [AL], Bweru, 29 Jul. 1976, leg. Jacques Verschuren (RBINS:
19912); 1 ad ♀, 2 ad ♂♂ [3 NS], Kanyinya, 6 Jun. 1969, leg. P.L. Niort (MHNG: 1607.064 to 1607.066);
1 ♀ [SK], Kirundo Province, 9 Mar. 1979, leg. Heinz Stephan (SMF: 57572); 2 ad ♂♂ [2 NS], Mpinga
Cave [= Mpinga Mission], 1 Jan. 1969, leg. P.L. Niort (MHNG: 1607.051, 1607.072); 1 ♀ [SK], Ngozi
Province, 13 Mar. 1979, leg. Heinz Stephan (SMF: 57573); 3 ad ♂♂ [3 NS], Nyankanda, 3 Jan. 1969,
leg. P.L. Niort (MHNG: 1607.074 to 1607.076); 1 ad ♂ [SO], Nyanza [= Nyanza-Lac], 28 Dec. 1932,
leg. Burgeon (RMCA: 12044); 4 ad ♂♂, 1 ♂ [5 NS], Rugari Mission, 7 Jun. 1969, 8 Jun. 1969, leg.
P.L. Niort (MHNG: 1607.067 to 1607.071); 1 ? [UN], Russissi, bank [= Rusizi Delta], unknown date,
leg. Rudolf Grauer (ZMB: T09305); 1 ad ♀, 1 ♂ [1 SA, 1 UN], Rutana, 6 Mar. 1979, leg. Heinz Stephan
(SMF: 57570, 57571); 1 ? [AL], Tora, 19 Nov. 1975, leg. Walter N. Verheyen (RMCA: 97.080-M-0232).
DEMOCRATIC REPUBLIC OF THE CONGO: 1 ad ♂, 1 juv ♂, 14 ♀♀, 8 ♂♂, 21 ?? [20 SS, 18 AL,
3 SK, 1 SO, 3 UN], unknown locality, unknown date, leg. Herbert Lang, James Paul Chapin and The
American Museum Congo Expedition, unknown date, between 1 Apr. 1999 and 30 Apr. 1999, leg.
unknown collector, unknown date, leg. The Alexander-Gosling Expedition, unknown date, leg. unknown
collector, unknown date, leg. Guy-Crispin Gembu Tungaluna, prior to 1 Aug. 1957, leg. Heinz Stephan,
prior to 22 Dec. 1943, leg. Gaston-François de Witte, prior to 4 Jul. 1958, leg. E. Jans (AMNH: 51279,
BMNH: 1906.7.1.10 to 1906.7.1.13, 1906.7.1.15 to 1906.7.1.17, 1906.7.1.7 to 1906.7.1.9, 1907.7.8.36,
1907.7.8.37, 1926.7.6.85 to 1926.7.6.91, 1930.11.11.166, RMCA: 89.004-M-0013, 89.004-M-0014,
a1.097-M-0565 to a1.097-M-0581, SMF: 93321, RMCA: 27468, 27469, 16639, 27369); 1 ad ♀, 3 ad
♂♂, 1 ♀, 1 ♂ [3 SS, 2 AL, 1 SO], Aba, 10 Aug. 1951, leg. H. de Saeger et al., 18 Dec. 1911, leg. Herbert
Lang, James Paul Chapin and The American Museum Congo Expedition (RBINS: 13862.1, 13862.2,
13866, AMNH: 48979 [holotype Pipistrellus abaensis J.A. Allen, 1917], 48980 [topotype Pipistrellus
abaensis J.A. Allen, 1917], 48981 [topotype Pipistrellus abaensis J.A. Allen, 1917]); 2 ad ♀♀, 3 ad
♂♂, 1 ♂ [4 SS, 1 SO, 1 SC], Abimva, 22 Jun. 1925, 23 Jun. 1925, leg. Henri Schouteden (RMCA: 9025,
9028a, 9030 to 9033); 1 ♀ [UN], Aketi, 22 Jun. 2015, leg. Vlir/Aketi (UNIKIS: AKETI607); 2 ??
[2 UN], Albert National Park [= Virunga National Park], unknown date, leg. unknown collector
(INPBC: V1397.1, V1708); 1 ad ♂, 1 sad ♂ [2 AL], Albertville [= Kalemie], prior to 18 Nov. 1959, leg.
H. Bomans, prior to 30 Jan. 1900, leg. Célestin Hecq (RMCA: 29302, 421); 1 ? [AL], Albertville-Moba
231
European Journal of Taxonomy 382: 1–327 (2017)
(road?) [= Kalemie-Moba (road?)], 23 Apr. 1954, leg. H. Bomans (RMCA: 23320); 1 ad ♀ [AL],
Ambarau, between 1 Jan. 1953 and 31 Dec. 1953, leg. H. Dewulf (RMCA: 21839); 6 ♀♀, 7 ♂♂
[13 SS], Bafuaboka [= Bafwabaka], 5 Jan. 1910, 6 Jan. 1910, 7 Jan. 1910, leg. Herbert Lang, James
Paul Chapin and The American Museum Congo Expedition (AMNH: 48951, 48952, 48954 to 48964);
1 ad ♂, 1 juv ♂, 6 ♀♀, 9 ♂♂ [8 SS, 6 SA, 3 AL], Bafwaboka [= Bafwabaka], 4 Jan. 1910, 5 Jan. 1910,
7 Jan. 1910, leg. Herbert Lang, James Paul Chapin and The American Museum Congo Expedition
(AMNH: 48943 to 48945, 49322, 48946, 48948 to 48950, 49295, RMCA: 12369, AMNH: 49293,
49294, 49296 to 49299, 49302); 1 ad ♀, 2 juv ♀♀, 1 juv ♂, 4 ?? [4 AL, 4 UN], Bagbele, 13 Apr. 1950,
leg. H. de Saeger et al., 15 Apr. 1950, leg. H. de Saeger (RBINS: 13802.3 to 13802.6, RMCA: 36867 to
36870); 17 ad ♀♀, 24 ad ♂♂, 1 ad ?, 1 ad? ♀, 2 ad? ♂♂, 2 juv ??, 1 ? [44 AL, 3 UN, 1 SC], Bagbele
Moko, unknown date, leg. unknown collector, 24 Feb. 1950, 13 Apr. 1950, 26 Jul. 1952, leg. H. de
Saeger et al., 26 Jul. 1952, leg. H. de Saeger (RBINS: 13867, 13802.1, 13802.2, 13861, 13864.01 to
13864.15, 13865.01 to 13865.12, RMCA: 36871 to 36887); 2 ♀♀ [2 UN], Bagisana, 21 Apr. 1952, leg.
H. de Saeger et al. (RBINS: 13863.1, 13863.2); 1 juv ♀ [SK], Bambesa, 18 Oct. 1940, leg. J.M. Vrijdagh
(RMCA: 17766); 1 ♂ [SS], Banalia, between 26 Jan. 1939 and 26 Dec. 1939, leg. Duncan M. Hodgson
and William F. Coultas (MCZ: 38029); 1 ad ♀, 3 ad ♂♂, 1 ad? ♀, 1 juv ♂ [6 AL], Banana, 18 Aug. 1951,
12 Apr. 1952, between 1 Jan. 1952 and 31 Jan. 1952, leg. Is. Mesmaekers (RMCA: 20763, 20764, 20525
to 20527, 20749); 2 ad ♀♀ [1 SS, 1 SO], Basongo, 19 Jul. 1921, leg. Henri Schouteden (RMCA: 6644,
6645); 1 ad ♀, 2 ad ♂♂, 1 ♀, 1 ♂ [2 SS, 3 AL], Batala’s Village, 13 Apr. 1925, 16 Apr. 1925, leg.
Edmund Heller (FMNH: 26495, 26496, ZMUC: 1541 to 1543); 1 ? [AL], Batoko (River) [= OlongoNsongo [River]], 18 Nov. 1913, leg. J. Bequaert (RMCA: 5388); 1 ad ♂, 4 juv ♀♀, 1 juv ♂, 1 juv ?
[6 SS, 1 SC], Bikoro, 27 Feb. 1921, leg. Henri Schouteden (RMCA: 6618 to 6620, 6622 to 6625); 1 ad
♀, 1 ad ♂, 1 sad ♀ [3 SS], Bitshumbi [= Vitshumbi], prior to 22 Dec. 1943, leg. Gaston-François de
Witte (RMCA: 16640 to 16642); 1 ad ♀, 1 ad ♂ [2 AL], Bodjoki - Bumba, between 1 Jan. 1986 and
31 Dec. 1986, between 1 Jan. 1985 and 31 Dec. 1986, leg. WHO and Marc Colyn (RMCA: 88.011-M0019, 88.011-M-0018); 21 ad ♀♀, 41 ad ♂♂, 8 ad? ♀♀, 9 ad? ♂♂, 1 sad ♀, 2 sad ♂♂, 4 juv ♀♀, 2 juv
♂♂ [88 AL], Boende, 18 Nov. 1966, between 1 Nov. 1969 and 30 Nov. 1969, between 1 Jan. 1969 and
31 Dec. 1974, between 1 Jan. 1969 and 31 Jan. 1969, between 1 Jan. 1967 and 31 Dec. 1967, between
1 Jan. 1966 and 31 Jan. 1966, leg. P. Lootens (RMCA: 38716, 34076, 34079 to 34081, 34083 to 34092,
34094, 34096 to 34099, 34101 to 34108, 34110 to 34113, 34115 to 34119, 34121 to 34124, 34126,
34127, 34129 to 34145, 34147 to 34155, 38715, 38719 to 38723, 38726 to 38730, 35640, 75.035-M0027, 75.035-M-0030 to 75.035-M-0035); 1 ? [AL], Bokala, prior to 13 Oct. 1913, leg. Maes (RMCA:
2401); 16 ad ♀♀, 20 ad ♂♂, 4 ad? ♀♀, 8 ad? ♂♂, 5 sad ♀♀, 2 sad ♂♂, 3 juv ♀♀, 6 juv ♂♂, 1 juv ?, 2
imm ♀♀, 1 imm ♂, 1 imm ?, 3 ?? [71 AL, 1 UN], Bokuma, 2 Mar. 1952, 15 Mar. 1952, 23 Mar. 1952,
25 Jul. 1952, 23 Feb. 1954, 10 Mar. 1954, prior to 6 May 1954, prior to 25 Aug. 1954, between
2 Mar. 1952 and 8 Mar. 1952, between 1 Jun. 1953 and 30 Jun. 1953, between 1 Feb. 1954 and
28 Feb. 1954, between 1 Jan. 1954 and 31 Dec. 1954, between 1 Jan. 1954 and 31 Jan. 1954, between
1 Jan. 1952 and 31 Dec. 1952, leg. P. Lootens (RMCA: 22180, 21403 to 21405, 21398 to 21401, 22156
to 22158, 21406 to 21408, 21671, 21397, 22154, 22159 to 22167, 22319, 23248 to 23269, 22155,
22039, 22042, 22043, 22045 to 22048, 21402, 22320, 22321, 22168 to 22179); 1 juv ♂ [SK], Boma,
between 1 Mar. 1887 and 31 Mar. 1887, leg. P. Hesse (SMF: 2523 [holotype Vesperus pusillus Noack,
1889]); 1 ♂ [AL], Bombia Bonelu, 19 Jun. 1979, leg. Frieder Meier (MHNG: 1870.081); 1 ♀ [SO],
Bomili, 17 Aug. 1938, leg. Duncan M. Hodgson and William F. Coultas (MCZ: 38028); 1 ad ♀ [SS],
Bosodula [= Boso-Dula], 25 Oct. 1942, leg. J.M. Vrijdagh (RMCA: 17755); 5 ad ♀♀, 1 ad ♂, 1 ad? ♂,
1 sad ♂, 2 imm ♀♀ [10 AL], Boteka, 25 Oct. 1985, between 22 Jul. 1984 and 15 Mar. 1985, between
1 Feb. 1985 and 28 Feb. 1985, leg. P. Lootens (RMCA: 85.052-M-0004 to 85.021-M-0012, 85.021-M0004); 5 ad ♀♀, 3 ad ♂♂, 1 ad? ♂, 1 juv ♀, 1 juv ♂, 10 ?? [11 AL, 10 UN], Bugara, unknown date, leg.
unknown collector, 3 Dec. 1953, leg. Gaston-François de Witte (INPBC: W710.02 to W710.11, RMCA:
37201 to 37211); 1 ad ♂ [AL], Bukavu, between 1 Jul. 1971 and 31 Jul. 1971, leg. Alexandre Prigogine
(RMCA: 38583); 10 ?? [10 UN], Bulaya, unknown date, leg. unknown collector (INPBC: W1178.1 to
232
VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
W1178.9, W868); 1 ♂ [SA], Bulundu, unknown date, leg. Uwe Rahm (BMNH: 1963.1179); 1 ad? ♂
[AL], Bumba, prior to 23 Jul. 1946, leg. P. Lootens (RMCA: 18114); 1 ♂ [AL], Bunia, between
1 Apr. 1927 and 30 Apr. 1927, leg. Harold Jefferson Coolidge (MCZ: 28544); 1 juv ? [UN], Bunkeya,
24 Mar. 1958, leg. Van Oost (RMCA: 28803); 3 ad ♀♀, 5 ad ♂♂, 1 ad? ♂, 2 sad ♂♂, 2 juv ♀♀, 2 juv
♂♂, 36 ?? [18 AL, 33 UN], Bunyangura, unknown date, leg. unknown collector, 9 Dec. 1953,
22 Nov. 1954, 11 Jan. 1955, leg. Gaston-François de Witte (INPBC: W795.01 to W795.33, RMCA:
37363, 37355 to 37360, 37212 to 37222); 6 ad ♀♀, 9 ad ♂♂, 1 sad ♂, 2 imm ♀♀, 1 imm ♂, 22 ??
[19 AL, 22 UN], Bunyenzi, unknown date, leg. unknown collector, 15 Sep. 1953, 17 Sep. 1953,
3 Feb. 1954, leg. Gaston-François de Witte (INPBC: W1007.01 to W1007.20, W624, W632, RMCA:
37189, 37190, 37193, 37248 to 37263); 1 ad ♀, 1 ad ♂ [2 AL], Busholinka River, 18 Feb. 1954, leg.
Gaston-François de Witte (RMCA: 37223, 37224); 4 ad ♂♂ [4 AL], Busu Mokalu [= Busu-Mokala],
between 5 Nov. 1938 and 10 Nov. 1938, leg. Baudart (RMCA: 18093 to 18096); 5 ad ♀♀, 2 ad ♂♂,
1 ad? ♀, 1 sad ♀, 4 juv ♂♂, 1 ? [13 AL, 1 UN], Buta, unknown date, prior to 7 Mar. 1940, prior to
3 Nov. 1939, leg. Jozef Hutsebaut (RMCA: 14563a to 14563c, 14581, 14586, 16572, 16573, 15731 to
15736, 15911); 1 ad ♀ [AL], Butembo, prior to 12 Sep. 1958, leg. P. Dyleff (RMCA: 27397); 3 ad ♀♀,
3 ad ♂♂, 1 ad ?, 1 ad? ♀ [8 AL], Congo da Lemba, prior to 13 Mar. 1913, leg. Mayné (RMCA: 1544a
to 1544h); 1 ad ? [SA], Congo River, 75 mi [= 121 km] up, between 1 Aug. 1897 and 31 Aug. 1897,
leg. George Latimer Bates (USNM: 102499); 1 ad ♂ [AL], Costermansville [= Bukavu], prior to
14 Feb. 1952, leg. H. Bomans (RMCA: 20656); 2 ?? [1 AL, 1 UN], Dika, 19 Mar. 1925, leg. Henri
Schouteden (RMCA: 16257, 16260); 1 ad ♂, 1 sad ♀ [2 AL], Dilolo, prior to 31 Mar. 1948, leg. Jozef
Hutsebaut (RMCA: 19076, 19077); 1 ad ♂ [SS], Dramba, 18 Apr. 1925, leg. Henri Schouteden (RMCA:
8991); 2 ad? ♂♂ [2 AS], Duma, between 1 Oct. 1910 and 31 Oct. 1910, leg. Ad. Tr. z. U. (ZMH:
41082?, 41083?); 3 ad ♀♀, 7 ad ♂♂, 1 ad? ♂ [11 AL], Dungu, 5 Oct. 1950, leg. H. de Saeger et al.,
between 1 Apr. 1925 and 30 Apr. 1925, leg. Henri Schouteden, prior to 11 Jul. 1912, leg. M. Hutereau
(RBINS: 13803, RMCA: 16192, 1105a to 1105i); 2 juv ♀♀ [1 SS, 1 SO], Eala, 15 Mar. 1921, leg. Henri
Schouteden (RMCA: 6626, 6627); 1 ad? ♀ [AL], Elila, 16 Jul. 1947, leg. Max Poll (RMCA: 18044);
4 ad ♀♀, 2 ad ♂♂, 1 ad ?, 1 sad ♀, 1 sad ♂ [7 AL, 2 SO], Elisabethville [= Lubumbashi], unknown
date, 22 Dec. 1955, leg. Félix / Michel Anciaux de Faveaux, 2 Sep. 1954, between 1 Dec. 1954 and
31 Dec. 1954, leg. De Smet, 22 Dec. 1955, leg. M. Lips (MNHN: ZM-MO-1957-126 A, RMCA: 22456,
26220, 26332, 26344, 26345, 26365, 22718, 22719); 1 ad ♀, 4 ad ♂♂, 2 juv ♂♂, 1 ♀, 5 ♂♂, 2 ?? [8 SS,
2 SA, 1 AL, 2 SO, 2 UN], Faradje, 18 Feb. 1911, 1 Mar. 1911, 8 Aug. 1911, 18 Feb. 1912, leg. Herbert
Lang, James Paul Chapin and The American Museum Congo Expedition, 1 Jul. 1925, 5 Jul. 1925,
6 Jul. 1925, leg. Henri Schouteden (AMNH: 48978, 52112, 52113, RMCA: 9897 to 9901, AMNH:
48977, 49303, 49307, 49332, RMCA: 9059, 9045, AMNH: 48976); 1 ? [SA], Faradje Territory,
18 Feb. 1912, leg. Herbert Lang, James Paul Chapin and The American Museum Congo Expedition
(AMNH: 49316); 1 ad ♀, 1 juv ♂, 1 ♀ [2 AL, 1 UN], Flandria [= Boteka], 27 Aug. 1929, between
1 Jan. 1930 and 31 Dec. 1930, leg. R.P. Hulsaert (RMCA: 89.004-M-0012, 90.007-M-0001, 16278); 1 ♂
[SA], Fomulac-Katana, between 2 Oct. 1963 and 3 Oct. 1963, leg. P. Kunkel (SMF: 31837); 4 ??
[4 UN], Fuko, unknown date, leg. unknown collector (INPBC: W1379.1 to W1379.3, W854); 1 ad ♀,
2 ad ♂♂ [3 AL], Fuko (River), 7 Apr. 1954, leg. Gaston-François de Witte (RMCA: 37282 to 37284);
1 ? [UN], Funda Biabo, 20 Mar. 1914, leg. Charliers (RMCA: 5384); 3 ?? [3 UN], Gahama, unknown
date, leg. unknown collector (INPBC: W1241.1 to W1241.3); 1 ♂ [SS], Gamangui, 27 Feb. 1910, leg.
Herbert Lang, James Paul Chapin and The American Museum Congo Expedition (AMNH: 48965);
1 ad ♂ [AL], Ganda-Sundi, 31 Mar. 1964, leg. Alexandre Fain (RMCA: 33276); 3 ♂♂ [1 SS, 2 SO],
Gangala [= Gangala-na-Bodio], 24 Mar. 1957, prior to 1 Aug. 1957, leg. Heinz Stephan (RMCA:
27470 to 27472); 2 ad ♀♀, 2 ad ♂♂, 1 ad ?, 1 ad? ♂ [1 SS, 5 AL], Gangala-na-Bodio, 26 Nov. 1947,
leg. M. Micha, 5 Apr. 1948, 9 Apr. 1948, 6 May 1948, 7 May 1948, leg. Hediger and Jacques Verschuren,
8 Nov. 1949, leg. H. de Saeger et al. (RBINS: 12947, 8191, 8193, 8194, 13860, 8192); 1 ? [SA],
Garamba National Park, 15 Mar. 1912, leg. Herbert Lang, James Paul Chapin and The American
Museum Congo Expedition (AMNH: 49311); 1 ad ♀, 2 ad ♂♂, 1 juv ♀, 3 juv ♂♂, 1 imm ♀, 1 imm ♂,
233
European Journal of Taxonomy 382: 1–327 (2017)
3 ?? [11 AL, 1 UN], Geramihasha River, unknown date, leg. unknown collector, 22 Mar. 1954, 3 May
1954, 28 Jun. 1954, 3 May 1955, between 3 May 1954 and 3 May 1955, leg. Gaston-François de Witte
(INPBC: W997, RMCA: 37292 to 37294, 37326 to 37329, 37302, 37299 to 37301); 2 ad ♀♀, 1 sad ♀,
2 sad ♂♂, 1 imm ♂ [6 AL], Ibambi, prior to 24 Jun. 1939, leg. Reitter (RMCA: 15390 to 15395); 1 sad
♀ [AL], Ibembo, prior to 22 Feb. 1952, leg. Jozef Hutsebaut (RMCA: 20662); 1 ad ♀, 1 ad ♂, 2 ♀♀,
2 ♂♂ [4 SS, 2 AL], Idjwi Island, 14 Jan. 1960, leg. Uwe Rahm, 4 Mar. 1939, leg. Arthur Loveridge
(RMCA: 28919, 28920, MCZ: 40763 to 40766); 2 ?? [2 AL], Ikela, prior to 4 Jan. 1958, leg. P. Lootens
(RMCA: 27023, 27024); 1 juv ♂, 4 ♀♀, 5 ♂♂, 4 ?? [4 AL, 9 SK, 1 UN], Irangi, 8 Jan. 1957, 25 Jan. 1984,
29 Jan. 1984, 1 Feb. 1984, leg. Heinz Stephan, 3 Jan. 1957, leg. J.J. Laarman (SMF: 65023, 65024,
65013 to 65022, RMNH: MAM.16372, RMCA: 27473); 1 ? [UN], Iringi, prior to 31 Aug. 1899, leg.
A. or G. Lindeman (RMCA: 407); 1 ♀ [SO], Irsac Labo [= Luiro], 14 Dec. 1955, leg. J.J. Laarman
(RMNH: MAM.16370); 1 ♂ [UN], Isai River, 19 Dec. 1974, leg. Michael D. Gallagher (HZM:
206.7973); 2 ?? [2 UN], Ishango River, unknown date, leg. unknown collector
(INPBC: W1372.1, W1372.2); 2 ad ♀♀, 1 ad? ♀, 1 sad ♀, 1 ♀ [4 AL, 1 UN], Jadotville [= Likasi],
1 Dec. 1957, 8 Jun. 1958, leg. Félix / Michel Anciaux de Faveaux, 18 Dec. 1959, leg. unknown collector
(RMCA: 27581, 27582, RBINS: 14531, RMCA: 27583, 27584); 1 ad ♀, 3 ♀♀, 4 ♂♂ [7 SS, 1 SK],
Kabengere, 13 Dec. 1926, 22 Dec. 1926, 24 Dec. 1926, leg. John Todd Zimmer (FMNH: 29418 to
29424, 30686); 2 ad ♂♂ [1 SS, 1 SC], Kabira, 12 Mar. 1959, 15 Mar. 1959, leg. Uwe Rahm (RMCA:
28915, 28914); 1 ad ♂ [AL], Kabongo, between 1 Jan. 1952 and 31 Dec. 1952, leg. M. Dierckx (RMCA:
21483); 1 ad ♀, 1 ad ♂, 1 ad? ♂, 19 ?? [3 AL, 19 UN], Kadehe River, unknown date, leg. unknown
collector, 18 Sep. 1953, leg. Gaston-François de Witte (INPBC: W1426.01 to W1426.14, W636, W638,
W660, W779, W953, RMCA: 37195 to 37197); 1 ad ♀ [AL], Kadjidji [= Kajuju], unknown date, leg.
unknown collector (MNHN: ZM-MO-1985-1069); 2 ad ♀♀, 3 ad ♂♂, 1 ad? ♀, 1 juv ♀, 1 juv ♂, 1 imm
♀ [9 AL], Kadjudju [= Kajuju], prior to 6 Jul. 1935, leg. Guy Babault (RMCA: 12847a to 12847c,
12848a to 12848f); 1 ad ♀, 2 ad ♂♂, 1 ad? ♀, 1 juv ♂, 1 imm ♀, 15 ?? [6 AL, 15 UN], Kahama
[= Gahama], unknown date, leg. unknown collector, 9 Jun. 1955, leg. Gaston-François de Witte
(INPBC: W1496, W926.01 to W926.14, RMCA: 37367 to 37372); 3 ad ♂♂, 1 ad? ♀ [3 AL, 1 SO],
Kakanda Cave, 22 Sep. 1960, leg. Félix / Michel Anciaux de Faveaux, 9 Aug. 1964, leg. Alexandre
Prigogine, between 21 Apr. 1955 and 22 Apr. 1955, leg. R.P. Th. de Caters (RBINS: 14529, 14530,
RMCA: 33068, 22888); 1 ad ♀ [AL], Kakanda Mutaka, between 6 Feb. 1954 and 15 Feb. 1954, leg.
R.P. Th. de Caters (RMCA: 22379); 1 ad ♀ [UN], Kakoi, 5 Aug. 1995, leg. Herwig Leirs et al. (MSB:
274920); 2 ad ♂♂, 1 ad? ♂ [3 AL], Kakonde, 20 Oct. 1956, leg. Félix / Michel Anciaux de Faveaux
(RMCA: 26239, 26240, 26244); 1 ? [UN], Kakondo, unknown date, leg. unknown collector (NMP:
16873); 1 ♀ [SS], Kalanda, 7 Aug. 1934, leg. B.L. Bennett (MCZ: 31982); 7 ad ♀♀, 2 ad ♂♂ [9 AL],
Kalulako, unknown date, 5 Jan. 1956, leg. Félix / Michel Anciaux de Faveaux, 5 Jan. 1956, 4 Nov. 1956,
leg. D’Haenens (MNHN: ZM-MO-1957-126 B, RMCA: 26388 to 26393, 26221, 26384); 5 ad ♀♀, 1 ad
♂, 1 sad ♀, 1 sad ♂, 1 ♂ [2 SS, 6 AL, 1 SO], Kamituga, 17 Feb. 1964, 20 Feb. 1964, 20 Mar. 1964,
between 21 Feb. 1964 and 21 Mar. 1964, leg. Alexandre Prigogine (RMCA: 32587, 32857 to 32861,
32585, 32586, 32862); 3 ad ♀♀, 2 ad ♂♂ [5 AL], Kamp U.M.H.K. [= Kamp u.m.H(aut).K(atanga).],
25 Aug. 1954, 27 Aug. 1954, 28 Aug. 1954, 2 Sep. 1954, leg. De Smet (RMCA: 22457, 22452 to 22455);
2 ?? [2 SS], Kampala, between 1 Mar. 1935 and 31 Mar. 1935, leg. The Grandidier Collection (MCZ:
57539, 57540); 2 ♀♀, 1 ♂ [1 SS, 2 AL], Kananga, 8 Sep. 1963, 13 Mar. 1964, leg. Antoon Emeric
Marcel De Roo, 30 Nov. 1973, leg. Western New Mexico University (AMNH: 212580, FMNH: 152772,
AMNH: 212579); 6 ad ♀♀, 18 ad ♂♂, 5 ad? ♂♂, 2 sad ♀♀, 30 ?? [31 AL, 30 UN], Kantundwe,
unknown date, leg. unknown collector, 18 Sep. 1953, 19 Mar. 1954, 30 Mar. 1954, 10 Jun. 1954,
5 Jul. 1954, 17 Sep. 1954, 17 May 1957, between 3 Nov. 1954 and 3 Dec. 1954, leg. Gaston-François de
Witte (INPBC: W1258, W1491.01 to W1491.25, W617, W634, W640, W721, RMCA: 37336 to 37342,
37173, 37350 to 37352, 37194, 37285 to 37291, 37295 to 37297, 37343 to 37349, 37353, 37354); 3 ??
[3 UN], Kanyawamukunzi River, unknown date, leg. unknown collector (INPBC: W754.1 to W754.3);
1 ad ♂, 9 ?? [1 AL, 9 UN], Kanzarue River, unknown date, leg. unknown collector, 7 Sep. 1953, leg.
234
VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
Gaston-François de Witte (INPBC: W766.1 to W766.9, RMCA: 37198); 1 ? [AL], Kapanga, prior to
10 Dec. 1958, leg. J. Allaer (RMCA: 27667); 1 ad ♀ [AL], Kapaso, 7 Oct. 1955, leg. M. Lips (RMCA:
26381); 1 ad ♀ [AL], Kaponda, 14 Oct. 1955, leg. M. Lips (RMCA: 26337); 1 ♂ [AL], Karambi,
3 Apr. 1992, leg. Ngenge Masumbuko Kamitongo (ZMA: MAM.24548); 5 ad ♀♀, 1 ad ♂, 4 juv ♂♂
[10 AL], Karavia [Mine], 30 Sep. 1955, 7 Nov. 1955, between 3 Sep. 1955 and 30 Sep. 1955, leg.
M. Lips (RMCA: 26346, 26360 to 26364, 26374 to 26376, 26347); 1 imm ♀ [AL], Karawa, prior to
24 Nov. 1936, leg. R.P. Wallin (RMCA: 14569); 1 ad ♂, 1 ♂, 4 ?? [1 AL, 5 UN], Kasai, between
1 Jan. 1923 and 31 Dec. 1923, leg. Achten, prior to 11 Jul. 1905, leg. La Compagnie du Kassai (RMCA:
89.004-M-0005, 90.005-M-0001, 739a to 739d); 1 ad ♂ [AL], Kasebaya River [= Kasebeya River],
15 Feb. 1954, leg. Gaston-François de Witte (RMCA: 37270); 1 ad ♀ [AL], Kasenga, between
1 Jul. 1954 and 31 Jul. 1954, leg. De Smet (RMCA: 22440); 1 ad ♀ [AL], Kasongo, 10 Jan. 1956, leg.
M. Lips (RMCA: 26335); 1 ad ♀ [AL], Katana, unknown date, leg. unknown collector (MNHN: ZMMO-1985-1068); 1 ♀, 2 ♂♂ [2 SA, 1 AL], Katanga (former) Province, 21 Sep. 1955, 20 Oct. 1956,
leg. unknown collector (SMF: 16870 to 16872); 1 ad? ♂ [AL], Kataye Sukunga, between 1 Jan. 1930
and 31 Dec. 1930, leg. Fourche (RMCA: 11548); 1 ♂ [SS], Kateke [= Kateke [River]], 26 Nov. 1947,
leg. l’ Institut des Parcs nationaux du Congo belge (RBINS: 10709); 1 ? [UN], Kiamanwa, 20 Feb. 1931,
leg. H.J. Bredo (RMCA: 11709); 1 ad ♀ [SS], Kiamokoto, 7 Jul. 1948, leg. l’ Institut des Parcs nationaux
du Congo belge (RBINS: 10710); 3 ♀♀ [3 AL], Kibanda, 3 Feb. 1956, leg. Félix / Michel Anciaux de
Faveaux (MHNG: 922.083 to 922.085); 1 sad ♂ [AL], Kifumwanshi, 13 Dec. 1955, leg. M. Lips
(RMCA: 26334); 5 ad ♀♀, 3 ad ♂♂, 2 ad? ♀♀, 1 ad? ♂, 1 sad ♂, 1 juv ♀ [13 AL], Kikanda, unknown
date, 3 Feb. 1956, 6 Feb. 1956, leg. Félix / Michel Anciaux de Faveaux, 23 Sep. 1955, 2 Feb. 1956,
6 Feb. 1956, leg. M. Lips (MNHN: ZM-MO-1957-126 C, RMCA: 26336, 26339, 26343, 26379, 26380,
23817, 23869, 26224, 26338, 26340 to 26342); 1 ♀ [SA], Kikole-Pecheur, 17 May 1980, leg.
M. Mutashia (SMF: 58069); 1 ? [AL], Kikwit, between 1 Jan. 1920 and 31 Dec. 1920, leg. R.P. Vanderijst
(RMCA: 5478); 1 ? [UN], Kilia Kaparata, unknown date, leg. unknown collector (INPBC: W509);
1 ad ♀, 1 ad ♂, 1 sad ♀, 1 imm ?, 6 ?? [4 AL, 6 UN], Kilinga, unknown date, leg. unknown collector,
26 Aug. 1953, 15 Mar. 1954, leg. Gaston-François de Witte (INPBC: W1256.1 to W1256.4, W610,
W750, RMCA: 37278 to 37280, 37183); 1 ad ♀, 2 ad ♂♂ [3 AL], Kilobelobe [= Kilobilobe], 6 Jan. 1956,
between 5 Jan. 1956 and 6 Jan. 1956, leg. D’Haenens (RMCA: 26386, 26387, 26385); 1 ad ♀ [AL],
Kilwa, between 1 Jan. 1938 and 31 Jan. 1938, leg. H.J. Bredo (RMCA: 14589); 1 ♀ [AL], Kinkole,
15 Aug. 1980, leg. unknown collector (ZMA: MAM.21153); 1 ? [AL], Kipopo, between 1 Jun. 1960
and 30 Jun. 1960, leg. Dirk Thys Van den Audenaerde (RMCA: 29142); 9 ad ♀♀, 13 ad ♂♂, 1 ad? ♀,
1 ad? ♂, 5 juv ♀♀, 3 juv ♂♂, 1 ♀, 1 ♂, 3 ?? [32 AL, 5 UN], Kisangani, 13 Jan. 1975, leg. Michael
D. Gallagher, between 1 Jun. 1982 and 30 Jun. 1982, leg. Marc Colyn (HZM: 207.7974, 208.7975,
RMCA: 82.031-M-0033, 82.031-M-0036 to 82.031-M-0069); 1 ad ♀ [AL], Kisantu, unknown date,
leg. Dumont (RMCA: 3433); 16 ?? [16 UN], Kisisile, unknown date, leg. unknown collector
(INPBC: W832.01 to W832.16); 5 ad ♀♀, 3 juv ?? [8 AL], Kiswishi [River], 11 Oct. 1955, leg. Félix /
Michel Anciaux de Faveaux, 21 Sep. 1955, 11 Oct. 1955, leg. M. Lips (RMCA: 26217, 26366, 26367,
26377, 26378, 26348 to 26350); 1 sad ♂ [SO], Kiulu Mgemgo, 7 Jan. 1977, leg. Tshinyenye (RMCA:
77.012-M-0010); 1 ad ♂, 1 ad? ♂ [2 AL], Kiumba, 18 Sep. 1957, leg. Uwe Rahm (RMCA: 28916,
28917); 4 ?? [4 AL], Kivu region, between 1 Jan. 1934 and 31 Dec. 1934, leg. Guy Babault (MCZ:
46066 to 46069); 4 ad ♀♀, 1 ad ♂, 6 ad ?? [11 AL], Kivu, near, unknown date, leg. Guy Babault
(MNHN: ZM-MO-1985-1065, ZM-MO-1985-1070 to ZM-MO-1985-1079); 1 ♂ [AL], Kokonde,
4 Feb. 1956, leg. unknown collector (SMF: 16874); 3 ♀♀, 6 ♂♂, 7 ?? [16 UN], Kongolo, 7 Nov. 1974,
leg. Michael D. Gallagher (HZM: 209.7976 to 212.7979, 213.7080 to 222.7089, 253.8020, 254.8021);
1 ad ♀, 2 ad ♂♂, 1 juv ♂ [2 SS, 2 AL], Koteli [= Kotili], 3 Jan. 1925, 23 Jan. 1925, between 1 Dec. 1924
and 31 Dec. 1924, leg. Henri Schouteden (RMCA: 8921, 8877, 16273, 16274); 5 ad ♀♀, 7 ad ♂♂,
1 ad ?, 1 juv ♀ [12 SS, 2 SO], Kunungu, prior to 19 Nov. 1937, between 1 Jan. 1922 and 31 Dec. 1922,
leg. Ngwe, prior to 22 Oct. 1930, prior to 1 Aug. 1939, leg. N’Kele (RMCA: 7752 to 7755, 15515, 14159
to 14166, 10616); 1 ? [AL], Lake Kivu, near, between 1 Jan. 1935 and 31 Dec. 1935, leg. Guy Babault
235
European Journal of Taxonomy 382: 1–327 (2017)
(MCZ: 46157); 1 ad? ♀, 1 ? [1 AL, 1 UN], Loondo Mugunga, between 1 Mar. 1945 and 31 Mar. 1945,
leg. l’ Expédition au Lac Tumba (Congo Belge), between 1 Mar. 1954 and 31 Mar. 1954, leg. Gonze de
Loneux (RBINS: 7468.1, 7468.2); 1 ad ♂ [AL], Lovoi, between 1 Mar. 1932 and 31 May 1932, leg.
Quarré (RMCA: 11737); 1 juv ♂ [AL], Luanza Mission, prior to 13 Mar. 1913, leg. de Paoli (RMCA:
1541); 1 ad ♂, 1 juv ♀ [2 AL], Lubwe, 28 Nov. 1955, leg. M. Lips (RMCA: 26330, 26331); 2 ad ♀♀,
7 ad ♂♂, 1 sad ♂, 1 juv ♀, 1 juv ♂ [9 SS, 2 SK, 1 SC], Luebo, 16 Dec. 1921, 17 Dec. 1921, 19 Dec. 1921,
leg. Henri Schouteden (RMCA: 6609, 6635 to 6638, 6610, 6639, 6640, 6641a, 6641b, 6642, 6643); 1 ad
♀, 1 sad ♀ [1 SS, 1 SC], Lukafu, 20 Dec. 1930, leg. Gaston-François de Witte and de Crawhez (RMCA:
11129, 11130); 1 ad ♂ [SO], Lukolela, between 1 Sep. 1923 and 30 Sep. 1923, leg. J. Ghesquière
(RMCA: 7039); 2 ad ♀♀, 1 ad ♂, 2 ad? ♀♀, 3 ad? ♂♂, 1 sad ♀, 1 sad ♂, 1 ? [11 AL], Lula, 6 Aug. 1956,
10 Aug. 1956, leg. Max Poll, prior to 4 Jul. 1958, leg. A.J. Jobaert (RMCA: 25632 to 25638, 25629 to
25631, 27200); 1 imm ♀ [SS], Lulenga [= Rugari], 24 Nov. 1925, leg. Henri Schouteden (RMCA:
9151); 6 ad ♀♀, 2 ad ♂♂, 7 ad ??, 2 ad? ♀♀, 1 ad? ♂, 2 ad? ??, 1 sad ♀, 1 sad ♂, 3 juv ??, 2 imm ??,
1 ♀, 1 ? [14 SS, 13 AL, 2 SO], Luluabourg [= Kananga], unknown date, 5 Apr. 1922, 15 Apr. 1922,
16 Apr. 1922, 6 May 1922, 11 May 1922, 16 May 1922, prior to 8 Jul. 1913, between 11 May 1922 and
22 May 1922, between 11 May 1922 and 15 May 1922, leg. Richard Callewaert, unknown date, leg. Jan
Deheegher, 24 Sep. 1963, 24 May 1964, leg. Antoon Emeric Marcel De Roo (BMNH: 1927.12.21.11,
RMCA: 31567 to 31569, 31571, 31574, 31575, 7556, 7564, 7557 to 7560, 7555, 7561, 7554, 7562,
32657, 32666, 7553, 7552, 7563, 7565, 1954, 1954cc, 1954d to 1954g); 1 ad ♂, 7 ad ?? [5 SS, 2 SO,
1 SC], Lusambo, unknown date, 25 Dec. 1921, between 1 Dec. 1921 and 31 Dec. 1921, between
1 Jan. 1922 and 31 Jan. 1922, leg. J. Ghesquière (RMCA: 7339, 7340, 7335, 7631, 7334, 7336 to 7338);
1 ad ♀ [AL], Lusinga, 12 Jun. 1947, leg. Gaston-François de Witte et al. (RBINS: 10805); 2 ad ♀♀,
1 sad ♂ [1 SS, 2 AL], Lwiro [= Luiro], 21 Feb. 1959, leg. Uwe Rahm, 8 Aug. 1964, leg. U. Goepel
(RMCA: 28028, ZMZ: 126637, 126638); 1 ad ♂, 1 ♀, 2 ♂♂ [4 AL], Mabali, 29 Oct. 1958, leg. Uwe
Rahm, between 1 Jun. 1956 and 30 Jun. 1956, leg. Alvin Novick (RMCA: 28918, MCZ: 48258 to
48260); 10 ?? [10 UN], Mabwe, 10 Aug. 1949, leg. unknown collector (RBINS: 10806.02, 10806.05,
10806.06, 10806.10 to 10806.16); 5 ad ♀♀, 1 ad ♂ [6 AL], Mabwe River, 10 Aug. 1949, leg. GastonFrançois de Witte et al. (RBINS: 10806.01, 10806.03, 10806.04, 10806.07 to 10806.09); 1 ad ♀, 4 juv
♀♀, 1 juv ? [6 AL], Madimao, 18 May 1955, between 18 May 1955 and 18 Oct. 1955, leg. M. Lips
(RMCA: 26351 to 26353, 26333, 26354, 26355); 1 ad ♂ [AL], Makisabu [= Makisabo], 8 Jul. 1953,
leg. Gaston-François de Witte (RMCA: 37174); 1 ad ♀ [AL], Matadi, prior to 4 Aug. 1937, leg.
Dartevelle (RMCA: 13966); 1 juv ♂ [AL], Matale, prior to 27 Feb. 1940, leg. Hautmann (RMCA:
15885); 3 ad ♀♀, 2 ad ??, 4 juv ♀♀, 1 juv ♂ [7 SS, 3 SO], Mauda, 26 Feb. 1925, 16 Mar. 1925,
18 Mar. 1925, 21 Mar. 1925, between 2 Mar. 1925 and 3 Mar. 1925, leg. Henri Schouteden (RMCA:
8969 to 8971, 8978, 8979, 8989, 8990, 8955, 9902, 8958); 1 ad ♂, 1 juv ♂ [2 AL], May ya Moto,
5 Apr. 1954, between 15 Apr. 1955 and 15 Apr. 1956, leg. Gaston-François de Witte (RMCA: 37281,
37375); 1 ad ♂, 1 ♂ [1 AL, 1 UN], Mayumbe, unknown date, leg. unknown collector, unknown date,
leg. unknown collector (RMCA: 89.004-M-0011, 90.012-M-0002); 1 ad ♂ [SA], Mbalaka (Plantation),
23 Jul. 1995, leg. Herwig Leirs et al. (RMCA: 97.021-M-0838); 1 ♂, 5 ?? [1 SS, 5 UN], Medje,
26 Mar. 1910, 16 Sep. 1910, 7 Oct. 1910, 26 Mar. 1913, leg. Herbert Lang, James Paul Chapin and The
American Museum Congo Expedition (AMNH: 49308, 49304, 49314, 49338, 49291, 48966); 1 ad ♀
[SS], M’gayu, 14 Dec. 1909, leg. Herbert Lang, James Paul Chapin and The American Museum Congo
Expedition (RMCA: 12368); 1 ad? ♂, 1 sad ♀ [2 AL], Mimbulu, 4 Jan. 1956, leg. D’Haenens (RMCA:
26382, 26383); 1 ad ♂ [SA], Moera, 12 May 1921, leg. Nils Carl Gustaf Fersen Gyldenstolpe and Swiss
Mission (RMCA: 16285); 1 ad ♀ [AL], Mosenge, 15 Aug. 1986, leg. WHO and Marc Colyn (RMCA:
88.011-M-0021); 5 ad ♀♀, 7 ad ♂♂, 1 ad ?, 1 ad? ♂, 1 juv ♂, 1 imm ♂, 3 imm ??, 2 ?? [14 SS, 7 AL],
Mount Wago [= Wago, Mount], between 1 Dec. 1950 and 31 Dec. 1950, between 1 Jul. 1951 and
31 Jul. 1951, between 1 Jan. 1951 and 31 Dec. 1951, leg. Alexandre Fain, prior to 31 Dec. 1990, leg.
unknown collector (RMCA: 20791 to 20801, 23301, 21469 to 21475, RBINS: 21658, 21659); 2 ad ♀♀,
3 ad ♂♂ [5 AL], M’Pala, Kazenze, 22 Nov. 1949, leg. R.P. Lefébure (RBINS: 14524 to 14528); 6 ♂♂
236
VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
[2 SB, 4 SN], Mugeri Mission, 25 Jul. 2002, leg. Michael H. Huhndorf (FMNH: 173297 to 173302);
2 ad ♀♀, 1 ad ♂, 1 ad? ♀, 1 sad ♀ [4 SS, 1 SO], Mugunga, Lake, prior to 22 Dec. 1943, leg. GastonFrançois de Witte, prior to 31 Dec. 1943, leg. Gaston-François de Witte et al. (RMCA: 16635 to 16637,
RBINS: 4761, 4762); 2 ad ♀♀, 2 ad ♂♂, 1 ad? ♀, 3 juv ?? [8 AL], Mukuta [= Mututa], 16 Oct. 1955,
leg. M. Lips (RMCA: 26394 to 26401); 11 ♀♀, 13 ♂♂, 6 ?? [30 UN], Mulongo, between 28 Oct. 1974
and 29 Oct. 1974, leg. Michael D. Gallagher (HZM: 223.7990 to 252.8019); 1 ad ♀, 1 ad ♂ [2 AL],
Mulungu, prior to 1 Feb. 1940, leg. F.L. Hendricks (RMCA: 15873, 15874); 1 imm ? [SO], Mulungu
Kivu, between 1 Jun. 1935 and 30 Jun. 1935, leg. J.V. Leroy (RMCA: 13095); 5 ad ♀♀, 6 ad ♂♂, 17 ??
[1 SS, 10 AL, 17 UN], Murambi, unknown date, leg. unknown collector, 27 Aug. 1953, 21 Sep. 1953,
30 Sep. 1953, leg. Gaston-François de Witte, 27 Mar. 1959, leg. Uwe Rahm (INPBC: W1193, W1321,
W1516.01 to W1516.10, W604, W606, W608, W642, W644, RMCA: 37181, 37182, 28029, 37175 to
37180, 37199, 37200); 2 ad ♀♀ [2 AL], Musosa, near, between 1 Jan. 1941 and 31 Dec. 1941, leg.
H.J. Bredo (RBINS: 14532, 14533); 7 ?? [7 UN], Musugeriza River, unknown date, leg. unknown
collector (INPBC: W785, W919.1 to W919.6); 3 ?? [3 UN], Mutsora, unknown date, leg. unknown
collector (INPBC: W333.1 to W333.3); 1 ad ♀ [SK], Mutwanga, between 1 Aug. 1937 and 31 Aug. 1937,
leg. Henri Martin Hackars (RMCA: 16600); 4 ad ♀♀, 11 ad ♂♂, 1 ad? ♂, 2 ♀♀, 3 ♂♂ [16 AL, 5 UN],
Mwenda-Katuka Piste, 4 Jan. 1946, leg. J. de Wilde, 4 Jan. 1946, leg. unknown collector (RBINS:
15452.01 to 15452.21); 1 ad ♂ [AL], Nakiluba [= Inakiluba], 24 Oct. 1955, leg. M. Lips (RMCA:
26329); 1 ad ♀, 1 ad ♂, 1 juv ♂ [3 AL], Nakilubo [= Inakiluba], 14 Nov. 1955, leg. M. Lips (RMCA:
26368 to 26370); 2 ad ♀♀ [2 SK], Netonna, between 1 Jan. 1889 and 31 Dec. 1889, leg. P. Hesse (SMF:
2524 [lectotype Vesperugo pagenstecheri Noack, 1889], 2525 [paralectotype Vesperugo pagenstecheri
Noack, 1889]); 1 juv ♀, 1 ♂, 1 ? [2 SS, 1 SA], N’Gayu area, 14 Dec. 1909, leg. Herbert Lang, James
Paul Chapin and The American Museum Congo Expedition (AMNH: 48942, 48953, 49343); 1 juv ♀
[SS], N’goma [= Ngoma], between 1 Jan. 1925 and 31 Jan. 1925, leg. Henri Schouteden (RMCA:
9903); 3 ad ♀♀, 3 ad ♂♂, 2 sad ♀♀, 1 juv ♀ [9 SS], N’gombe [= Ngombe], 9 Nov. 1921, 12 Nov. 1921,
leg. Henri Schouteden (RMCA: 6629, 6630, 6611, 6628, 6631 to 6633, 7311, 7312); 6 ♀♀, 3 ♂♂ [8 SS,
1 SO], Niangara, 11 Nov. 1910, 14 Nov. 1910, leg. Herbert Lang, James Paul Chapin and The American
Museum Congo Expedition (AMNH: 48967 to 48975); 11 ad ♀♀, 10 ad ♂♂, 1 sad ♀, 1 sad ♂, 3 juv
♀♀, 2 juv ♂♂, 41 ?? [28 AL, 41 UN], Nyabikere, unknown date, leg. unknown collector, 1 Nov. 1953,
1 Feb. 1954, 18 Aug. 1954, 7 Feb. 1955, 30 Aug. 1955, leg. Gaston-François de Witte (INPBC: W1246,
W1334, W1375, W1532.01 to W1532.35, W620, W622, RMCA: 37225 to 37234, 37187, 37188, 37312
to 37325, 37364, 37361, 37362); 7 ?? [1 AL, 6 UN], Nyabitembe River, unknown date, leg. unknown
collector, 9 Apr. 1954, leg. Gaston-François de Witte (INPBC: W856.1 to W856.6, RMCA: 37298); 2 ad
♀♀, 1 ad ♂, 2 ?? [5 AL], Nyakariba River, 20 Jan. 1954, 10 Feb. 1954, leg. Gaston-François de Witte
(RMCA: 37235 to 37239); 2 ad ♂♂ [2 SO], Nzulu, prior to 31 Dec. 1943, leg. Gaston-François de Witte
et al. (RBINS: 4763, 4764); 1 ad ♂ [SO], N’Zulu [= Nzulu], prior to 22 Dec. 1943, leg. Gaston-François
de Witte (RMCA: 16638); 1 ad? ♀ [AL], Omaniumdu, 27 Jul. 1959, leg. Laurent (RMCA: 29537);
1 ad ? [SO], Pili Pili [= Pilipili], 16 Dec. 1912, leg. Cuthbert Christy (RMCA: 1610); 1 ? [UN], Poko,
23 Jul. 1914, leg. Cuthbert Christy (RMCA: 4797); 1 ? [SA], Poko Zone [= Poko Territory],
30 Aug. 1913, leg. Herbert Lang, James Paul Chapin and The American Museum Congo Expedition
(AMNH: 49329); 1 juv ♀ [AL], Popokabaka, between 1 Nov. 1955 and 30 Nov. 1955, leg. René Close
(RMCA: 90.009-M-0001); 1 ? [AL], Risimu Zone, 9 Sep. 1909, leg. Herbert Lang, James Paul Chapin
and The American Museum Congo Expedition (AMNH: 49346); 3 ad ♀♀, 4 ad ♂♂, 1 ad? ♀, 1 juv ♀,
12 ?? [9 AL, 12 UN], Rodahira River, unknown date, leg. unknown collector, 28 Oct. 1953, 16 Jul. 1954,
5 Aug. 1955, leg. Gaston-François de Witte (INPBC: W1195, W1197.1 to W1197.8, W612, W614,
W875.2, RMCA: 37330 to 37335, 37185, 37365, 37366); 7 ?? [7 UN], Rumangabo, unknown date, leg.
unknown collector (INPBC: V1461.2.1 to V1461.2.6, V1461.27); 2 ad ♀♀, 1 ad ♂, 1 juv ♀, 1 imm ♀,
1 imm ♂, 7 ?? [6 AL, 7 UN], Rumanura River, unknown date, leg. unknown collector, 9 Feb. 1954,
30 Apr. 1954, leg. Gaston-François de Witte (INPBC: W939.01 to W939.07, RMCA: 37307 to 37311,
37264); 3 ad ♀♀, 5 ad ♂♂, 1 imm ♂ [9 AL], Rusenyi River, 22 Sep. 1953, 10 Mar. 1954, leg. Gaston237
European Journal of Taxonomy 382: 1–327 (2017)
François de Witte (RMCA: 37271 to 37277, 37184, 37186); 2 ad ♀♀, 14 ad ♂♂, 1 ad? ♀, 1 sad ♀,
1 imm ♀, 4 ♀♀, 411 ?? [1 SS, 25 AL, 1 SO, 407 UN], Rutshuru, unknown date, leg. unknown collector,
16 Sep. 1953, 20 Jan. 1954, 9 Feb. 1954, 22 Apr. 1954, 5 Dec. 1955, leg. Gaston-François de Witte,
9 Aug. 1925, 30 Aug. 1925, leg. Edmund Heller, 31 Mar. 1938, leg. Serge Frechkop, between 6 Jan. 1931
and 6 Jan. 1938, leg. J. Ghesquière (INPBC: V121, V1248, V1348, V1349, V1350.2, V1352.1, V1352.2,
V1353.1, V1355, V1357, V1358, V1366, V1368 to V1370, V1372 to V1374, V1376 to V1380, V1382.1,
V1383 to V1385, V1386.1, V1388, V1390 to V1396, V1398, V1400, V1401, V1404, V1405, V1420.1,
V1421, V1424, V1425, V1428, V1522.1, V1527, V1528, V1530, V1531, V1594.001 to V1594.345,
V371, V377, W1164, W1525, W626, W628, W630, W739, W775, W989.01, W989.02, RMCA: 37191,
37192, 37240 to 37247, 37303 to 37306, FMNH: 30669, 30670, RMCA: 17090, 37373, 37374, 37265
to 37269, FMNH: 26499, 30671, RMCA: 14445); 2 ad ♂♂, 1 juv ♀, 1 ? [3 SS, 1 UN], Ruwenzori,
unknown date, leg. Rudolf Grauer and Hermann Rolle, between 22 Jan. 1906 and 31 Dec. 1911, between
21 Jan. 1906 and 31 Dec. 1911, between 20 Jan. 1906 and 31 Dec. 1911, leg. Rosenberg (ZMB: T09311,
RMCA: 937 to 939); 1 ad ♂ [AL], Sakania, prior to 31 Dec. 1913, leg. Coulon (RMCA: 2821); 3 ad
♀♀, 1 ad ♂ [4 AL], Shindaika, 27 Sep. 1955, leg. M. Lips (RMCA: 26356 to 26359); 1 ♂ [SS],
St Joseph Mission [= Mikalaya], unknown date, leg. Richard Callewaert (BMNH: 1934.10.24.19);
6 ♀♀, 7 ♂♂, 1 ? [12 SS, 2 UN], St. Jos. (Mission) [= Mikalaya], 14 Apr. 1924, 15 Apr. 1924,
18 Apr. 1924, 5 May 1924, 8 May 1924, 10 May 1924, 8 Jul. 1924, leg. Richard Callewaert (AMNH:
86163, 86164, 117649, 86153 to 86162, 86142); 2 ad ♀♀, 1 ad ♂, 1 ad ? [2 SK, 2 SO], St. Joseph de
Luluabourg Mission [= Mikalaya], 2 May 1939, 7 May 1939, leg. Jean Jacques Deheyn, 27 Jun. 1923,
between 15 Feb. 1923 and 15 Mar. 1923, leg. Richard Callewaert (RMCA: 15373, 7182, 15379, 7169);
1 ad ♀, 3 ad ♂♂, 1 ad? ♀, 1 ad? ♂, 1 juv ♀, 1 juv ♂, 2 juv ??, 1 ♀, 4 ♂♂, 1 ? [4 SS, 2 SA, 10 AL],
Stanleyville [= Kisangani], 1 Mar. 1928, leg. Collart, 21 Jun. 1931, leg. Schwetz, 25 Aug. 1909,
5 Sep. 1909, 9 Sep. 1909, leg. Herbert Lang, James Paul Chapin and The American Museum Congo
Expedition, prior to 6 Oct. 1910, prior to 31 Jul. 1899, leg. Weyns (RMCA: 16262, 11707, AMNH:
48938 to 48940, 49342, 48937, 49325, RMCA: 282, 282a to 282f, 894); 8 ad ♀♀, 3 ad ♂♂, 5 ♀♀, 8 ♂♂
[2 SS, 22 AL], Tandala, 7 Jul. 1979, 9 Jul. 1979, 10 Jul. 1979, 11 Jul. 1979, 12 Jul. 1979, 13 Jul. 1979,
leg. Lynn W. Robbins and C. Brian Robbins, 9 Jul. 1979, 10 Jul. 1979, 11 Jul. 1979, 12 Jul. 1979,
13 Jul. 1979, leg. unknown collector, 7 Jul. 1979, 11 Jul. 1979, 12 Jul. 1979, 13 Jul. 1979, leg. C. Brian
Robbins (CM: 86797 to 86799, MHNG: 1870.083, CM: 86800, MHNG: 1870.084, USNM: 537723,
CM: 86802, MHNG: 1870.085, USNM: 463502, CM: 86803, 86804, MHNG: 1870.086 to 1870.088,
USNM: 463503, CM: 86794, USNM: 463498 to 463501, 537722, CM: 86796, MHNG: 1870.082); 1 ad
♀, 1 ad ♂ [2 SS], Teturi, 24 May 1948, 17 Sep. 1948, leg. Jean Hiernaux (RBINS: 7051, 7050); 4 ad
♀♀, 2 ad ♂♂, 1 juv ♀ [3 SS, 2 SO, 2 SC], Tondu, 24 Feb. 1921, between 24 Feb. 1921 and 27 Feb. 1921,
leg. Henri Schouteden (RMCA: 6612 to 6617, 6621); 2 ad ♀♀ [2 AL], Tshakala, between 1 Jan. 1982
and 31 Jan. 1982, leg. Marc Colyn (RMCA: 82.019-M-0005, 82.019-M-0006); 1 ad ♂ [AL], Tshibaka
[= Tshibaka-Kaji], 25 Aug. 1956, leg. Max Poll (RMCA: 25628); 1 ad ♂, 1 ad? ♂ [2 SS], Tshikapa,
28 Nov. 1921, between 16 Nov. 1921 and 26 Nov. 1921, leg. Henri Schouteden (RMCA: 6634, 7310); 1
ad ♀, 1 ad ♂, 1 juv ? [3 AL], Tshisangwe, 27 Sep. 1955, leg. M. Lips (RMCA: 26371 to 26373); 4 ♀♀,
2 ♂♂ [6 SS], Upper Congo, unknown date, leg. Alexander Douglas Mitchell Carruthers (BMNH:
1907.6.14.10, 1907.6.14.19, 1907.6.14.21, 1907.6.14.7 to 1907.6.14.9); 1 juv ♀, 1 ? [1 AL, 1 UN],
Uvira, unknown date, leg. Rudolf Grauer, between 1 Mar. 1955 and 31 Mar. 1955, leg. Narcisse Leleup
(ZMB: T09307, RMCA: 22913); 1 ad ♀, 1 ♀ [2 SS], Vitshumbi, prior to 31 Dec. 1943, leg. unknown
collector, prior to 31 Dec. 1943, leg. Gaston-François de Witte et al. (RBINS: 4765, 4766); 6 ad ♂♂,
97 ?? [101 AL, 2 UN], Wafanya [= Wafania], unknown date, 28 Nov. 1987, 3 Jan. 1988, 5 Dec. 1988,
2 Sep. 1991, 4 Sep. 1991, 14 Sep. 1991, 17 Sep. 1991, 20 Sep. 1991, 23 Sep. 1991, 25 Sep. 1991,
7 Oct. 1991, 12 Oct. 1991, 14 Oct. 1991, 28 Apr. 1992, 29 Apr. 1992, 19 May 1992, 26 May 1992,
29 May 1992, between 12 Oct. 1991 and 18 Oct. 1991, between 1 Dec. 1991 and 31 Dec. 1991, between
1 Nov. 1991 and 30 Nov. 1991, between 1 Oct. 1987 and 31 Oct. 1987, between 1 Aug. 1991 and
31 Aug. 1991, between 1 Jul. 1991 and 31 Jul. 1991, between 1 May 1992 and 31 May 1992, between
238
VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
1 May 1991 and 31 May 1991, between 1 Apr. 1992 and 30 Apr. 1992, between 1 Mar. 1992 and
31 Mar. 1992, between 1 Mar. 1991 and 31 Mar. 1991, between 1 Jan. 1992 and 31 Jan. 1992, between
1 Jan. 1991 and 31 Jan. 1991, leg. P. Lootens (RMCA: 92.079-M-0012, 92.079-M-0127 to 92.079-M0139, 92.079-M-0153, 92.079-M-0154, 91.076-M-0108, 91.076-M-0084, 91.076-M-0085, 91.076-M0112, 91.076-M-0087, 92.079-M-0084, 92.079-M-0093, 91.076-M-0066, 91.076-M-0088, 91.076-M0089, 91.076-M-0091, 91.076-M-0093 to 91.076-M-0097, 92.079-M-0097 to 92.079-M-0100,
92.079-M-0045, 92.079-M-0046, 92.079-M-0048, 92.079-M-0054, 92.079-M-0055, 92.079-M-0101,
92.079-M-0102, 88.005-M-0004, 92.079-M-0052, 92.079-M-0044, 92.079-M-0047, 92.079-M-0050,
92.079-M-0051, 88.005-M-0005, 88.005-M-0006, 88.005-M-0013, 91.076-M-0068 to 91.076-M-0071,
89.020-M-0047, 91.076-M-0105, 91.076-M-0001, 92.079-M-0053, 92.079-M-0109, 91.076-M-0015,
92.079-M-0004, 92.079-M-0005, 92.079-M-0007, 92.079-M-0011, 92.079-M-0081, 92.079-M-0087 to
92.079-M-0089, 92.079-M-0091, 92.079-M-0092, 92.079-M-0094, 92.079-M-0071, 92.079-M-0085,
92.079-M-0086, 92.079-M-0095, 92.079-M-0096, 92.079-M-0106, 92.079-M-0107, 91.076-M-0030,
91.076-M-0042, 91.076-M-0043, 91.076-M-0049, 91.076-M-0050, 92.079-M-0080, 92.079-M-0082,
92.079-M-0083, 92.079-M-0090, 91.076-M-0053, 91.076-M-0054, 91.076-M-0062, 88.005-M-0015,
92.079-M-0008, 92.079-M-0006, 92.079-M-0009, 92.079-M-0010, 92.079-M-0013, 92.079-M-0049,
92.079-M-0056, 92.079-M-0105, 92.079-M-0108, 91.076-M-0103); 2 ad ♀♀, 2 ad ♂♂ [4 AL], Wilibadi
River, 16 Apr. 1948, leg. Hediger and Jacques Verschuren (RBINS: 8195.1 to 8195.4); 1 ? [SA],
Yakuluku, 4 Nov. 1912, leg. Herbert Lang, James Paul Chapin and The American Museum Congo
Expedition (AMNH: 49344); 1 ad ♀, 2 ad ♂♂, 3 ♀♀, 5 ♂♂ [11 AL], Yalosemba, 10 Jun. 1979,
17 Jun. 1979, 23 Jun. 1979, leg. Lynn W. Robbins and C. Brian Robbins, 15 Jun. 1979, leg. Frieder
Meier, 19 Jun. 1979, 22 Jun. 1979, 23 Jun. 1979, leg. C. Brian Robbins, 23 Jun. 1979, leg. unknown
collector (CM: 86790, MHNG: 1870.078, CM: 86791, USNM: 463488 to 463490, 463495, CM: 86792,
MHNG: 1870.079, USNM: 463496, 463497); 1 ad ♀, 1 ad ♂ [2 AL], Yalusaka, 28 Sep. 1957,
16 Mar. 1958, leg. F. Carbone (RMCA: 27164, 27165); 3 ♀♀, 2 ♂♂ [5 AL], Yamazo Yamono,
19 Jun. 1979, leg. Frieder Meier, 19 Jun. 1979, leg. C. Brian Robbins (MHNG: 1870.080, USNM:
463491 to 463494); 2 ad ♂♂, 1 ♂ [2 SS, 1 SC], Yangambi, 29 Jul. 1946, 8 Aug. 1946, leg. G. Gilbert,
8 Aug. 1946, leg. unknown collector (RBINS: 6989 to 6991); 1 ad ♂ [SO], Yangambi Division Forest,
8 Aug. 1946, leg. G. Gilbert (RMCA: 37900).
RWANDA: 2 ad ♀♀, 1 ad ♂, 1 juv ♀, 1 juv ♂, 2 ♀♀, 2 ♂♂, 1 ? [2 SS, 7 UN, 1 AS], Astrida [= Butare],
11 Nov. 1966, leg. Félix / Michel Anciaux de Faveaux, 13 Oct. 1976, leg. Frieder Meier, 24 Feb. 1966,
leg. unknown collector (RBINS: 16001, 16003, MHNG: 1873.084, RBINS: 15488.1, 15488.2, 15489.1,
15489.2, 15491.1 to 15491.3); 1 ad ♂ [SA], Butare, 7 Oct. 1981, leg. Frits De Vree, Benny Baeten
and Victor Van Cakenberghe (RMCA: 82.006-M-0201); 16 ?? [16 UN], Gabiro, unknown date, leg.
unknown collector (INPBC: V3102, V3129.01 to V3129.15); 5 ad ?? [5 SS], Gafumba, 24 Nov. 1966,
leg. Félix / Michel Anciaux de Faveaux (RBINS: 15996 to 15999, 16002); 1 ad ♂ [SA], Gafumba
Mugusa, 7 Feb. 1985, leg. Théoneste (RMCA: 87.047-M-0129); 58 ?? [58 UN], Gihinga, unknown
date, leg. unknown collector (INPBC: V3044.1.01 to V3044.1.58); 2 ♂♂ [2 AL], Gisenyi, 28 Jun. 1980,
13 Sep. 1980, leg. Hans H. de Iongh (ZMA: MAM.21232, MAM.21231); 1 ad ♀ [SA], Gisovu,
3 Oct. 1980, leg. Danny Meirte (RMCA: 80.055-M-0001); 3 ad ♀♀, 2 ad ♂♂, 1 juv ♂ [5 SA, 1 AL],
Kibuye, 6 Oct. 1980, 7 Oct. 1980, 8 Oct. 1980, leg. Danny Meirte (RMCA: 80.055-M-0002 to 80.055M-0004, 80.055-M-0006, 80.055-M-0005, 80.055-M-0007); 1 ad ♀ [SA], Kitabi, 5 Nov. 1981, leg.
Frits De Vree, Benny Baeten and Victor Van Cakenberghe (RMCA: 82.006-M-0202); 1 ad ♀ [SA],
Mabanza, 10 Oct. 1980, leg. Danny Meirte (RMCA: 80.055-M-0008); 69 ?? [69 UN], Mihindi, Lake,
unknown date, leg. unknown collector (INPBC: V3090, V3112.01 to V3112.68); 1 ad ? [SS], Mugogwe,
19 Dec. 1966, leg. Félix / Michel Anciaux de Faveaux (RBINS: 16000); 1 ♂ [UN], Nyamasheke,
8 Apr. 1966, leg. unknown collector (RBINS: 15486); 24 ad ♀♀, 49 ad ♂♂, 2 sad ♀♀, 3 juv ♀♀,
3 juv ♂♂, 2 imm ?? [66 SA, 16 AL, 1 UN], Nyamata, 15 Nov. 1981, 16 Nov. 1981, 17 Nov. 1981, leg.
Frits De Vree, Benny Baeten and Victor Van Cakenberghe (RMCA: 82.006-M-0203 to 82.006-M-0207,
239
European Journal of Taxonomy 382: 1–327 (2017)
82.006-M-0218 to 82.006-M-0267, 82.006-M-0208 to 82.006-M-0217, 82.006-M-0268 to 82.006-M0285); 1 ad ♀ [SA], Nyambumba, between 18 Dec. 1984 and 5 Jan. 1985, leg. Benny Baeten and Luc
Janssens (RMCA: 85.006-M-0095); 1 ad ♂ [SA], Nyundo, 19 Oct. 1980, leg. Danny Meirte (RMCA:
80.055-M-0009); 3 ad ♂♂, 1 ♂ [3 AL, 1 UN], Rubona, 20 Jan. 1966, 23 Jan. 1967, leg. Félix / Michel
Anciaux de Faveaux (RBINS: 15487, 15934.1 to 15934.3); 1 ad ♂ [SA], Shangi, between 18 Dec. 1984
and 5 Jan. 1985, leg. Benny Baeten and Luc Janssens (RMCA: 85.006-M-0096).
Neoromicia rendalli (Thomas, 1889)
DEMOCRATIC REPUBLIC OF THE CONGO: 6 ad ♀♀, 2 ad ♂♂, 2 sad ♀♀ [10 AL], unknown locality,
27 Jul. 1965, prior to 27 Jul. 1965, between 3 Jul. 1965 and 27 Jul. 1965, leg. Antoon Emeric Marcel
De Roo (RMCA: 33547 to 33551, 33543 to 33546, 33552); 1 sad ♀ [AL], Albertville [= Kalemie],
2 Apr. 1960, leg. G. Verhoustraete (RMCA: 28666); 1 ad ♀, 1 juv ♀, 1 juv ♂, 3 ?? [3 AL, 3 UN],
Bagbele, unknown date, leg. unknown collector, 30 Apr. 1950, leg. H. de Saeger (INPBC: 1182.1 to
1182.3, RMCA: 36848 to 36850); 1 ad ♂ [AL], Bagbele Moko, 25 Jul. 1952, leg. H. de Saeger (RMCA:
36851); 3 ad ♀♀, 2 ad ♂♂ [5 SS], Bolobo, prior to 23 Jul. 1946, leg. N’Kele (RMCA: 17404 to 17408);
1 ad ♂ [SS], Cel II, 8 Mar. 1952, leg. H. de Saeger et al. (RBINS: 13850); 1 ♀ [SA], Duma, between
1 Oct. 1910 and 31 Oct. 1910, leg. unknown collector (SMF: 6393); 3 ad ♀♀, 2 ad ♂♂, 1 juv ♀, 1 juv ♂,
5 ♀♀, 5 ♂♂ [11 SS, 2 SA, 2 SO, 2 UN], Faradje, 18 Feb. 1911, 21 Feb. 1911, 1 Mar. 1911, 2 Mar. 1911,
29 Mar. 1911, 30 Mar. 1911, 2 Aug. 1911, leg. Herbert Lang, James Paul Chapin and The American
Museum Congo Expedition, 2 Aug. 1911, leg. unknown collector (AMNH: 49047, 49048 [topotype
Eptesicus faradjius J.A. Allen, 1917], 49044 [topotype Eptesicus faradjius J.A. Allen, 1917], 52114,
52115, 49054 [topotype Eptesicus faradjius J.A. Allen, 1917], 49055 [topotype Eptesicus faradjius
J.A. Allen, 1917], MCZ: 17379, AMNH: 49045 [holotype Eptesicus faradjius J.A. Allen, 1917],
49046, 49053 [topotype Eptesicus faradjius J.A. Allen, 1917], 49049 [topotype Eptesicus faradjius
J.A. Allen, 1917], 49050, 49051 [topotype Eptesicus faradjius J.A. Allen, 1917], 49327, 49345,
RMCA: 12387); 1 ad ♂ [AL], Gangala-na-Bodio, between 1 Nov. 1956 and 30 Nov. 1956, leg. Max
Poll (RMCA: 25807); 7 ad ♀♀, 4 ad ♂♂, 1 ad? ♂, 1 juv ♀, 2 ♀♀, 6 ♂♂ [3 SS, 18 AL], Kabwe,
9 Feb. 1964, 9 Feb. 1965, 30 Apr. 1965, leg. Antoon Emeric Marcel De Roo (RMCA: 33352 to 33359,
AMNH: 212581 to 212590, RMCA: 33596 to 33598); 1 ad ♂ [SS], Katobwe, 9 Dec. 1926, leg. John
Todd Zimmer (FMNH: 29417); 1 ad ?, 1 ? [2 SS], Kawa (Forest), 24 Jul. 1951, leg. unknown collector,
24 Jul. 1952, leg. Alexandre Fain (RBINS: 21661, RMCA: 23153); 9 ad ♀♀, 6 ad ♂♂, 3 ad ??, 2 juv
♀♀, 1 juv ?, 5 ♀♀ [17 SS, 4 AL, 2 SK, 2 SO, 1 SC], Luluabourg [= Kananga], unknown date,
13 Apr. 1922, 15 May 1922, 28 May 1922, 29 May 1922, 7 Jun. 1922, 11 Jun. 1922, 19 Jun. 1922,
26 Jun. 1922, 17 Jul. 1922, 28 May 1924, between 16 Jun. 1922 and 19 Jun. 1922, leg. Richard
Callewaert, 24 Sep. 1963, 3 Jul. 1965, between 26 Apr. 1964 and 18 Aug. 1965, leg. Antoon Emeric
Marcel De Roo, between 1 Jan. 1963 and 31 Dec. 1963, leg. Jan Deheegher (BMNH: 1926.11.1.54,
1926.11.1.55, 1926.7.6.29, 1926.7.6.77, 1926.7.6.78, RMCA: 7573, 7575, 7576, 7578, 7586, 7572,
7582, 7579, 32668, 7574, 7583, 7571, 7577, 7580, AMNH: 86166, RMCA: 7581, 33542, 7585, 31572,
7584, 33605); 1 ad ♂ [SA], Mulongo, 31 Oct. 1974, leg. Michael D. Gallagher (HZM: 1.8024); 3 ad
♀♀, 1 ad ♂, 5 ♀♀, 4 ♂♂ [13 SS], Niangara, 16 Nov. 1910, 26 Nov. 1910, 12 Dec. 1910, 16 Dec. 1910,
26 Dec. 1910, leg. Herbert Lang, James Paul Chapin and The American Museum Congo Expedition,
16 Dec. 1910, leg. unknown collector (AMNH: 49039, 49040, FMNH: 43870, AMNH: 49034, FMNH:
43869, RMCA: 12388, AMNH: 49041, MCZ: 17378, AMNH: 49036 to 49038, 49042, 49043); 1 ♀,
2 ♂♂ [2 SS, 1 UN], St. Jos. (Mission) [= Mikalaya], 20 May 1924, 28 May 1924, 18 Oct. 1924,
leg. Richard Callewaert (AMNH: 86146, 86165, 86167); 8 ad ♀♀, 3 ad ♂♂ [10 SS, 1 SA], Tandala,
7 Jul. 1979, 9 Jul. 1979, 13 Jul. 1979, 16 Jul. 1979, 20 Jul. 1979, leg. Lynn W. Robbins and C. Brian
Robbins (CM: 86675, 86668, 86669, 86782, 86676 to 86679, 86672 to 86674).
240
VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
Neoromicia somalica (Thomas, 1901)
DEMOCRATIC REPUBLIC OF THE CONGO: 1 ? [UN], unknown locality, between 1 Apr. 1999 and
30 Apr. 1999, leg. unknown collector (SMF: 93322); 1 ad ? [SA], Lukolela, 4 Aug. 1930, leg. James
Paul Chapin (AMNH: 86927); 1 ♂ [SS], Mboga [= Boga], unknown date, leg. Howard de Walden
(BMNH: 1930.11.11.172); 1 ad ♂ [SA], Mbwambala, 20 Jun. 1995, leg. Herwig Leirs et al. (RMCA:
97.021-M-0749).
RWANDA: 1 ad ♀, 1 ad ♂ [2 SK], Karama, 11 Oct. 1981, 12 Oct. 1981, leg. Frits De Vree, Benny
Baeten and Victor Van Cakenberghe (RMCA: 82.006-M-0918, 82.006-M-0919).
Neoromicia tenuipinnis (Peters, 1872)
BURUNDI: 3 ad ♂♂, 1 juv ♀, 6 ♂♂ [5 AL, 4 SB, 1 SN], Bujumbura, 24 Jan. 1970, 26 Jan. 1970,
17 Feb. 1970, 20 May 1970, 6 Feb. 1972, 24 Jan. 1973, 12 Feb. 1973, leg. P.L. Niort, between 1 Jan. 1928
and 31 Dec. 1928, leg. Douce (MHNG: 1608.019, 1607.096, 1608.005, 1608.006, 1608.011, 1608.018,
1624.003, 1895.089, 1895.090, RMCA: 16271); 1 ♀ [UN], Russissi, bank [= Rusizi Delta], unknown
date, leg. Rudolf Grauer (ZMB: 54067); 1 ♀ [SS], Usumbura [= Bujumbura], 14 Oct. 1938, leg.
Duncan M. Hodgson and William F. Coultas (MCZ: 38033).
DEMOCRATIC REPUBLIC OF THE CONGO: 1 ad ♀, 4 juv ♂♂, 1 imm ♂, 2 ♀♀, 2 ♂♂ [5 SS, 1 SA,
3 AL, 1 UN], unknown locality, unknown date, 1 Jun. 1939, leg. unknown collector, unknown date, leg.
Henri Schouteden, unknown date, leg. P. Hesse, 4 Jul. 1958, prior to 4 Jul. 1958, leg. E. Jans (BMNH:
1926.11.1.56, 1926.7.6.81 to 1926.7.6.83, RMCA: 9193, SMF: 2526, RBINS: 6277.2, RMCA: 27372,
27367, 27370); 1 ad ♀ [SS], Alima, 12 Feb. 1960, leg. P. Dyleff (RMCA: 28827); 2 ad ♀♀ [2 AL],
Bamanya, 20 Mar. 1956, leg. R.P. Hulsaert, between 1 Jan. 1952 and 31 Dec. 1952, leg. Leonet (RMCA:
26328, 21699); 1 ad ♂, 2 ad ??, 1 sad ?, 1 juv ? [1 SS, 4 SO], Bambesa, 24 Apr. 1940, between 1 Jun. 1937
and 30 Jun. 1937, leg. J.M. Vrijdagh (RMCA: 17753, 14075 to 14078); 1 juv ♀ [AL], Banana, between
1 May 1952 and 31 May 1952, leg. Is. Mesmaekers (RMCA: 21375); 1 ad ♂ [AL], Banana Creek,
20 Apr. 1949, leg. The MBizi expedition (RBINS: 14511); 1 juv ♀ [AL], Bena Tshiadi [= Bena Tshadi],
between 1 Apr. 1979 and 30 Apr. 1979, leg. Robertson (RMCA: 79.019-M-0001); 2 ad ♀♀, 1 ad? ♂,
1 juv ♀, 1 juv ♂, 1 imm ♂ [6 AL], Beno, between 1 Jan. 1949 and 31 Dec. 1949, leg. P. Bayet (RMCA:
19475, 19828 to 19830, 19849, 19850); 1 ? [UN], Biangolo, unknown date, leg. unknown collector
(INPBC: W53); 1 ad ♂ [SS], Bitshumbi [= Vitshumbi], prior to 22 Dec. 1943, leg. Gaston-François de
Witte (RMCA: 16632); 1 juv ♀ [AL], Bokoro, between 1 Jan. 1952 and 31 Dec. 1952, leg. E. Jans
(RMCA: 21523); 2 ad ♂♂, 1 juv ♂ [3 AL], Boteka, 26 Aug. 1984, between 1 Feb. 1985 and 28 Feb. 1985,
leg. P. Lootens (RMCA: 85.021-M-0001 to 85.021-M-0003); 1 juv ? [SO], Bukavu, between
16 Aug. 1952 and 14 Apr. 1955, leg. P.G. Vercammen (RMCA: 23173); 2 ad ♀♀, 3 ad ♂♂, 3 sad ♀♀,
3 juv ♀♀, 8 juv ♂♂, 1 imm ♀, 1 imm ♂, 1 imm ?, 1 ♂, 1 ? [22 AL, 2 UN], Buta, unknown date,
8 Jan. 1936, prior to 31 May 1939, prior to 16 Jun. 1939, prior to 16 Jun. 1937, between 1 Jul. 1949 and
31 Jul. 1949, between 1 Jan. 1935 and 31 Dec. 1935, between 1 Jan. 1930 and 31 Dec. 1930, between
1 Jan. 1925 and 31 Dec. 1925, leg. Jozef Hutsebaut, 10 Dec. 1946, leg. unknown collector (RMCA:
14564a to 14564c, 14584a to 14584c, 16186, 16280, 16281, RBINS: 14510, RMCA: 13709a to 13709c,
16245, 16188, 13176, 13177, 19459, 19460, 13859, 15397, 15325, 15327, 15328); 4 ad ♀♀, 1 ad ♂,
1 ad? ♀, 2 juv ♂♂ [8 AL], Butembo, 16 Mar. 1959, between 16 Mar. 1959 and 26 May 1959, leg.
P. Dyleff (RMCA: 27878 to 27882, 27884, 27877, 27883); 1 ad ♀, 1 ad? ♀ [1 SS, 1 AL], Eala, between
1 Aug. 1936 and 31 Aug. 1936, between 1 Jul. 1935 and 31 Jul. 1935, leg. J. Ghesquière (RMCA: 13096,
13410); 1 ♀ [SS], Edouard, Lac [= Edward, Lake], prior to 31 Dec. 1943, leg. unknown collector
(RBINS: 4760); 1 juv ♀, 1 ? [1 AL, 1 UN], Elisabetha, prior to 5 Oct. 1920, prior to 19 Apr. 1926, leg.
Tinant (RMCA: 8220, 5369); 1 ad ♂, 1 ad ?, 1 ? [2 SS, 1 AL], Faradje, 1 Mar. 1911, 12 Jan. 1913, leg.
Herbert Lang, James Paul Chapin and The American Museum Congo Expedition, 6 Jul. 1925, leg. Henri
241
European Journal of Taxonomy 382: 1–327 (2017)
Schouteden (AMNH: 48988 [holotype Eptesicus ater J.A. Allen, 1917], 49423 [topotype Eptesicus
ater J.A. Allen, 1917], RMCA: 9043); 1 ad ♂ [AL], Ganda-Sundi, 25 Mar. 1964, leg. Alexandre Fain
(RMCA: 33268); 1 ? [UN], Ibembo, between 1 Sep. 1949 and 30 Sep. 1949, leg. Jozef Hutsebaut
(RMCA: 19461); 1 ad ♀ [AL], Irumu, between 1 Jan. 1926 and 31 Dec. 1926, leg. Leopold prince of
Belgium (RMCA: 16256); 2 ad ♂♂ [2 AL], Kabale, 10 May 1927, 11 May 1927, leg. James Paul
Chapin, Dewitt L. Sage, Frank P. Mathews and The Ruwenzori-Kivu Expedition (AMNH: 82384,
82383); 1 ad? ♂ [AL], Kabongo, between 1 Jan. 1952 and 31 Dec. 1952, leg. M. Dierckx (RMCA:
21484); 1 ? [UN], Kalenda, 1 Aug. 1952, leg. unknown collector (RBINS: 7742); 1 ? [UN], Kalenda
[Kasai], unknown date, leg. unknown collector (RBINS: 1742); 1 ad ♀ [AL], Kamponde, between
1 Apr. 1965 and 30 Apr. 1965, leg. Antoon Emeric Marcel De Roo (RMCA: 33389); 1 ad ? [SS],
Katana, 29 Oct. 1932, leg. Burgeon (RMCA: 12057); 1 ad ♀ [UN], Katanga (former) Province,
1 Jun. 1939, leg. unknown collector (RBINS: 6277.1); 1 ad ♂ [UN], Kinzambi Mission, 30 Jul. 1995,
leg. Herwig Leirs et al. (MSB: 274830); 1 ad ♂ [AL], Kisangani, between 1 Jun. 1982 and 30 Jun. 1982,
leg. Marc Colyn (RMCA: 82.031-M-0035); 1 ad ♂, 1 sad ♂ [1 SS, 1 SO], Kisantu, 11 Dec. 1920,
12 Dec. 1920, leg. Henri Schouteden (RMCA: 7313, 7314); 1 ad ♂ [SS], Koteli [= Kotili], 23 Jan. 1925,
leg. Henri Schouteden (RMCA: 8922); 1 ad ♂ [SS], Kunungu, between 1 Mar. 1921 and 31 Mar. 1921,
leg. Henri Schouteden (RMCA: 7783); 1 ad ♀, 1 ad ? [2 SO], Léopoldville [= Kinshasa], between
1 Jan. 1949 and 31 Dec. 1949, leg. Richard, between 1 Nov. 1942 and 30 Nov. 1942, leg. R. Fiasse
(RMCA: 18809, 17661); 1 ad ♀, 3 ?? [2 AL, 2 UN], Leverville [= Lusanga], between 1 Feb. 1929 and
28 Feb. 1929, leg. Tinant (RMCA: 9987a, 9988a, 9988c, 9989); 1 ad ♂, 1 sad ♀, 1 ? [2 AL, 1 SO],
Lukolela, unknown date, leg. J. Ghesquière, 1 Aug. 1930, leg. James Paul Chapin, 25 Jun. 1931, leg.
Lecocq (RMCA: 16182, AMNH: 86914, RMCA: 11735); 2 ad ♀♀, 1 ad ♂, 1 sad ♀, 1 juv ♂, 1 ♀ [2 SS,
3 AL, 1 SC], Luluabourg [= Kananga], unknown date, 22 Jun. 1922, 4 Jul. 1922, leg. Richard
Callewaert, 24 Sep. 1963, 3 Jul. 1965, leg. Antoon Emeric Marcel De Roo (BMNH: 1927.12.21.12,
RMCA: 7587, 32667, 33540, 33541, 7569); 3 ad ♀♀, 1 ad ♂ [4 AL], Luluabourg Ravine, 29 Aug. 1963,
leg. Antoon Emeric Marcel De Roo (RMCA: 32773 to 32776); 1 ad ♀, 1 ad ♂, 6 ♀♀, 2 ♂♂ [1 SA,
9 AL], Mabali, 19 Dec. 1958, leg. Jean Georges Baer, 21 Jun. 1956, between 1 Jun. 1956 and 30 Jun. 1956,
leg. Alvin Novick, 9 Oct. 1958, leg. Uwe Rahm (MHNG: 985.067, MCZ: 48245, RMCA: 28912, 28913,
MCZ: 48246 to 48251); 1 ad ♀ [AL], Makaw, prior to 15 Nov. 1958, leg. E. Jans (RMCA: 27501); 2 ??
[2 AL], Masako, unknown date, leg. Ndey B. Ifuta (RMCA: 93.079-M-0346, 93.079-M-0370); 1 juv ♀
[AL], Mauda, between 1 Mar. 1925 and 31 Mar. 1925, leg. Henri Schouteden (RMCA: 16228); 1 ad ♀,
1 sad ♂, 1 juv ♀ [3 AL], Mayumbe, unknown date, leg. De Pauw, unknown date, prior to 31 Dec. 1913,
leg. Georges Gilson (RMCA: 16495, 3432, 2668); 1 ad ♀ [SS], M’gayu, 17 Dec. 1909, leg. Herbert
Lang, James Paul Chapin and The American Museum Congo Expedition (RMCA: 12396); 1 ad ?, 2 juv
♂♂, 1 ♂ [3 SS, 1 UN], Mulungu, 1 Oct. 1946, leg. unknown collector, between 1 Jan. 1948 and
31 Dec. 1948, leg. F.L. Hendricks (RBINS: 6403, RMCA: 19646 to 19648); 2 ad ♀♀, 1 ad ♂ [3 SS],
Ngayu [River], 17 Dec. 1909, leg. Herbert Lang, James Paul Chapin and The American Museum Congo
Expedition (AMNH: 48982, 48983, 48985); 3 ?? [3 AL], N’Gayu area, 17 Dec. 1909, leg. Herbert
Lang, James Paul Chapin and The American Museum Congo Expedition (AMNH: 49310, 49319,
49337); 1 sad ♀, 1 sad ♂ [2 SS], Niangara, 16 Nov. 1910, 20 Nov. 1910, leg. Herbert Lang, James Paul
Chapin and The American Museum Congo Expedition (AMNH: 48986, 48987); 1 ad ?, 1 juv ♂, 1 juv ?
[1 AL, 2 SO], Paulis [= Isiro], 10 Dec. 1950, between 1 Jan. 1947 and 31 Dec. 1947, leg. Abbeloos
(RMCA: 20512, 17929, 17930); 2 ad ♀♀, 3 ad ♂♂, 2 juv ♂♂, 1 ♀, 2 ♂♂, 3 ?? [3 SS, 4 AL, 6 UN],
Rutshuru, unknown date, 25 Jan. 1938, 24 Feb. 1938, prior to 31 Dec. 1943, leg. unknown collector,
11 May 1953, 30 Aug. 1954, 19 Sep. 1955, prior to 22 Dec. 1943, leg. Gaston-François de Witte, prior
to 31 Dec. 1943, leg. Gaston-François de Witte et al. (INPBC: V1350.1, V1353.2, W1247, RMCA:
37163, 37166, RBINS: 4767, 4768, RMCA: 37164, 37165, 16633, 16634, RBINS: 4758, 4759); 3 ♀♀,
2 ♂♂ [3 SS, 2 UN], St. Jos. (Mission) [= Mikalaya], 31 May 1924, 5 Aug. 1924, 5 Oct. 1924,
27 Oct. 1924, leg. Richard Callewaert (AMNH: 86149, 86169, 86173, 86174, 86141); 2 juv ♀♀ [2 AL],
Stanleyville [= Kisangani], 2 Jul. 1929, leg. Collart, 24 Apr. 1929, leg. Geldof (RMCA: 16187, 9995);
242
VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
1 ad ♂, 1 ♀ [1 SS, 1 AL], Tandala, 16 Jul. 1979, leg. Lynn W. Robbins and C. Brian Robbins, 18 Jul. 1979,
leg. C. Brian Robbins (CM: 86667, USNM: 463506); 1 ad ♂ [AL], Tshakala, between 1 Jan. 1982 and
31 Jan. 1982, leg. Marc Colyn (RMCA: 82.019-M-0007); 1 ad ♂ [SS], Tshikapa, 24 Nov. 1921, leg.
Henri Schouteden (RMCA: 7255); 1 ad ♂, 1 juv ♂ [2 AL], Tshuapa River, upper, between 1 Jun. 1925
and 30 Jun. 1925, leg. J. Ghesquière (RMCA: 16267, 16268); 3 ♀♀, 3 ♂♂ [6 SS], Upper Congo,
unknown date, leg. Alexander Douglas Mitchell Carruthers (BMNH: 1907.6.14.15 to 1907.6.14.18,
1907.6.14.4, 1907.6.14.5); 6 ad ♀♀, 1 ad ♂, 1 juv ♀, 1 ♀, 1 ♂, 1 ? [8 AL, 3 UN], Uvira, unknown date,
leg. Rudolf Grauer, 2 Mar. 1955, between 1 Mar. 1955 and 31 Mar. 1955, leg. Narcisse Leleup, between
1 Apr. 1951 and 30 Apr. 1951, leg. F.L. Lambrecht, between 1 Nov. 1956 and 30 Nov. 1956, leg. Laurent
(ZMB: 67026 to 67028, RMCA: 27209 to 27213, 22912, 90.004-M-0001, 35688); 1 ad ♀, 1 juv ?
[2 UN], Vitshumbi, prior to 31 Dec. 1943, leg. unknown collector (RBINS: 4757.1, 4757.2); 2 ad ♀♀,
1 ad ♂, 2 ♀♀, 4 ♂♂ [3 SS, 6 AL], Yalosemba, 28 Jun. 1979, 16 Jul. 1979, leg. unknown collector,
17 Jun. 1979, leg. C. Brian Robbins, 28 Jun. 1979, leg. Lynn W. Robbins and C. Brian Robbins,
9 Jul. 1979, leg. Frieder Meier (ZMMU: S-180533, USNM: 463504, 463505, 537724, 537725, CM:
86666, MHNG: 1871.010, 1871.011, ZMMU: S-180534); 1 ♂ [UN], Zobia, 18 Apr. 1946, leg. unknown
collector (RBINS: 14509).
RWANDA: 1 ♂, 1 ? [2 UN], Gabiro, unknown date, 1 Jul. 1948, leg. unknown collector (RBINS: 13921,
13021); 1 juv ♂, 1 ♀, 2 ♂♂ [4 AL], Gisenyi, 5 Oct. 1980, leg. Hans H. de Iongh (ZMA: MAM.21233
to MAM.21236); 1 ad ♂, 1 ? [1 SA, 1 UN], Rwankwi, between 1 Apr. 1948 and 30 Apr. 1948, leg.
J.V. Leroy (RMCA: 19070, 19071); 1 ad ♂ [SA], Shangi Schad [= Shangi], 5 Jan. 1985, leg. Benny
Baeten and Luc Janssens (RMCA: 85.006-M-0457).
Neoromicia zuluensis (Roberts, 1924)
RWANDA: 1 ad ♀ [SA], Mimuli, 31 Oct. 1964, leg. U. Goepel (ZMZ: 126640).
Nycticeinops Hill & Harrison, 1987
Nycticeinops schlieffenii (Peters, 1859)
DEMOCRATIC REPUBLIC OF THE CONGO: 2 ♂♂ [2 UN], Aketi, 21 Jun. 2015, leg. Vlir/Aketi
(UNIKIS: AKETI601, AKETI602); 2 ♀♀, 1 ? [1 AL, 2 UN], Bagunda, unknown date, leg. unknown
collector, 18 Jan. 1951, leg. H. de Saeger et al., 18 Jan. 1951, leg. H. de Saeger (INPBC: 2751, RBINS:
13870, RMCA: 36888); 1 ? [UN], Bas Congo (former) Province [= Kongo Central Province],
unknown date, leg. Alphonse Cabra (RMCA: 65b); 1 ♀, 1 ♂ [2 SS], Dikul River [= Dikulwe Cave],
unknown date, leg. S.A. Neave (BMNH: 1907.12.13.3, 1907.12.13.4); 1 ♂ [UN], Gangala-na-Bodio,
8 Nov. 1949, leg. H. de Saeger et al. (RBINS: 13868); 3 ♀♀, 1 ♂ [4 SS], Kabenge, 3 Apr. 1949, leg.
l’ Institut des Parcs nationaux du Congo belge, 3 Apr. 1949, leg. unknown collector (RBINS: 10719 to
10722); 1 ? [SA], Katanga, prior to 23 Dec. 1947, leg. soeur de notre dame de Bethanie (RMCA: 18177);
1 ♀, 1 ♂ [2 SS], Kateke [= Kateke [River]], 1 Dec. 1947, leg. l’ Institut des Parcs nationaux du Congo
belge, 1 Dec. 1947, leg. unknown collector (RBINS: 10714, 10715); 1 ♀, 1 ♂ [2 SS], Luira River,
upper, unknown date, leg. S.A. Neave (BMNH: 1907.12.13.5, 1907.12.13.6); 1 ♀ [SS], Luluabourg
[= Kananga], unknown date, leg. Richard Callewaert (BMNH: 1926.7.6.112); 2 ♂♂ [2 SS], Mabwe,
13 Dec. 1948, leg. l’ Institut des Parcs nationaux du Congo belge, 22 Dec. 1948, leg. unknown collector
(RBINS: 10716, 10717); 1 ♂ [SS], Masombwe, 24 Dec. 1948, leg. unknown collector (RBINS: 10798);
1 ♂ [SS], Munoi [= Munoi conluence], 18 Jun. 1948, leg. l’ Institut des Parcs nationaux du Congo
belge (RBINS: 10718); 2 ♀♀ [2 UN], Musosa, near, 1 Jun. 1940, between 1 Jun. 1940 and 30 Jun. 1940,
leg. unknown collector (RBINS: 14515, 14514).
243
European Journal of Taxonomy 382: 1–327 (2017)
Pipistrellus Kaup, 1829
BURUNDI: 2 ♂♂ [2 NS], Mpinga Cave [= Mpinga Mission], 25 Mar. 1970, leg. P.L. Niort (MHNG:
1607.099, 1607.100); 1 ♀ [AL], Nyanza [= Nyanza-Lac], 28 Feb. 1920, leg. Henry Cushier Raven
(USNM: 237232); 1 ♂ [NS], Rugari Mission, between 1 Oct. 1969 and 31 Oct. 1969, leg. P.L. Niort
(MHNG: 1607.089); 1 ♀ [NS], Rwintare Mission [= Rwintare], 19 Jun. 1969, leg. P.L. Niort (MHNG:
1607.083).
DEMOCRATIC REPUBLIC OF THE CONGO: 10 ♀♀, 24 ♂♂ [34 AL], Batala’s Village, 13 Apr. 1925,
16 Apr. 1925, leg. Edmund Heller (FMNH: 30635 to 30668); 1 ? [AL], Belgian River, 30 Dec. 1926,
leg. unknown collector (MCZ: 25124); 2 ♂♂, 6 ?? [2 SS, 6 SA], Bumba, 1 Jan. 1927, leg. David
H. Linder, 1 Jan. 1927, leg. Harold Jefferson Coolidge, 30 Dec. 1926, 30 Dec. 1930, leg. unknown
collector (MCZ: 23380, 23381, 25125 to 25130); 9 ♀♀, 7 ♂♂ [16 AL], Kabengere, 22 Dec. 1926,
24 Dec. 1926, 27 Dec. 1926, 28 Dec. 1926, leg. John Todd Zimmer (FMNH: 30682 to 30685, 30687 to
30698); 1 ♀, 5 ♂♂ [6 AL], Mabali, between 1 Jun. 1956 and 30 Jun. 1956, leg. Alvin Novick (MCZ:
48252 to 48257); 1 juv ? [UN], Rutshuru, unknown date, leg. unknown collector (INPBC: W3.2).
Pipistrellus grandidieri (Dobson, 1876)
BURUNDI: 1 ad ♀, 1 ad ♂ [1 SA, 1 SK], Resha, 1 Mar. 1979, 4 Apr. 1979, leg. Heinz Stephan (SMF:
57586, 57587).
Pipistrellus hesperidus (Temminck, 1840)
BURUNDI: 1 ad ♂ [SA], Nyamugari Hill, 20 Apr. 1991, leg. L. Davenport (FMNH: 148368).
RWANDA: 2 ad ♀♀ [2 SA], Kitabi, 2 Nov. 1981, 3 Nov. 1981, leg. Frits De Vree, Benny Baeten
and Victor Van Cakenberghe (RMCA: 82.006-M-0926, 82.006-M-0927); 1 ad ♂ [SA], Mutura,
24 Oct. 1981, leg. Frits De Vree, Benny Baeten and Victor Van Cakenberghe (RMCA: 82.006-M-0925).
Pipistrellus nanulus Thomas, 1904
DEMOCRATIC REPUBLIC OF THE CONGO: 1 ad ♀ [SA], Avakubi, 26 Feb. 1914, leg. Herbert
Lang, James Paul Chapin and The American Museum Congo Expedition (AMNH: 49323); 1 ad ♂ [SK],
Mosenge, 15 Aug. 1986, leg. WHO and Marc Colyn (RMCA: 88.011-M-0020).
Pipistrellus rueppellii (Fischer, 1829)
DEMOCRATIC REPUBLIC OF THE CONGO: 1 juv ♀, 1 juv ♂ [1 SA, 1 AL], Albertville [= Kalemie],
prior to 18 Nov. 1959, leg. H. Bomans (RMCA: 29300, 29301); 1 juv ♂ [AL], Beno, between 1 Jan. 1949
and 31 Dec. 1949, leg. P. Bayet (RMCA: 19853); 1 ad ♂ [AL], Ibembo, between 1 Nov. 1951 and
30 Nov. 1951, leg. Jozef Hutsebaut (RMCA: 20641); 1 ad ♀ [SA], Jadotville [= Likasi], 2 Jun. 1958, leg.
Félix / Michel Anciaux de Faveaux (RMCA: 27586); 1 ? [UN], Kasindi, unknown date, leg. unknown
collector (INPBC: W979.1); 1 ad ? [SS], Kawa (Forest), between 14 Jun. 1952 and 24 Jul. 1952, leg.
Alexandre Fain (RMCA: 23152); 1 ad ♀ [AL], Lukula, between 5 Jan. 1912 and 26 Jan. 1912, leg.
Daniel (RMCA: 993); 1 ? [UN], Moanda [= Muanda], unknown date, leg. unknown collector (RMCA:
12693); 1 juv ♀ [SA], Moba, 29 Jan. 1954, leg. H. Bomans (RMCA: 22125); 1 ad ♂ [AL], Mongbwalu,
13 Feb. 1934, leg. Jordens (RMCA: 12247); 1 ? [AL], Mweka Kasai [= Mweka], between 1 Jan. 1930
and 31 Dec. 1930, leg. Heusgens (RMCA: 11549); 1 ad ♀ [SA], Nepoko Bomili, between 1 Jan. 1946
and 31 Dec. 1946, leg. M. Henrion (RMCA: 18138); 1 ad ♂ [SA], Poko, 22 Aug. 1913, leg. Herbert
Lang, James Paul Chapin and The American Museum Congo Expedition (AMNH: 49326).
244
VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
Scotoecus Thomas, 1901
Scotoecus albofuscus (Thomas, 1890)
DEMOCRATIC REPUBLIC OF THE CONGO: 1 ? [SA], Baudouinville [= Moba], 12 Dec. 1953, leg.
H. Bomans (RMCA: 22412).
Scotoecus hindei Thomas, 1901
DEMOCRATIC REPUBLIC OF THE CONGO: 1 ? [SA], Kapolowe, prior to 24 May 1954, leg.
De Smet (RMCA: 22192); 1 ? [SA], Niangara, 20 Jun. 1913, leg. Herbert Lang, James Paul Chapin and
The American Museum Congo Expedition (AMNH: 49320).
245
European Journal of Taxonomy 382: 1–327 (2017)
Summary table of museum specimens
Overview of the museum acronyms.
Acronym
Name + Contact name + Database website
AMNH
American Museum of Natural History, New York, USA; Nancy B. Simmons
http://sci-web-001.amnh.org/db/emuwebamnh/index.php
BMNH
The Natural History Museum (formerly British Museum of Natural History), London, UK; Paulina
Jenkins
http://data.nhm.ac.uk/
CAS
California Academy of Sciences, San Francisco, USA; Maureen Flannery
http://researcharchive.calacademy.org/research/bmammals/MamColl/index.asp
CM
Carnegie Museum, Pittsburgh, USA; Suzanne B. McLaren
DNSM
Durban Natural Science Museum, Durban, South Africa; Leigh Richards
FMNH
Field Museum of Natural History, Chicago, USA; Bruce D. Patterson
http://emuweb.ieldmuseum.org/mammals/taxonomic.php?_ga=1.142122738.1331713266.1463234212
HNHM
Hungarian Natural History Museum, Budapest, Hungary; Gábor Csorba
HZM
Harrison Institute (formerly Harrison Zoological Museum), Sevenoaks, UK; Paul Bates
INPBC
Institute of the National Parks of Belgian Congo, Brussels, Belgium
KU
University of Kansas, Natural History Museum, Lawrence, USA; Maria Eifler
http://collections.biodiversity.ku.edu/KUMammals/
MCZ
Museum of Comparative Zoology, Harvard University, Cambridge, USA; Judith Chupasko
http://mczbase.mcz.harvard.edu/SpecimenSearch.cfm
MHNG
Muséum d’Histoire Naturelle, Genève, Switzerland; Manuel Ruedi
MNHN
Muséum national d’Histoire naturelle, Paris, France; Jean Marc Pons
https://science.mnhn.fr/institution/mnhn/collection/zm/item/search
MSB
Museum of Southwestern Biology, Albuquerque, USA; Joseph Cook
http://arctos.database.museum/SpecimenSearch.cfm
NMBA
Naturhistorisches Museum, Basel, Switzerland; Denis Vallan, Raffael Winkler
NMP
National Museum (Natural History), Prague, Czech Republic; Petr Benda
NMW
Naturhistorisches Museum, Vienna, Austria; Frank Emmanuel Zachos
NMZL
National Museum of Zambia, Livingstone, Zambia; Clare Mateke
OSU
Oklahoma State University, Stillwater, USA; Karen McBee
RBINS
Royal Belgian Institute for Natural Sciences, Brussels, Belgium; Georges Lenglet, Tom Geerinckx,
Olivier Pauwels
https://darwin.naturalsciences.be/
RMCA
Royal Museum of Central Africa, Tervuren, Belgium; Wim Wendelen, Emmanuel Gilissen
RMNH
Naturalis - National Natuurhistorish Museum (formerly Rijksmuseum voor Natuurlijke Historie),
Leiden, the Netherlands; Pepijn Kamminga, Steven van der Mije
http://bioportal.naturalis.nl/
246
VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
Acronym
Name + Contact name + Database website
SMF
Senckenberg Museum, Frankfurt, Germany; Katrin Krohmann
http://sesam.senckenberg.de/page/index.htm
SMNS
Staatliches Museum für Naturkunde, Stuttgart, Germany; Stefan Merker
TM
Ditsong National Museum of Natural History (formerly Transvaal Museum), Pretoria, South
Africa; Teresa Kearney
TTU
Texas Tech University (The Museum), Lubbock, USA; Heath J. Garner
UNIKIS
Université de Kisangani, Democratic Republic of the Congo
USNM
United States National Museum, Washington DC, USA; Kristofer M. Helgen, Darrin P. Lunde
http://collections.nmnh.si.edu/search/mammals/
WNMU
Western New Mexico University, Silver City, USA; Randy Jennings
ZFMK
Zoologisches Forschungsinstitut und Museum Alexander Koenig, Bonn, Germany; Jan Decher
ZMA
Zoologisch Museum, Amsterdam, The Netherlands (currently at Naturalis, Leiden)
ZMB
Zoologisches Museum Berlin, Germany; Frieder Mayer
ZMH
Zoologisches Institut und Zoologisches Museum, Hamburg, Germany; Thomas M. Kaiser
ZMMU
Zoological Museum Moscow University, Moscow, Russia; Sergei V. Kruskop
ZMUC
Natural History Museum of Denmark (formerly Zoologisk Museum), University of Copenhagen,
Denmark; Eline Lorenzen, Daniel Klingberg Johansson
ZMZ
Zoologisches Museum der Universität, Zurich, Switzerland; Barbara Oberholzer
In the following table (pages 248–259), we provide an overview of the number of CRB specimens in
the various museums. In most cases, the number of specimens we report is higher than the number
reported by Hayman et al. (1966: 77). The only exceptions are INPBC, where part of the material
has been divided between the RBINS and RMCA collections, and the BMNH, where Hayman et al.
(1966) did not provide any registration numbers and we were unable to trace the specimens involved.
Hayman et al. (1966: 77) also reported an additional 45 specimens (1 Epomophorus anurus, 1 Epomops
franqueti, 1 Hypsignatus monstrosus, 2 Taphozous mauritianus, 4 Saccolaimus peli, 1 Nycteris arge, 1
N. hispida, 7 Lavia frons, 4 Hipposideros caffer, 4 H. ruber, 4 Myotis bocagii, 6 Neoromicia nana, 7
Pipistrellus rueppellii, 1 Nycticeinops schlieffeni and 1 Miniopterus inlatus) from literature data only
(Noack 1887; Gyldenstolpe 1928; Sanborn 1936; Hatt 1940; Allen & Loveridge 1942).
247
7
1
1
4
34
14
32
328
70
3
Epomophorus Bennett, 1836
11
4
1
14
28
92
583
1
Epomophorus anselli Bergmans & Van Strien, 2004
3
Epomophorus crypturus Peters, 1852
26
10
1
12
1
25
1
54
158
3
1
1
5
Epomophorus minimus Claessen & De Vree, 1991
Epomophorus minor Dobson, 1880
2
Epomophorus wahlbergi (Sundevall, 1846)
11
2
4
2
4
11
4
3
7
89
13
4
3
7
357
1
12
116
2
Epomops Gray, 1866
2
Epomops dobsonii (Bocage, 1889)
1
248
Epomops franqueti (Tomes, 1860)
31
12
9
Hypsignathus monstrosus H. Allen, 1862
44
6
4
32
3
7
1
1
6
Micropteropus Matschie, 1899
4
Micropteropus intermedius Hayman, 1963
Micropteropus pusillus (Peters, 1868)
15
15
1
31
1
4
55
37
11
450
10
164
3
Nanonycteris veldkampii (Jentink, 1888)
Megaloglossus woermanni Pagenstecher, 1885
3
2
20
Myonycteris angolensis (Bocage, 1898)
4
8
61
1
4
13
12
15
6
230
1
Myonycteris relicta Bergmans, 1980
Myonycteris torquata (Dobson, 1878)
4
2
12
5
1
1
Plerotes anchietae (Seabra, 1900)
Casinycteris argynnis Thomas, 1910
1
3
203
1
2
2
6
Scotonycteris bergmansi Hassanin, Khouider, Gembu,
Goodman, Kadjo, Nesi, Pourrut, Nakouné & Bonillo, 2015
Rousettus Gray, 1821
5
8
1
3
3
1
1
83
25
European Journal of Taxonomy 382: 1–327 (2017)
135
Rousettinae K. Andersen, 1912
Epomophorus labiatus (Temminck, 1837)
RMCA
RBINS
OSU
NMZL
NMW
NMP
NMBA
MSB
MNHN
MHNG
MCZ
KU
INPBC
HZM
HNHM
FMNH
88
DNSM
Eidolon helvum (Kerr, 1792)
CM
4
CAS
CHIROPTERA Blumenbach, 1779
BMNH
AMNH
Taxon/Museum
RBINS
RMCA
OSU
NMZL
NMW
9
NMP
11
NMBA
MNHN
14
MSB
MHNG
2
MCZ
19
KU
4
INPBC
6
HZM
Stenonycteris lanosus (Thomas, 1906)
HNHM
27
FMNH
5
DNSM
8
CM
BMNH
6
CAS
AMNH
Rousettus aegyptiacus (E. Geoffroy St.-Hilaire, 1810)
Taxon/Museum
29
294
7
46
HIPPOSIDERIDAE Lydekker, 1891
44
Doryrhina cyclops (Temminck, 1853)
14
6
6
25
33
1
2
6
Hipposideros beatus (K. Andersen, 1906)
Hipposideros caffer (Sundevall, 1846)
7
34
22
162
85
17
11
1
Hipposideros Gray, 1831
Hipposideros abae J.A. Allen, 1917
1
3
5
Hipposideros camerunensis Eisentraut, 1956
1
Hipposideros fuliginosus (Temminck, 1853)
2
4
18
110
249
3
6
2
3
1
127
548
15
63
1
204
211
1
5
3
24
66
102
37
8
3
Hipposideros ruber (Noack, 1893)
73
40
18
Macronycteris gigas (Wagner, 1845)
1
1
2
3
3
2
1
1
Macronycteris vittatus (Peters, 1852)
Cardioderma cor (Peters, 1872)
Lavia frons (E. Geoffroy St.-Hilaire, 1810)
28
2
7
1
1
1
2
Rhinolophus Lacépède, 1799
Rhinolophus alcyone Temminck, 1853
15
1
2
4
1
1
2
1
Rhinolophus blasii Peters, 1867
Rhinolophus clivosus Cretzschmar, 1828
Rhinolophus eloquens K. Andersen, 1905
5
Rhinolophus fumigatus Rüppell, 1842
9
1
2
4
2
18
23
10
3
33
1
2
Rhinolophus hildebrandtii Peters, 1878
1
9
21
28
4
1
14
26
50
150
1
Rhinolophus hilli Aellen, 1973
1
Rhinolophus kahuzi Fahr & Kerbis Peterhans, 2013
Rhinolophus landeri Martin, 1838
361
4
1
1
3
1
1
18
108
VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
18
Asellia tridens (E. Geoffroy St.-Hilaire, 1813)
RMCA
3
12
22
Rhinolophus simulator K. Andersen, 1904
2
Rhinolophus swinnyi Gough, 1908
35
17
1
11
3
1
9
6
3
1
21
39
147
4
Rhinolophus willardi Kerbis Peterhans & Fahr, 2013
Cloeotis percivali Thomas, 1901
Coleura afra (Peters, 1852)
14
8
Saccolaimus peli (Temminck, 1853)
25
6
4
Taphozous mauritianus E. Geoffroy St.-Hilaire, 1818
48
5
10
2
6
2
1
1
Taphozous nudiventris Cretzschmar, 1830
Taphozous perforatus E. Geoffroy St.-Hilaire, 1818
8
2
2
Nycteris arge Thomas, 1903
18
17
2
4
250
5
Nycteris grandis Peters, 1865
Nycteris hispida (Schreber, 1774)
1
1
Nycteris G. Cuvier & E. Geoffroy, 1795
29
14
1
3
1
8
3
2
Nycteris intermedia Aellen, 1959
Nycteris macrotis Dobson, 1876
2
Nycteris major (K. Andersen, 1912)
1
22
10
4
2
21
4
1
3
4
1
6
7
2
Nycteris nana (K. Andersen, 1912)
3
4
2
3
10
41
3
17
1
13
69
1
2
21
3
83
151
1
1
1
2
2
1
1
68
79
30
19
1
Nycteris parisii (de Beaux, 1924)
3
Nycteris thebaica E. Geoffroy St.-Hilaire, 1818
19
2
11
5
MOLOSSIDAE Gervais, 1856
2
Chaerephon Dobson, 1874
Chaerephon aloysiisabaudiae (Festa, 1907)
1
Chaerephon ansorgei (Thomas, 1913)
24
Chaerephon bemmeleni (Jentink, 1879)
Chaerephon bivittatus (Heuglin, 1861)
17
263
4
19
2
2
2
1
4
58
1
1
15
97
1
1
European Journal of Taxonomy 382: 1–327 (2017)
RBINS
OSU
5
NMZL
NMW
NMP
NMBA
MSB
MNHN
MHNG
MCZ
KU
INPBC
HZM
HNHM
FMNH
DNSM
3
CM
1
CAS
BMNH
Rhinolophus ruwenzorii J. Eric Hill, 1942
AMNH
Taxon/Museum
RMCA
RBINS
2
OSU
NMZL
NMW
NMP
NMBA
MSB
MNHN
MHNG
MCZ
KU
INPBC
HZM
HNHM
FMNH
DNSM
CM
CAS
BMNH
Chaerephon chapini J.A. Allen, 1917
AMNH
Taxon/Museum
3
1
Chaerephon gallagheri (Harrison, 1975)
28
Chaerephon major (Trouessart, 1897)
34
Chaerephon nigeriae Thomas, 1913
1
Chaerephon pumilus (Cretzschmar, 1826)
12
2
2
5
18
3
1
17
117
Chaerephon russatus J.A. Allen, 1917
24
1
Mops (Mops) condylurus (A. Smith, 1833)
2
2
Mops (Mops) congicus J.A. Allen, 1917
15
Mops (Mops) demonstrator (Thomas, 1903)
2
2
Mops (Mops) midas (Sundevall, 1843)
6
9
Mops (Mops) niangarae J.A. Allen, 1917
1
1
2
5
1
54
171
1
251
1
Mops (Xiphonycteris) brachypterus (Peters, 1852)
23
4
2
Mops (Xiphonycteris) nanulus J.A. Allen, 1917
6
14
1
Mops (Xiphonycteris) spurrelli (Dollman, 1911)
1
Mops (Xiphonycteris) thersites (Thomas, 1903)
8
10
Myopterus daubentonii Desmarest, 1820
7
1
Myopterus whitleyi (Scharff, 1900)
4
7
6
16
39
1520
1
15
10
1
2
5
2
2
87
3
2
23
11
22
23
2
119
1
3
628
2
8
Mops (Mops) trevori J.A. Allen, 1917
103
4
2
3
Mops (Mops) niveiventer Cabrera & Ruxton, 1926
5
1
6
1
38
9
1
1
9
1
1
Tadarida Rainesque, 1814
2
1
1
1
1
Tadarida fulminans (Thomas, 1903)
21
2
9
Otomops martiensseni (Matschie, 1897)
48
1
3
12
2
1
1
17
Tadarida ventralis (Heuglin, 1861)
1
VESPERTILIONOIDEA Gray, 1821
44
Miniopterus fraterculus Thomas & Schwann, 1906
17
7
7
7
3
5
10
23
VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
Chaerephon leucogaster (A. Grandidier, 1869)
VESPERTILIONIDAE Gray, 1821
1
33
2
12
115
1
5
75
381
2
1
2
Kerivoula Gray, 1842
4
Kerivoula argentata Tomes, 1861
1
3
3
Kerivoula cuprosa Thomas, 1912
1
Kerivoula lanosa (A. Smith, 1847)
Kerivoula phalaena Thomas, 1912
3
1
1
Kerivoula smithii Thomas, 1880
3
2
Myotis bocagii (Peters, 1870)
31
5
3
5
1
16
8
5
252
Myotis tricolor (Temminck, 1832)
2
4
Myotis welwitschii (Gray, 1866)
4
2
10
Scotophilus Leach, 1821
Scotophilus dinganii (A. Smith, 1833)
2
Scotophilus leucogaster (Cretzschmar, 1826)
3
2
Scotophilus nigrita (Schreber, 1774)
7
1
Scotophilus nux Thomas, 1904
2
2
1
11
25
7
11
1
8
4
3
16
139
1
9
3
5
1
10
1
1
Glauconycteris argentata (Dobson, 1875)
10
Glauconycteris beatrix Thomas, 1901
2
1
3
1
1
22
71
1
9
1
Glauconycteris curryae Eger & Schlitter, 2001
Glauconycteris humeralis J.A. Allen, 1917
6
Glauconycteris poensis (Gray, 1842)
1
1
1
3
1
2
2
Glauconycteris superba Hayman, 1939
Glauconycteris variegata (Tomes, 1861)
155
1
Glauconycteris Dobson, 1875
Glauconycteris alboguttata J.A. Allen, 1917
2
7
20
2
5
2
2
20
European Journal of Taxonomy 382: 1–327 (2017)
1
RMCA
5
RBINS
1
OSU
10
NMZL
NMW
NMP
NMBA
MSB
MNHN
3
MHNG
MCZ
KU
26
3
INPBC
Miniopterus natalensis (A. Smith, 1833)
HZM
1
HNHM
Miniopterus minor Peters, 1867
FMNH
6
DNSM
CAS
10
CM
BMNH
AMNH
Miniopterus inlatus Thomas, 1903
Taxon/Museum
RMCA
RBINS
OSU
NMZL
NMW
NMP
NMBA
MSB
1
1
2
1
Laephotis angolensis Monard, 1935
2
Laephotis botswanae Setzer, 1971
2
4
1
6
1
3
1
4
33
1
7
1
18
36
137
1085
2
55
1
Mimetillus thomasi Hinton, 1920
1
Neoromicia Roberts, 1926
1
253
Neoromicia brunnea (Thomas, 1880)
1
Neoromicia capensis (A. Smith, 1829)
1
Neoromicia guineensis (Bocage, 1889)
2
Neoromicia nana (Peters, 1852)
87
29
12
21
49
835
18
Neoromicia rendalli (Thomas, 1889)
38
5
11
3
1
3
2
Neoromicia somalica (Thomas, 1901)
1
1
Neoromicia tenuipinnis (Peters, 1872)
18
11
1
2
35
16
1
1
3
2
4
8
12
15
4
1
17
130
14
3
Neoromicia zuluensis (Roberts, 1924)
5
Nycticeinops schlieffenii (Peters, 1859)
1
50
Pipistrellus Kaup, 1829
1
Pipistrellus grandidieri (Dobson, 1876)
1
Pipistrellus hesperidus (Temminck, 1840)
Pipistrellus nanulus Thomas, 1904
1
Pipistrellus rueppellii (Fischer, 1829)
1
3
1
1
12
1
Scotoecus albofuscus (Thomas, 1890)
Scotoecus hindei Thomas, 1901
1
1
Grand Total
1031 403
Number in Hayman et al. (1966)
760
537
20
29
3
393
4
213 1564
2143
10
191
647
170
250
250
2
1
13
2
1
1904 10730
1383
3529
VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
1
Hypsugo musciculus (Thomas, 1913)
Mimetillus moloneyi (Thomas, 1891)
MNHN
MHNG
MCZ
KU
INPBC
HZM
HNHM
FMNH
DNSM
CM
CAS
2
Hypsugo crassulus (Thomas, 1904)
Hypsugo cf. eisentrauti Hill, 1968
BMNH
Hypsugo anchietae (Seabra, 1900)
AMNH
Taxon/Museum
203
64
3
3
18
4
2
2
560
19
22
3
1
Epomophorus anselli Bergmans & Van Strien, 2004
1
Epomophorus crypturus Peters, 1852
Epomophorus labiatus (Temminck, 1837)
347
70
Rousettinae K. Andersen, 1912
Epomophorus Bennett, 1836
Total in
Grand
Hayman et al.
Total
(1966)
1
37
2
2
2
19
103
37
974
253
5
Epomophorus minimus Claessen & De Vree, 1991
Epomophorus minor Dobson, 1880
3
Epomophorus wahlbergi (Sundevall, 1846)
3
1
Epomops dobsonii (Bocage, 1889)
254
Epomops franqueti (Tomes, 1860)
1
Hypsignathus monstrosus H. Allen, 1862
1
9
9
Micropteropus intermedius Hayman, 1963
1
Micropteropus pusillus (Peters, 1868)
63
77
21
8
194
6
7
1
680
125
8
3
1
2
214
132
23
23
Micropteropus Matschie, 1899
17
4
4
Epomops Gray, 1866
11
136
1
1
3
5
2
688
113
3
Nanonycteris veldkampii (Jentink, 1888)
Megaloglossus woermanni Pagenstecher, 1885
32
Myonycteris angolensis (Bocage, 1898)
10
169
6
1
6
479
7
10
1
309
66
1
Myonycteris relicta Bergmans, 1980
22
Myonycteris torquata (Dobson, 1878)
318
2
5
Plerotes anchietae (Seabra, 1900)
583
9
5
4
Casinycteris argynnis Thomas, 1910
2
17
33
6
149
4
Scotonycteris bergmansi Hassanin, Khouider, Gembu, Goodman, Kadjo,
Nesi, Pourrut, Nakouné & Bonillo, 2015
1
5
19
1
63
2
Rousettus Gray, 1821
4
European Journal of Taxonomy 382: 1–327 (2017)
ZMZ
ZMUC
ZMMU
ZMH
ZMB
ZMA
ZFMK
WNMU
USNM
Eidolon helvum (Kerr, 1792)
UNIKIS
CHIROPTERA Blumenbach, 1779
TTU
TM
SMNS
SMF
RMNH
Taxon/Museum
95
6
ZMZ
4
ZMUC
4
3
Stenonycteris lanosus (Thomas, 1906)
ZMMU
186
ZMH
694
ZMB
14
ZMA
11
ZFMK
USNM
34
WNMU
UNIKIS
206
TTU
16
TM
10
SMNS
SMF
Rousettus aegyptiacus (E. Geoffroy St.-Hilaire, 1810)
RMNH
Taxon/Museum
Total in
Grand
Hayman et al.
Total
(1966)
44
Asellia tridens (E. Geoffroy St.-Hilaire, 1813)
18
Doryrhina cyclops (Temminck, 1853)
7
Hipposideros Gray, 1831
8
Hipposideros abae J.A. Allen, 1917
1
Hipposideros beatus (K. Andersen, 1906)
3
Hipposideros caffer (Sundevall, 1846)
28
126
3
15
6
1
13
9
2
13
7
1
Hipposideros camerunensis Eisentraut, 1956
Hipposideros fuliginosus (Temminck, 1853)
1
Hipposideros ruber (Noack, 1893)
32
1
9
13
255
1
Macronycteris gigas (Wagner, 1845)
17
Macronycteris vittatus (Peters, 1852)
1
1
1
2
1
281
57
16
883
475
1
1
7
3
707
671
12
14
47
6
237
4
176
50
2
Rhinolophus Lacépède, 1799
284
1
1
Cardioderma cor (Peters, 1872)
Lavia frons (E. Geoffroy St.-Hilaire, 1810)
20
101
Rhinolophus alcyone Temminck, 1853
4
2
Rhinolophus blasii Peters, 1867
1
3
1
Rhinolophus clivosus Cretzschmar, 1828
5
420
46
23
Rhinolophus eloquens K. Andersen, 1905
223
Rhinolophus fumigatus Rüppell, 1842
Rhinolophus hildebrandtii Peters, 1878
11
1
Rhinolophus hilli Aellen, 1973
Rhinolophus ruwenzorii J. Eric Hill, 1942
328
69
246
163
2
1
Rhinolophus kahuzi Fahr and Kerbis Peterhans, 2013
Rhinolophus landeri Martin, 1838
1
10
1
148
116
24
6
VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
HIPPOSIDERIDAE Lydekker, 1891
52
30
Rhinolophus simulator K. Andersen, 1904
Total in
Grand
Hayman et al.
Total
(1966)
Rhinolophus swinnyi Gough, 1908
54
Rhinolophus willardi Kerbis Peterhans & Fahr, 2013
4
5
Cloeotis percivali Thomas, 1901
Coleura afra (Peters, 1852)
1
Saccolaimus peli (Temminck, 1853)
2
Taphozous mauritianus E. Geoffroy St.-Hilaire, 1818
4
1
1
Taphozous nudiventris Cretzschmar, 1830
3
Taphozous perforatus E. Geoffroy St.-Hilaire, 1818
Nycteris arge Thomas, 1903
4
1
Nycteris grandis Peters, 1865
5
2
256
Nycteris hispida (Schreber, 1774)
2
9
Nycteris intermedia Aellen, 1959
1
1
Nycteris macrotis Dobson, 1876
2
5
1
3
1
1
3
3
1
4
1
Nycteris major (K. Andersen, 1912)
1
1
1
1
17
38
27
65
42
264
208
1
1
70
57
1
1
169
90
49
20
341
252
11
1
175
84
2
1
63
46
1
Nycteris parisii (de Beaux, 1924)
Nycteris thebaica E. Geoffroy St.-Hilaire, 1818
20
22
Nycteris G. Cuvier & E. Geoffroy, 1795
Nycteris nana (K. Andersen, 1912)
77
3
5
65
MOLOSSIDAE Gervais, 1856
263
Chaerephon Dobson, 1874
23
53
4
2
Chaerephon ansorgei (Thomas, 1913)
206
70
Chaerephon bemmeleni (Jentink, 1879)
2
2
Chaerephon bivittatus (Heuglin, 1861)
1
Chaerephon chapini J.A. Allen, 1917
5
Chaerephon gallagheri (Harrison, 1975)
1
Chaerephon aloysiisabaudiae (Festa, 1907)
1
4
European Journal of Taxonomy 382: 1–327 (2017)
ZMZ
ZMUC
ZMMU
ZMH
ZMB
ZMA
ZFMK
WNMU
USNM
UNIKIS
TTU
TM
SMNS
SMF
RMNH
Taxon/Museum
ZMZ
ZMUC
ZMMU
Chaerephon nigeriae Thomas, 1913
40
1
2
34
31
10
31
8
1872
530
1
30
28
1093
324
27
18
38
49
65
53
30
74
1
Mops (Mops) congicus J.A. Allen, 1917
13
5
Mops (Mops) demonstrator (Thomas, 1903)
2
Mops (Mops) midas (Sundevall, 1843)
Mops (Mops) niangarae J.A. Allen, 1917
10
2
1
1
142
20
1
Mops (Mops) trevori J.A. Allen, 1917
257
Mops (Xiphonycteris) brachypterus (Peters, 1852)
3
39
9
Mops (Xiphonycteris) nanulus J.A. Allen, 1917
6
76
63
Mops (Xiphonycteris) spurrelli (Dollman, 1911)
3
13
1
13
112
31
11
9
30
25
8
5
Mops (Xiphonycteris) thersites (Thomas, 1903)
1
Myopterus daubentonii Desmarest, 1820
1
Myopterus whitleyi (Scharff, 1900)
Otomops martiensseni (Matschie, 1897)
1
Tadarida Rainesque, 1814
2
Tadarida fulminans (Thomas, 1903)
19
2
Tadarida ventralis (Heuglin, 1861)
1
1
VESPERTILIONOIDEA Gray, 1821
44
Miniopterus fraterculus Thomas & Schwann, 1906
79
Miniopterus inlatus Thomas, 1903
6
Miniopterus minor Peters, 1867
Miniopterus natalensis (A. Smith, 1833)
26
38
203
309
8
3
550
49
VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
60
Chaerephon russatus J.A. Allen, 1917
Mops (Mops) niveiventer Cabrera & Ruxton, 1926
ZMH
8
Chaerephon major (Trouessart, 1897)
Mops (Mops) condylurus (A. Smith, 1833)
Total in
Grand
Hayman et al.
Total
(1966)
42
Chaerephon leucogaster (A. Grandidier, 1869)
Chaerephon pumilus (Cretzschmar, 1826)
ZMB
ZMA
ZFMK
WNMU
USNM
UNIKIS
TTU
TM
SMNS
SMF
RMNH
Taxon/Museum
Total in
Grand
Hayman et al.
Total
(1966)
4
1
2
Kerivoula Gray, 1842
2
Kerivoula argentata Tomes, 1861
2
12
Kerivoula cuprosa Thomas, 1912
3
Kerivoula lanosa (A. Smith, 1847)
4
3
3
1
Kerivoula phalaena Thomas, 1912
8
21
14
277
133
16
6
Myotis welwitschii (Gray, 1866)
13
8
Scotophilus Leach, 1821
11
6
Kerivoula smithii Thomas, 1880
Myotis bocagii (Peters, 1870)
5
Myotis tricolor (Temminck, 1832)
2
18
9
1
224
73
Scotophilus leucogaster (Cretzschmar, 1826)
5
2
Scotophilus nigrita (Schreber, 1774)
18
7
18
9
Scotophilus dinganii (A. Smith, 1833)
1
7
4
5
258
6
Scotophilus nux Thomas, 1904
2
1
Glauconycteris Dobson, 1875
Glauconycteris alboguttata J.A. Allen, 1917
14
Glauconycteris argentata (Dobson, 1875)
Glauconycteris beatrix Thomas, 1901
2
3
1
1
4
3
123
72
18
1
1
2
Glauconycteris humeralis J.A. Allen, 1917
3
15
6
Glauconycteris poensis (Gray, 1842)
1
4
2
3
1
52
40
Glauconycteris curryae Eger & Schlitter, 2001
1
Glauconycteris superba Hayman, 1939
1
Glauconycteris variegata (Tomes, 1861)
2
Hypsugo anchietae (Seabra, 1900)
Hypsugo crassulus (Thomas, 1904)
5
7
Hypsugo cf. eisentrauti Hill, 1968
1
4
2
European Journal of Taxonomy 382: 1–327 (2017)
ZMZ
ZMUC
ZMMU
ZMH
ZMB
ZMA
ZFMK
VESPERTILIONIDAE Gray, 1821
WNMU
USNM
UNIKIS
TTU
TM
SMNS
SMF
RMNH
Taxon/Museum
ZMZ
ZMUC
ZMMU
ZMH
ZMB
ZMA
ZFMK
WNMU
USNM
UNIKIS
TTU
TM
SMNS
SMF
RMNH
Taxon/Museum
Total in
Grand
Hayman et al.
Total
(1966)
Hypsugo musciculus (Thomas, 1913)
2
2
Laephotis angolensis Monard, 1935
2
13
11
3
Laephotis botswanae Setzer, 1971
53
1
19
1
Mimetillus thomasi Hinton, 1920
13
3
Neoromicia Roberts, 1926
Neoromicia brunnea (Thomas, 1880)
1
Neoromicia capensis (A. Smith, 1829)
59
Neoromicia guineensis (Bocage, 1889)
2
67
2389
1574
Neoromicia rendalli (Thomas, 1889)
1
121
63
Neoromicia somalica (Thomas, 1901)
1
6
Neoromicia tenuipinnis (Peters, 1872)
1
Neoromicia nana (Peters, 1852)
2
27
1
19
4
259
5
4
3
2
4
3
2
219
2
1
Neoromicia zuluensis (Roberts, 1924)
2
2
Pipistrellus grandidieri (Dobson, 1876)
22
71
1
Pipistrellus Kaup, 1829
1
25
2
Nycticeinops schlieffenii (Peters, 1859)
155
Pipistrellus hesperidus (Temminck, 1840)
4
Pipistrellus nanulus Thomas, 1904
2
1
Pipistrellus rueppellii (Fischer, 1829)
14
16
Scotoecus albofuscus (Thomas, 1890)
1
1
Scotoecus hindei Thomas, 1901
2
1
Grand Total
Number in Hayman et al. (1966)
44
568
2
540
2
1036
200
1
3
101
41
3
2
3
24
20231
8522
VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
6
Mimetillus moloneyi (Thomas, 1891)
European Journal of Taxonomy 382: 1–327 (2017)
Collecting localities for the bats from the CRB region
In this section we provide a list of all the localities in the three countries from which bats were
collected. For each of the localities, the name, country-province, coordinates and altitude are given.
We also provide alternative spellings and linguistic variations (English, Dutch and French). The
English version of the current name was chosen as “master locality” and the alternative versions are
mentioned in brackets preceded by “see alias:”. The aliases are followed by “see:” referring to the
“master locality”.
The coordinates for the localities were found and/or checked from various sources, e.g., “Global
Gazetteer Version 2.3” (http://www.fallingrain.com/world/), NGA GEOnet Names Server (GNS)
(http://geonames.nga.mil/gns/html/), literature data (including Hayman et al. 1966), (online) museum
catalogs and specimen labels.
An overview of the current provincial names and abbreviations used in the gazetteer (source:
http://www.statoids.com) and those used in Hayman et al. (1966) or historically is provided before the
actual list of collecting localities.
Province name
Abbreviation
Burundi
BY
Bubanza
BB
Bujumbura Mairie
BM
Bujumbura Rural
BL
Bururi
BR
Cankuzo
CA
Cibitoke
CI
Gitega
GI
Karuzi
KR
Kayanza
KY
Kirundo
KI
Makamba
MA
Muramvya
MU
Muyinga
MY
Mwaro
MW
Ngozi
NG
Rumonge
RM
Rutana
RT
Former province names
Abbreviation
BU
Ruyigi
RY
Democratic Republic of the Congo
CG
Bas-Uélé
BU
Orientale
PO
Équateur
ET
Équateur
EQ
Haut-Katanga
HK
Katanga
KT
Haut-Lomami
HL
Katanga
KT
Haut-Uélé
HU
Orientale
PO
Ituri
IT
Orientale
PO
Kasaï
KS
Kasai (Kasaï-Occidental)
KS
260
VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
Province name
Abbreviation
Former province names
Abbreviation
Kasaï-Central
LL
Kasai (Kasaï-Occidental)
KS
Kasaï Oriental
KO
Kasai (Kasaï-Oriental)
KS
Kinshasa
KN
Léopoldville (Kinshasa City)
LE
Kongo Central
BC
Léopoldville (Bas-Congo)
LE
Kwango
KG
Léopoldville (Bandundu)
LE
Kwilu
KU
Léopoldville (Bandundu)
LE
Lomami
LM
Kasai (Kasaï-Oriental)
KS
Lualaba
LB
Katanga
KT
Mai-Ndombe
MA
Léopoldville (Bandundu)
LE
Maniema
MN
Katanga (Maniema)
KT
Mongala
MO
Équateur
EQ
Nord-Kivu
NK
Kivu (Nord-Kivu)
KI
Nord-Ubangi
NU
Équateur
EQ
Sankuru
SN
Kasai (Kasaï-Oriental)
KS
Sud-Kivu
SK
Kivu (Sud-Kivu)
KI
Sud-Ubangi
SU
Équateur
EQ
Tanganyika
TG
Katanga
KT
Tshopo
TO
Orientale
PO
Tshuapa
TP
Équateur
Rwanda
RW
Eastern
ES
Umutara, Kibungo, Kigali-Rural (part)
Kigali City
KV
Kigali-Ville
Northern
NO
Byumba, Kigali-Rural (part), Ruhengeri
Southern
SU
Butare, Gikongoro, Gitarama
Western
OU
Cyangugu, Gisenyi, Kibuye
EQ
RU
Aba
Abatupi River
(see also: Amatupi River)
Abialosa River
Abimva
Abysse de Swanepoel II
Aka
Aka River
Akagera Park
(see also: Parc National de la Kagera, Kagera National Park)
Aka-Inimvua conluence
Akenge
(see: Akengei)
Akengei
(see also: Akenge)
Aketi
(see also: Port Chaltin)
Aketi Territory
Alambi
Albert Edward Nyanza
(see: Edward, Lake)
Albert Meer
(see: Albert, Lake)
Albert National Park
(see: Virunga National Park)
Albert Nyanza
(see: Albert, Lake)
261
CG-HU
CG-NK
03°52′ N, 30°14′ E
00°45′ N, 29°49′ E
890 m
700 m
CG-NK
CG-HU
CG-LB
CG-HU
CG-HU
RW-ES
00°42′ N, 29°41′ E
03°09′ N, 29°46′ E
10°51′ S, 26°41′ E
04°35′ N, 29°42′ E
04°30′ N, 29°40′ E
01°30′ S, 30°30′ E
800 m
1100 m
ca 1350 m
CG-HU
CG-BU
04°35′ N, 29°42′ E
02°56′ N, 26°50′ E
654 m
CG-BU
02°56′ N, 26°50′ E
654 m
CG-BU
02°50′ N, 23°56′ E
ca 500 m
CG-BU
CG-IT
CG-NK
02°07′ N, 28°49′ E
ca 00°08′ S, 29°35′ E
913 m
CG-IT
ca 01°40′ N, 30°40′ E
CG-NK
01°00′ S, 29°15′ E
CG-IT
ca 01°40′ N, 30°40′ E
European Journal of Taxonomy 382: 1–327 (2017)
Albert, Lac
(see: Albert, Lake)
Albert, Lake
(see also: Albert Meer, Albert Nyanza, Albert, Lac, Bantu Lueru, Bentu
Lueru, Lac Albert, Lake Albert, Lake Mobutu Sese Seko, Luta Nzige,
Mobutu Sese Seko, Lake, Muite Nzige, Mwittanzige)
Albertstad
(see: Kalemie)
Albertville
(see: Kalemie)
Albertville-Moba (Road?)
(see: Kalemie-Moba (Road?))
Alima
Alipago
(see also: Alipango, Alipaye, Alipei)
Alipango
(see: Alipago)
Alipaye
(see: Alipago)
Alipei
(see: Alipago)
Amangundu
Amatupi River
(see: Abatupi River)
Ambarao
(see: Ambarau)
Ambarau
(see also: Ambarao)
Ancien Beni
(see: Vieux Beni)
Ango
Ango Cave
(see also: Ango Grot, Grotte d’Ango)
Ango Grot
(see: Ango Cave)
Angu, near
Angumu
(see also: Angumu Etat)
Angumu Etat
(see: Angumu)
Ankoro
Arebi
Arebi Bondo Mabe
Astrida
(see: Butare)
Athens Mission
Avakubi
Avakubi, near
Azande Game Reserve
Ba Balaye
Babeke
Babongena
Badjoki - Bumba
(see: Bodjoki - Bumba)
Bafuaboka
(see: Bafwabaka)
Bafwabaka
(see also: Bafwaboka, Bafuaboka)
Bafwaboka
(see: Bafwabaka)
Bafwasende Territory
Baga Cave
(see: Baya Cave)
Baga Cave
Bagbele
(see also: Cel I, Cellule I)
Bagbele Moko
Bagisana
Bagunda
Bakwakenge
(see: Bakwa-Kenge)
Bakwa-Kenge
(see also: Bakwakenge)
Bakwanga
(see: Mbuji-Mayi)
Bala
(see: Bata)
262
CG-IT
ca 01°40′ N, 30°40′ E
CG-IT
ca 01°40′ N, 30°40′ E
CG-TG
05°56′ S, 29°12′ E
763 m
CG-TG
05°56′ S, 29°12′ E
763 m
CG-TG
05°57′ S, 29°01′ E
CG-NK
CG-BU
00°35′ N, 29°18′ E
03°34′ N, 25°44′ E
565 m
CG-BU
03°34′ N, 25°44′ E
565 m
CG-BU
03°34′ N, 25°44′ E
565 m
CG-BU
03°34′ N, 25°44′ E
565 m
CG-NK
CG-NK
00°45′ N, 29°48′ E
00°45′ N, 29°49′ E
700 m
700 m
CG-HU
03°10′ N, 29°44′ E
867 m
CG-HU
03°10′ N, 29°44′ E
867 m
CG-NK
00°26′ N, 29°33′ E
585 m
CG-BU
CG-BU
04°02′ N, 25°52′ E
04°02′ N, 25°52′ E
610 m
610 m
CG-BU
04°02′ N, 25°52′ E
610 m
CG-BU
CG-TO
ca 03°33′ N, 24°28′ E
00°07′ S, 27°42′ E
859 m
CG-TO
00°07′ S, 27°42′ E
859 m
CG-TG
CG-HU
CG-HU
RW-SU
06°45′ S, 26°57′ E
02°52′ N, 29°40′ E
02°47′ N, 29°35′ E
02°36′ S, 29°44′ E
1733 m
CG-KU
CG-TO
CG-TO
CG-HU
CG-KS
CG-IT
CG-TO
CG-MO
05°02′ S, 18°49′ E
01°19′ N, 27°33′ E
01°20′ N, 27°35′ E
05°50′ S, 22°30′ E
01°24′ N, 28°04′ E
00°29′ N, 25°42′ E
02°59′ N, 22°18′ E
608 m
469 m
441 m
CG-HU
02°07′ N, 27°40′ E
631 m
CG-HU
02°07′ N, 27°40′ E
631 m
CG-HU
02°07′ N, 27°40′ E
631 m
CG-TO
CG-HK
01°05′ N, 27°16′ E
11°52′ S, 27°27′ E
1279 m
CG-HK
CG-HU
ca 11°40′ S, 27°28′ E
04°21′ N, 29°17′ E
742 m
CG-HU
CG-HU
CG-HU
CG-KS
04°21′ N, 29°17′ E
04°35′ N, 29°32′ E
04°04′ N, 29°35′ E
04°51′ S, 22°04′ E
990 m
461 m
CG-KS
04°51′ S, 22°04′ E
461 m
CG-KO
06°09′ S, 23°36′ E
549 m
CG-KU
06°01′ S, 19°40′ E
590 m
589 m
742 m
VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
Baliko
Bamangwa
CG-TP
CG-HU
00°16′ S, 20°49′ E
04°21′ N, 29°27′ E
CG-ET
CG-BU
CG-IT
00°01′ N, 18°24′ E
03°28′ N, 25°43′ E
01°48′ N, 30°14′ E
CG-HU
04°21′ N, 29°27′ E
CG-HU
04°21′ N, 29°27′ E
CG-TO
CG-BC
01°33′ N, 25°20′ E
06°01′ S, 12°24′ E
432 m
0m
CG-BC
CG-BC
CG-BC
CG-BC
05°47′ S, 12°31′ E
05°58′ S, 12°27′ E
ca 05°58′ S, 12°27′ E
06°01′ S, 12°24′ E
0m
CG-ET
00°04′ N, 18°16′ E
306 m
CG-KU
CG
CG-BC
04°39′ S, 18°58′ E
CG-BC
05°16′ S, 14°53′ E
CG-IT
ca 01°40′ N, 30°40′ E
CG-BC
05°16′ S, 14°52′ E
ca 611 m
CG-BC
05°16′ S, 14°52′ E
ca 611 m
CG-BC
05°43′ S, 14°28′ E
CG-BC
05°43′ S, 14°28′ E
CG-SK
CG-SK
CG
04°05′ S, 29°05′ E
ca 04°05′ S, 29°05′ E
ca 05°00′ S, 15°00′ E
CG
ca 05°00′ S, 15°00′ E
(see: Bamungwa [River])
Bamanya
Bambesa
Bambu
(see: Kilomines)
Bamongwa
(see: Bamungwa [River])
Bamungwa [River]
(see also: Bamongwa, Bamangwa)
Banalia
Banana
(see also: Banane)
Banana - Boma Road, km 74, 25 km from coast in Forest
Banana Creek
Banana or Netonna
Banane
(see: Banana)
Bandaka
(see also: Cocquilhatville, Coquilhalville, Coquilhatstad, Coquilhatville,
Wangata)
Bandimbu
Bandungu (former) Province
Banjan Caves
(see also: Banjan Grotten, Grottes de Banjan, Thysville, Cave, Thysville,
Cave b 13 a)
Banjan Grotten
(see: Banjan Caves)
Bantu Lueru
(see: Albert, Lake)
Banza Ninda Caves
(see also: Banza Ninda Grot, Grotte de Banza Ninda)
Banza Ninda Grot
(see: Banza Ninda Caves)
Barainda Cave B 13
(see also: Barainda Grotte B 13)
Barainda Grotte B 13
(see: Barainda Cave B 13)
Baraka
Baraka, 5 mi [=8 km] N
Bas Congo (former) Province
(see: Kongo Central Province)
Bas Congo Province
(see: Kongo Central Province)
Bas Uele Province
(see also: Lower Uele Province)
Basoko
Basoko Territory
Basongo
Bata
(see also: Bala)
Batala’s Village
(see also: Beni, 30 mi [=48 km] W, Mambabwanga Hill)
Batoko (River)
(see: Olongo-Nsongo [River])
Baudhuinville
(see: Moba)
Baudouinville
(see: Moba)
Bauelende
Bauelendu
Baya carrière
(see: Baya quarry)
Baya Cave
(see also: M’baya Cave, Grotte de M’baya, M’baya Grot, Grotte de Baya,
Baga Cave)
(see: Baya quarry)
Baya Grot
(see: Baya quarry)
Baya quarry
(see also: Baya carrière, Baya Cave, Baya Grot, Carrière Baya, Carrière
Pierkat, Lubumbashi, 30 km S, Pierkat quarry)
Baye via Bondo
Befale
Bela
263
358–388 m
614 m
1466 m
05°16′ S, 14°53′ E
772 m
CG
CG-TO
CG-TO
CG-KS
CG-KU
01°14′ N, 23°36′ E
01°14′ N, 23°36′ E
04°20′ S, 20°24′ E
06°01′ S, 19°40′ E
329 m
590 m
CG-NK
00°30′ N, 29°15′ E
914 m
CG-MA
01°39′ S, 18°05′ E
CG-TG
07°03′ S, 29°47′ E
763 m
CG-TG
07°03′ S, 29°47′ E
763 m
CG-NK
CG-NK
CG-HK
00°16′ N, 29°36′ E
00°16′ N, 29°36′ E
11°52′ S, 27°27′ E
925 m
925 m
1279 m
CG-HK
11°52′ S, 27°27′ E
1279 m
CG-HK
11°52′ S, 27°27′ E
1279 m
CG-HK
11°52′ S, 27°27′ E
1279 m
CG-SN
CG-TP
CG-NK
04°25′ N, 23°04′ E
00°28′ N, 20°58′ E
00°38′ N, 29°14′ E
393 m
913 m
429 m
European Journal of Taxonomy 382: 1–327 (2017)
Belgian River
Bena Mbala
Bena Tshadi
(see also: Bena Tshandi, Bena Tshiadi)
Bena Tshandi
(see: Bena Tshadi)
Bena Tshiadi
(see: Bena Tshadi)
Bena-Kaniki
(see: Kaniki Kapangu)
Bene Longo
Bengale
Beni
(see also: New Beni)
Beni Territory
Beni Territory
Beni, 25 km NW
(see: Moera’s village)
Beni, 30 mi [=48 km] W
(see: Batala’s Village)
Beni, 36 km N
(see: Kisiki)
Beno
Bentu Lueru
(see: Albert, Lake)
Benza Masola
Benzale
Beyi - Kalonji
Biadimbi
Biakabomba
Biangolo
Bikoro
Bilalu
(see also: Bilule)
Bilule
(see: Bilalu)
Binga
Bionga
(see also: Biyonga)
Bitale
Bitshumbi
(see: Vitshumbi)
Biyonga
(see: Bionga)
Blukwa
Blukwa, 10 km E
(see: Wago, mount)
Boangi
Bobandana
Bodjoki - Bumba
(see also: Badjoki - Bumba)
Boende
Boende, 14 km N by Road
(see: Quatorze)
Boga
(see also: Mboga)
Bokala
Bokalakala
Bokoro
Bokuma
Bokungu
Bolafa
Bolobo
Bolobo Territory
Boma
Boma, Zone
Bomane
(see: Bomaneh)
Bomaneh
(see also: Bomane)
Bombanga - Bumba
Bombia Bonelu
Bombo Lumene
Bomili
Bondo
(see also: Lukula, 15 km W)
Bondo Mabe
264
CG
CG-LM
CG-KS
06°48′ S, 23°47′ E
04°40′ S, 22°49′ E
615 m
CG-KS
04°40′ S, 22°49′ E
615 m
CG-KS
04°40′ S, 22°49′ E
615 m
CG-KO
06°45′ S, 23°31′ E
805 m
CG-KS
CG-KN
CG-NK
04°52′ S, 21°40′ E
04°08′ S, 16°40′ E
00°30′ N, 29°28′ E
853 m
CG-NK
CG-NK
CG-NK
00°30′ N, 29°28′ E
00°30′ N, 29°28′ E
ca 00°35′ N, 29°25′ E
914 m
CG-NK
00°30′ N, 29°15′ E
914 m
CG-NK
00°45′ N, 29°34′ E
1177 m
CG-MA
CG-IT
03°37′ S, 17°48′ E
ca 01°40′ N, 30°40′ E
281 m
CG-BC
CG-MA
CG-KS
CG-HU
CG-KS
CG-NK
CG-ET
RW-ES
05°12′ S, 13°00′ E
04°08′ S, 16°40′ E
05°49′ S, 22°30′ E
04°13′ N, 29°21′ E
06°27′ S, 21°02′ E
00°25′ N, 29°45′ E
00°45′ S, 18°07′ E
01°30′ S, 30°18′ E
608 m
1300 m
344 m
1600 m
RW-ES
01°30′ S, 30°18′ E
1600 m
CG-MA
CG-TP
02°23′ N, 20°30′ E
00°34′ S, 23°00′ E
397 m
444 m
CG-SK
CG-NK
02°13′ S, 28°38′ E
00°41′ S, 29°23′ E
907 m
CG-TP
00°34′ S, 23°00′ E
444 m
CG-IT
CG-IT
01°45′ N, 30°36′ E
01°45′ N, 30°40′ E
2001 m
2240 m
CG-TP
CG
CG-MO
01°51′ S, 20°56′ E
01°42′ S, 29°01′ E
02°59′ N, 22°18′ E
419 m
CG-TP
CG-TP
00°13′ S, 20°52′ E
00°10′ N, 20°56′ E
371 m
326 m
CG-IT
01°03′ N, 29°56′ E
1562 m
CG-MA
CG-MA
CG-MA
CG-ET
CG-TP
CG-MO
CG-TP
CG-MA
CG-BC
CG-BC
CG-TO
03°07′ S, 17°02′ E
02°08′ S, 16°22′ E
02°50′ S, 18°23′ E
00°06′ S, 18°42′ E
00°41′ S, 22°19′ E
02°08′ N, 21°37′ E
02°09′ S, 16°14′ E
307 m
252 m
252 m
340 m
402 m
05°51′ S, 13°03′ E
ca 05°51′ S, 13°03′ E
01°18′ N, 23°47′ E
61 m
429 m
CG-TO
01°18′ N, 23°47′ E
429 m
CG-MO
CG-MO
CG-MA
CG-TO
CG-BC
02°48′ N, 22°16′ E
02°35′ N, 21°47′ E
03°58′ S, 15°59′ E
01°40′ N, 27°01′ E
05°24′ S, 12°54′ E
CG
02°36′ N, 29°34′ E
277 m
441 m
444 m
VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
Bongale
Bongeya River
Bongolo Mission
Bonzale
Bosobolo
Bosodula
(see: Boso-Dula)
Boso-Dula
(see also: Bosodula)
Bossainda
Boteka
(see also: Boteke, Flandria, Frandria)
Boteka
Boteke
(see: Boteka)
Boudewijnstad
(see: Moba)
Bozade
Brouwerij Baai
(see: Gisenyi)
Bubanza Province
Bufundi
Bugara
Bugarula, 6 km N
(see: Washiha Forest)
Bugarura Island
Bugesera
(see also: Kanazi)
Bugoye Kisenyi
Bugundi
Buhanga
Buhatu Valley
(see: Butahu Valley, Cave)
Buhonga
Buhoro
(see also: Mabaye, 10 km W)
Bujumbura
(see also: Usumbura)
Bujumbura, 60 km S
(see: Resha)
Bujumbura, 7 km NW
(see also: Bujumbura, 7 km NW)
Bujumbura-Mairie Province
Bujumbura-Rural Province
Bukama
Bukama Territory
Bukama, 40 mi [=64 km] below
(see: Katobwe)
Bukara
Bukavu
(see also: Costermansstad, Costermansville)
Bukavu, 100 km W
(see: Irangi Forest)
Bukeye Commune
Bukinanyana Commune
Bulaga
(see: Kasenga Road, km 36)
Bulaya
Buluku
Buluku River
Bulundu
Bumali
Bumba
Bumba Territory
Bunia
(see also: Ngioni)
Bunkeya
(see also: Bunkeye)
Bunkeye
(see: Bunkeya)
Bunyangura
Bunyenzi
Burundi
Bururi
Bururi Forest Reserve
(see also: Bururi, Réserve naturelle forestière de)
Bururi Province
06°36′ S, 16°46′ E
02°58′ N, 19°22′ E
04°11′ N, 19°54′ E
04°17′ N, 20°29′ E
CG-NU
04°17′ N, 20°29′ E
599 m
CG-BC
CG-ET
ca 05°15′ S, 14°52′ E
00°20′ S, 19°06′ E
333 m
CG-ET
CG-ET
00°19′ S, 19°07′ E
00°19′ S, 19°07′ E
337 m
337 m
CG-TG
07°03′ S, 29°47′ E
763 m
CG-SU
RW-OU
02°57′ N, 19°18′ E
01°42′ S, 29°15′ E
1481 m
BY
RW-NO
CG-NK
CG-SK
01°21′ S, 29°51′ E
01°10′ S, 29°28′ E
ca 02°00′ S, 29°04′ E
1200 m
1500 m
RW-OU
RW-ES
01°55′ S, 29°16′ E
02°09′ S, 30°05′ E
1432 m
RW-OU
RW-NO
BY-GI
CG-NK
01°42′ S, 29°35′ E
01°29′ S, 29°55′ E
03°39′ S, 29°59′ E
ca 00°20′ N, 29°49′ E
1828 m
2286 m
BY-BL
BY-CI
03°26′ S, 29°25′ E
02°42′ S, 29°11′ E
1529 m
1453 m
BY-BM
03°23′ S, 29°22′ E
794 m
BY-RU
03°54′ S, 29°24′ E
BY-BM
03°21′ S, 29°18′ E
BY
BY
CG-HL
CG-HL
CG-HL
09°12′ S, 25°51′ E
09°12′ S, 25°51′ E
08°54′ S, 26°04′ E
547 m
BY-MA
CG-SK
04°07′ S, 29°47′ E
02°30′ S, 28°52′ E
1498 m
CG-SK
01°54′ S, 28°27′ E
BY-MV
BY-CI
CG-HK
03°12′ S, 29°38′ E
02°53′ S, 29°20′ E
11°37′ S, 27°40′ E
CG-NK
CG-HU
CG-HU
CG-SK
CG-NK
CG-MO
CG-MO
CG-IT
01°12′ S, 29°27′ E
04°33′ N, 29°49′ E
04°33′ N, 29°49′ E
ca 02°42′ S, 27°20′ E
00°21′ N, 29°45′ E
02°11′ N, 22°28′ E
02°11′ N, 22°28′ E
01°34′ N, 30°15′ E
1332 m
CG-HK
10°24′ S, 26°58′ E
1008 m
CG-HK
10°24′ S, 26°58′ E
1008 m
CG-NK
CG-NK
BY
BY-BI
BY-BI
01°09′ S, 29°26′ E
01°10′ S, 29°27′ E
1200 m
1225 m
03°57′ S, 29°37′ E
03°56′ S, 29°36′ E
1836 m
BY
265
06°00′ S, 18°41′ E
CG-KU
CG-NK
CG-KG
CG-SU
CG-NU
CG-NU
1350 m
383 m
508 m
599 m
601 m
5000–5100
ft
1875 m
1543 m
1160 m
1300 m
419 m
European Journal of Taxonomy 382: 1–327 (2017)
Bururi, Réserve naturelle forestière de
(see: Bururi Forest Reserve)
Bushanganya
Bushdinka
(see: Busholinka River)
Bushokwe Caves
(see also: Bushokwe Grottes, Grottes de Bushokwe, Grottoes de Bushokwe)
Bushokwe Grottes
(see: Bushokwe Caves)
Busholinka River
(see also: Bushdinka)
Bushushu
Busiga
Busu Mokalu
(see: Busu-Mokala)
Busu-Mokala
(see also: Busu Mokalu)
Buta
Butagu Valley
(see: Butahu Valley, Cave)
Butahu Valley, Cave
(see also: Buhatu Valley, Butagu Valley, Butawu Valley, Butatu Valley)
Butare
(see also: Astrida, Butare INRS)
Butare INRS
(see: Butare)
Butatu Valley
(see: Butahu Valley, Cave)
Butawu Valley
(see: Butahu Valley, Cave)
Butembo
Butembo region
Buye Bala
Bwamanda [Mission]
Bwandza
Bweru
Bwirina
Byangore River
Byonga
Byumba
Camp Putnam
(see also: Putnam, Camp)
Cankuzo Province
Cape Cassa Mabongo
(see: Kasambongo)
Cape Lopez
Carrière Baya
(see: Baya quarry)
Carrière Pierkat
(see: Baya quarry)
Catchpole’s Farm
Cel I
(see: Bagbele)
Cel II
(see also: Cellule II)
Cellule I
(see: Bagbele)
Cellule II
(see: Cel II)
Chibambo
(see: Shibambo)
Chikapa
(see: Tshikapa)
Chinkolobwe
(see: Shinkolobwe)
Cibitoke
Cibitoke Province
Cinputu
(see also: Tschinputu)
Cocquilhatville
(see: Bandaka)
Cohoha, Lake
(see also: Lake Cohoha, Lac Cohoha)
Colline Giserama
(see: Giserama, Colline)
266
BY-BI
03°56′ S, 29°36′ E
CG-SK
CG-NK
02°19′ S, 28°47′ E
01°11′ S, 29°29′ E
CG-SK
ca 02°59′ S, 28°55′ E
CG-SK
ca 02°59′ S, 28°55′ E
CG-NK
01°11′ S, 29°29′ E
1350 m
CG-SK
BY-NG
CG-SU
02°01′ S, 28°54′ E
02°53′ S, 29°44′ E
02°20′ N, 20°15′ E
1455 m
1829 m
CG-SU
02°20′ N, 20°15′ E
CG-BU
CG-NK
02°48′ N, 24°44′ E
ca 00°20′ N, 29°49′ E
434 m
2286 m
CG-NK
ca 00°20′ N, 29°49′ E
2286 m
RW-SU
02°36′ S, 29°44′ E
1733 m
RW-SU
02°36′ S, 29°44′ E
1733 m
CG-NK
ca 00°20′ N, 29°49′ E
2286 m
CG-NK
ca 00°20′ N, 29°49′ E
2286 m
CG-NK
00°09′ N, 29°17′ E
CG-NK
CG
CG-SU
CG-MO
BY-RY
CG-NK
CG-NK
CG-SK
RW-NO
CG-IT
ca 00°09′ N, 29°17′ E
00°58′ S, 00°27′ E
03°10′ N, 19°15′ E
02°37′ N, 21°45′ E
03°19′ S, 30°28′ E
00°39′ N, 29°43′ E
1381–1760
m
03°21′ S, 28°08′ E
01°35′ S, 30°04′ E
01°24′ N, 28°35′ E
1350 m
508 m
1684 m
900 m
2300 m
675 m
BY
CG-SK
03°24′ S, 29°08′ E
CG-SK
CG-HK
03°07′ S, 28°43′ E
11°52′ S, 27°27′ E
1279 m
CG-HK
11°52′ S, 27°27′ E
1279 m
CG-IT
CG-HU
ca 01°28′ N, 30°04′ E
04°21′ N, 29°17′ E
1280 m
742 m
CG-HU
ca 03°55′ N, 29°22′ E
CG-HU
04°21′ N, 29°17′ E
CG-HU
03°55′ N, 29°25′ E
CG-HK
10°18′ S, 28°37′ E
928 m
CG-KS
06°25′ S, 20°48′ E
485 m
CG-LB
11°02′ S, 26°29′ E
1334 m
BY-CI
BY
CG-LL
02°53′ S, 27°07′ E
1020 m
05°22′ S, 22°21′ E
CG-ET
00°04′ N, 18°16′ E
BY-KI
02°30′ S, 30°07′ E
BY-CI
02°50′ S, 29°25′ E
742 m
306 m
2045 m
VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
Colline Gitovu
(see: Gitovu)
Colline Gosenyi
(see: Gisenyi)
Colline Kidaho
(see: Kidaho)
Colline Kwogofe
(see: Kwogofe, Colline)
Colline Mugogwe
(see: Mugogwe)
Colline Murunda
(see: Muhazi)
Colline Musanze
(see: Musanze)
Colline Musha
(see: Musha)
Colline Nyakibanda
(see: Nyakibanda)
Colline Nyamata
(see: Nyamata)
Colline Rubona
(see: Rubona)
Colline Shangi
(see: Shangi)
Colline Shyanda
(see: Shyanda)
Congo (Democratic Republic of the)
Congo da Lemba
Congo River, 75 mi [=121 km] up
Congo-Nil-Aka
Coquilhalville
(see: Bandaka)
Coquilhatstad
(see: Bandaka)
Coquilhatville
(see: Bandaka)
Costermansstad
(see: Bukavu)
Costermansville
(see: Bukavu)
Crevettes Cave
Cyangugu
(see: Shangugu)
Cyili
(see also: Cyili Ntyano)
Cyili Ntyano
(see: Cyili)
Cyohaha, Lake
(see also: Tshyohaha, Lake)
Dambi
(see: Ibambi)
Dedegwa River
Defrenne Cave
(see: Mwanga Cave)
Defrenne Grot
(see: Mwanga Cave)
Demba Territory
Dembelenge
(see: Dimbelenge)
Dembelenge Territory
Dethioux Farm, Cave I
(see: Shinkolobwe)
Dethioux Farm, Cave II
(see: Shinkolobwe)
Dethioux Farm, Tantara Cave
(see: Shinkolobwe)
Dethioux Hoeve, Grot I
(see: Shinkolobwe)
Dethioux Hoeve, Grot II
(see: Shinkolobwe)
Dethioux Hoeve, Tantara Grot
(see: Shinkolobwe)
Dethiox Caves
(see: Kambove)
Dethiox Grotten
(see: Kambove)
267
RW-SU
02°19′ S, 29°09′ E
1649 m
RW-OU
01°42′ S, 29°15′ E
1481 m
RW-NO
01°22′ S, 29°45′ E
1944 m
BY-CI
02°50′ S, 29°25′ E
RW-SU
02°31′ S, 29°49′ E
RW-OU
01°54′ S, 29°22′ E
RW-NO
01°29′ S, 29°36′ E
RW-SU
02°32′ S, 29°52′ E
RW-SU
02°38′ S, 29°41′ E
RW-ES
02°10′ S, 30°06′ E
RW-SU
02°19′ S, 29°56′ E
RW-OU
02°24′ S, 29°00′ E
RW-SU
02°33′ S, 29°48′ E
1711 m
2067 m
1346 m
1646 m
CG
CG-KU
CG-BC
CG-HU
CG-ET
04°34′ N, 29°45′ E
00°04′ N, 18°16′ E
306 m
CG-ET
00°04′ N, 18°16′ E
306 m
CG-ET
00°04′ N, 18°16′ E
306 m
CG-SK
02°30′ S, 28°52′ E
1498 m
CG-SK
02°30′ S, 28°52′ E
1498 m
CG-HK
RW-OU
10°58′ S, 26°47′ E
02°29′ S, 28°54′ E
1537 m
RW-SU
02°30′ S, 29°46′ E
RW-SU
02°30′ S, 29°46′ E
RW-ES
02°15′ S, 30°00′ E
CG-HU
02°22′ N, 27°37′ E
CG-HU
CG-HK
04°35′ N, 29°43′ E
10°58′ S, 26°39′ E
CG-HK
10°58′ S, 26°39′ E
CG-LL
CG-LL
05°30′ S, 22°16′ E
05°33′ S, 23°07′ E
682 m
CG-LL
CG-LB
11°02′ S, 26°29′ E
1334 m
CG-LB
11°02′ S, 26°29′ E
1334 m
CG-LB
11°02′ S, 26°29′ E
1334 m
CG-LB
11°02′ S, 26°29′ E
1334 m
CG-LB
11°02′ S, 26°29′ E
1334 m
CG-LB
11°02′ S, 26°29′ E
1334 m
CG-LB
10°50′ S, 26°39′ E
CG-LB
10°50′ S, 26°39′ E
05°42′ S, 13°40′ E
687 m
European Journal of Taxonomy 382: 1–327 (2017)
Detioux Cave
(see: Kambove)
Digba
Dika
Dikul River
(see: Dikulwe Cave)
Dikuluwe Cave
(see: Dikulwe Cave)
Dikuluwe Grot
(see: Dikulwe Cave)
Dikulwe Cave
(see also: Dikuluwe Cave, Dikuluwe Grot, Grotte de Dikuluwe, Dikul River)
Dikulwe River
Dilolo
(see also: Kayombe, Dilolo Station, Dilolo, Poste)
Dilolo Station
(see: Dilolo)
Dilolo Territory
Dilolo, Poste
(see: Dilolo)
Dimbelenge
(see also: Dembelenge)
Djabir
Djamba
Djambala (Hill)
Djeka
Djelube River, lower
Djelube River, middle
Djemba
(see also: Djomba)
Djomba
(see: Djemba)
Djugu Bos
(see: Djugu Forest)
Djugu Forest
(see also: Djugu Bos, Djugu, forêt de, Forêt de Djugu)
Djugu Territory
Djugu Territory
Djugu, forêt de
(see: Djugu Forest)
Djuma River, upper
Djumma River, middle
Dramba
Drodo
(see: Drodro)
Drodro
(see also: Drodo)
Dubulu
Duma
Dungu
Durba
Eala
East Kasai (former) Province
(see: Kasai Oriental (former) Province)
East Kasai Province
(see: Kasai Oriental Province)
Eastern Congo
Eastern Prefecture
(see also: Kibungo (former) Prefecture)
Edouard, Lac
(see: Edward, Lake)
Edward Meer
(see: Edward, Lake)
Edward, Lake
(see also: Lake Edward, Lac Edouard, Edouard, Lac, Albert Edward Nyanza,
Edward Meer, Eward Nyanza, Lake Idi Amin Dada, Idi Amin Dada, Lake,
Lac Idi Amine, Lake Rutanzige, Rutanzige, Lake)
Eidolon land
Elila
Elisabetha
(see also: Elizabetha)
Elisabethstad
(see: Lubumbashi)
Elisabethville
(see: Lubumbashi)
Elizabetha
(see: Elisabetha)
268
CG-LB
10°50′ S, 26°39′ E
CG-BU
CG-HU
CG-LB
04°24′ N, 25°47′ E
04°22′ N, 27°40′ E
10°45′ S, 25°22′ E
CG-LB
10°45′ S, 25°22′ E
CG-LB
10°45′ S, 25°22′ E
CG-LB
10°45′ S, 25°22′ E
CG-LB
CG-LB
10°45′ S, 25°22′ E
10°28′ S, 22°28′ E
1069 m
CG-LB
10°28′ S, 22°28′ E
1069 m
CG-LB
CG-LB
10°28′ S, 22°28′ E
1069 m
CG-LL
05°33′ S, 23°07′ E
682 m
CG-TO
CG-BU
CG-BC
CG-SN
CG-NK
CG-NK
CG-NK
00°32′ N, 24°05′ E
02°53′ N, 24°02′ E
04°42′ S, 13°44′ E
02°47′ S, 23°55′ E
00°35′ N, 29°40′ E
00°31′ N, 29°44′ E
01°20′ S, 29°34′ E
437 m
419 m
850 m
1100 m
1500 m
CG-NK
01°20′ S, 29°34′ E
1500 m
CG-IT
01°55′ N, 30°30′ E
1530 m
CG-IT
01°55′ N, 30°30′ E
1530 m
CG-IT
CG-IT
CG-IT
01°55′ N, 30°30′ E
01°55′ N, 30°30′ E
01°55′ N, 30°30′ E
1530 m
CG-NK
CG-NK
CG-HU
CG-IT
00°38′ N, 29°34′ E
00°42′ N, 29°41′ E
03°40′ N, 30°19′ E
01°46′ N, 30°32′ E
850 m
800 m
877 m
1854 m
CG-IT
01°46′ N, 30°32′ E
1854 m
CG-NU
CG-SU
CG-HU
CG-HU
CG-ET
CG
04°18′ N, 20°16′ E
03°53′ N, 18°41′ E
03°37′ N, 28°34′ E
03°08′ N, 29°35′ E
00°04′ N, 18°17′ E
613 m
356 m
669 m
629 m
689 m
301 m
CG
CG
RW
CG-NK
ca 00°08′ S, 29°35′ E
CG-NK
ca 00°08′ S, 29°35′ E
CG-NK
ca 00°08′ S, 29°35′ E
CG-HK
CG-TO
CG-TO
ca 10°30′ S, 27°30′ E
02°43′ S, 25°53′ E
01°09′ N, 23°36′ E
486 m
428 m
CG-HK
11°40′ S, 27°26′ E
1209 m
CG-HK
11°40′ S, 27°26′ E
1209 m
CG-TO
01°09′ N, 23°36′ E
428 m
VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
Elizabethville
(see: Lubumbashi)
Embe
Epulu
Epulu, 13 km E
Epulu, 2 km W
Epulu, 6 km S, 2 km W
Equateur (former) Province
(see also: Evenaarsprovincie)
Equateur Province
Eringete
(see: Eringite)
Eringite
(see also: Eringete, Iringite)
Etembo
Evenaarsprovincie
(see: Equateur (former) Province)
E’ville
(see: Lubumbashi)
Eward Nyanza
(see: Edward, Lake)
Faradje
(see also: Irumu, 241 km N)
Faradje Territory
(see also: Faradje, Zone)
Faradje, Zone
(see: Faradje Territory)
Farm ‘des Grands Bois’
(see: Likasi)
Fataki
Fataki area
Ferme Wasseige
(see: Wasseige Farm)
Flandria
(see: Boteka)
FomuLac-Katana
Forêt de Djugu
(see: Djugu Forest)
Forêt d’Irangi
(see: Irangi Forest)
Forêt Naturelle de Nyungwe
(see: Nyungwe Forest Reserve)
Fort Beni
(see: Vieux Beni)
François-Joseph, Chutes
(see: Franz-Josef Falls)
Frandria
(see: Boteka)
Franz-Josef Falls
(see also: François-Joseph, Chutes)
Fuko
Fuko (River)
Funda Biabo
(see also: Funda Biako)
Funda Biako
(see: Funda Biabo)
Fundi
Fundi
Fwa Meer
(see: Fwa, Lake)
Fwa, Lake
(see also: Fwa Meer, Lac Fwa, Lake Fwa)
Gabiro
Gafumba
Gafumba Mugusa
Gafunzo
Gafunzo Kigaga
Gahama
(see also: Kahama)
Gahinga (Vulcano)
Gakoma (Mission)
Gala
(see: Gumba)
Gamangui
Ganda
(see: Ganda-Sundi)
Gandajika
269
CG-HK
11°40′ S, 27°28′ E
1208 m
CG-HU
CG-IT
CG-IT
CG-IT
CG-IT
CG
04°40′ N, 29°31′ E
01°24′ N, 28°35′ E
ca 01°24′ N, 28°35′ E
ca 01°24′ N, 28°35′ E
ca 01°24′ N, 28°35′ E
750 m
750 m
750 m
750 m
CG
CG-NK
00°51′ N, 29°37′ E
1270 m
CG-NK
00°51′ N, 29°37′ E
1270 m
CG-IT
CG
00°55′ N, 28°39′ E
832 m
CG-HK
11°40′ S, 27°26′ E
1209 m
CG-NK
ca 00°08′ S, 29°35′ E
CG-HU
03°44′ N, 29°43′ E
CG-HU
03°44′ N, 29°43′ E
CG-HU
03°44′ N, 29°43′ E
CG-HK
10°59′ S, 26°44′ E
1318 m
CG-IT
CG-IT
CG-HK
02°00′ N, 30°34′ E
02°01′ N, 30°38′ E
11°50′ S, 27°20′ E
1611 m
ca 1750 m
CG-ET
00°19′ S, 19°07′ E
337 m
CG-SK
CG-IT
02°13′ S, 28°50′ E
01°55′ N, 30°30′ E
1530 m
CG-SK
01°54′ S, 28°27′ E
RW-OU
02°31′ S, 29°18′ E
5000–5100
ft
2500 m
CG-NK
00°26′ N, 29°33′ E
585 m
CG-KG
ca 07°36′ S, 17°14′ E
CG-ET
00°19′ S, 19°07′ E
CG-KG
ca 07°36′ S, 17°14′ E
CG-NK
CG-NK
CG-LB
01°15′ S, 29°28′ E
01°15′ S, 29°28′ E
09°50′ S, 25°33′ E
1300 m
1300 m
693 m
CG-LB
09°50′ S, 25°33′ E
693 m
CG-TO
CG-IT
CG-LL
00°52′ N, 27°34′ E
02°06′ N, 30°46′ E
05°43′ S, 23°25′ E
1942 m
CG-LL
05°43′ S, 23°25′ E
RW-ES
RW-SU
RW-SU
RW-OU
RW-OU
CG-NK
01°31′ S, 30°28′ E
02°28′ S, 29°50′ E
02°28′ S, 29°50′ E
02°24′ S, 29°03′ E
02°24′ S, 29°03′ E
01°13′ S, 29°28′ E
1457 m
1457 m
1250 m
RW-NO
RW-SU
CG-MO
01°24′ S, 29°38′ E
02°29′ S, 29°55′ E
02°57′ N, 21°26′ E
412 m
CG-HU
CG-BC
02°10′ N, 27°15′ E
04°52′ S, 12°52′ E
600 m
294 m
CG-LM
06°45′ S, 23°57′ E
806 m
775 m
337 m
1400 m
European Journal of Taxonomy 382: 1–327 (2017)
Ganda-Sundi
(see also: Ganda)
Gandu
Gangala
(see: Gangala-na-Bodio)
Gangala na Bodio
(see: Gangala-na-Bodio)
Gangala-na-Bodio
(see also: Gangala, Gangara, Gangara na Bodio, Gangala na Bodio)
Gangara
(see: Gangala-na-Bodio)
Gangara na Bodio
(see: Gangala-na-Bodio)
Ganza
Garamba National Park
(see also: Parc National de Garamba)
Gasiza
Gayu [River]
(see: Ngayu [River])
Gemena
Gemena Territory
Geramihasha River
Gifuruzi
(see: Kifuruzi)
Gihanga
(see also: Kihanga)
Giheta
Gihinda
(see: Gihinga)
Gihinga
(see also: Gihinda)
Gisenyi
(see also: Colline Gosenyi, Gosenyi, Brouwerij Baai)
Gisenyi
(see also: Kisenyi, Kisenyi, Road to Sake, Kissenji)
Giserama, Colline
(see also: Colline Giserama, Kirumura, 1.1 - 2.1 km N, 0.8 - 1.1 km E)
Gisovu
Gitega
(see also: Kitega)
Gitega Province
Gitovu
(see also: Colline Gitovu)
Goma
(see also: Ngoma)
Gombe
(see: Ngombe)
Goroumbwa Mine
Gosenyi
(see: Gisenyi)
Grand Ebouli de Lemera
(see: Itombwe)
Grotte d’Ango
(see: Ango Cave)
Grotte de Banza Ninda
(see: Banza Ninda Caves)
Grotte de Baya
(see: Baya Cave)
Grotte de Defrenne
(see: Mwanga Cave)
Grotte de Dikuluwe
(see: Dikulwe Cave)
Grotte de Kaboyaboya
(see: Kaboyaboya Cave)
Grotte de Kakanda
(see: Kakanda Cave)
Grotte de Kakindo
(see: Kakondo Cave)
Grotte de Kakondo
(see: Kakondo Cave)
Grotte de Kakontwe
(see: Kakontwe Cave)
Grotte de Kalomoni
(see: Kakontwe)
Grotte de Kalumbu
(see: Kalumbu Cave)
270
CG-BC
04°52′ S, 12°52′ E
294 m
CG
CG-HU
03°41′ N, 29°08′ E
701 m
CG-HU
03°41′ N, 29°08′ E
701 m
CG-HU
03°41′ N, 29°08′ E
701 m
CG-HU
03°41′ N, 29°08′ E
701 m
CG-HU
03°41′ N, 29°08′ E
701 m
CG-HL
CG-HU
09°14′ S, 26°42′ E
04°10′ N, 29°30′ E
860 m
RW
CG-HU
01°35′ N, 27°13′ E
CG-SU
CG-SU
CG-NK
BY-MA
03°15′ N, 19°46′ E
03°15′ N, 19°46′ E
01°13′ S, 29°27′ E
04°19′ S, 29°35′ E
BY-BB
03°12′ S, 29°18′ E
814 m
BY
RW-ES
03°20′ S, 29°50′ E
01°24′ S, 30°37′ E
1400 m
RW-ES
01°24′ S, 30°37′ E
1400 m
RW-OU
01°42′ S, 29°15′ E
1481 m
RW-OU
01°41′ S, 29°15′ E
1450 m
BY-CI
02°50′ S, 29°25′ E
2045 m
RW-SU
BY-GI
02°15′ S, 29°20′ E
03°26′ S, 29°56′ E
2335 m
1845 m
BY
RW-SU
02°19′ S, 29°09′ E
1649 m
CG-NK
01°41′ S, 29°13′ E
1499 m
CG-KS
06°35′ S, 20°42′ E
528 m
CG-HU
RW-OU
03°07′ N, 29°34′ E
01°42′ S, 29°15′ E
1481 m
CG-SK
04°04′ S, 28°40′ E
CG-BU
04°02′ N, 25°52′ E
CG-BC
05°16′ S, 14°52′ E
CG-HK
11°52′ S, 27°27′ E
CG-HK
10°58′ S, 26°39′ E
CG-LB
10°45′ S, 25°22′ E
CG-LB
09°56′ S, 25°58′ E
CG-LB
10°47′ S, 26°36′ E
CG-SK
02°16′ S, 28°51′ E
CG-SK
02°16′ S, 28°51′ E
CG-HK
10°59′ S, 26°38′ E
1294 m
CG-HK
10°59′ S, 26°40′ E
1294 m
CG-HK
10°51′ S, 26°39′ E
555 m
1250 m
610 m
1279 m
1438 m
VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
Grotte de Kando
(see: Kando Cave)
Grotte de Kasoma
(see: Kasoma Cave)
Grotte de Kasowena
(see: Kasowena Cave)
Grotte de Katana
(see: Mahyusha Cave)
Grotte de Kele 1
(see: Kele Cave 1)
Grotte de Kiasala
(see: Kiasala Cave)
Grotte de Kibwe Lya Mikako
(see: Kibwe Lya Mikako Cave)
Grotte de Kimuenza
(see: Kimuenza Cave)
Grotte de Kiwakishi
(see: Kiamokoto)
Grotte de Kondo
(see: Kakanda Cave)
Grotte de Kwama Kwama
(see: Kwama Kwama Cave)
Grotte de Kyantapo
(see: Kyantapo Cave)
Grotte de Kyasala
(see: Kiasala Cave)
Grotte de Lubuka
(see: Lubuka Cave)
Grotte de Lukatu
(see: Ntoto Cave)
Grotte de Lusolo
(see: Lusolo Grot)
Grotte de Mahysha
(see: Mahyusha Cave)
Grotte de Mahyusha
(see: Mahyusha Cave)
Grotte de Mani ou Grotte de Nkela de Mbumba-Nsi
(see: Mani Cave or Nkela Cave or Mbumba-Nsi)
Grotte de Matupi
(see: Matupi Cave)
Grotte de M’baya
(see: Baya Cave)
Grotte de Montembessa
(see: Montembessa Cave)
Grotte de Mpopola
(see: Mpopola Cave)
Grotte de Mugaba
(see: Mugaba Cave)
Grotte de Mulonga
(see: Mulonga Cave)
Grotte de Mulungwishi
(see: Mulungwishi, Cave)
Grotte de Mwanakusu
(see: Mwanakusu Cave)
Grotte de Mwanga
(see: Mwanga Cave)
Grotte de Pempere
(see: Pempere Cave)
Grotte de Ribue Lya Mikako
(see: Kibwe Lya Mikako Cave)
Grotte de Rubindi
(see: Rubindi)
Grotte de Saga Saga
(see: Saga Saga Cave)
Grotte de Tantara, Ferme Dethioux
(see: Shinkolobwe)
Grotte de Tshamalenge
(see: Tshamalenge Cave)
Grotte de Yolohotiri
(see: Yolohotiri Cave)
Grotte I, Ferme Dethioux
(see: Shinkolobwe)
Grotte II, Ferme Dethioux
(see: Shinkolobwe)
Grotte Swanepoel I
(see: Swanepoel I Cave)
CG-HK
10°49′ S, 26°07′ E
CG-HL
09°25′ S, 26°37′ E
CG-HK
10°58′ S, 26°39′ E
CG-SK
02°13′ S, 28°50′ E
CG-BC
05°31′ S, 14°36′ E
CG-LB
09°56′ S, 25°58′ E
1300 m
CG-NK
00°15′ S, 29°01′ E
1500 m
CG-BC
04°28′ S, 15°17′ E
CG-HL
09°10′ S, 27°04′ E
900 m
CG-LB
10°47′ S, 26°36′ E
1438 m
CG-NK
01°19′ N, 29°52′ E
1200 m
CG-LB
09°56′ S, 25°58′ E
1525 m
CG-LB
09°56′ S, 25°58′ E
1300 m
CG-SK
03°12′ S, 28°51′ E
CG-TG
06°59′ S, 29°43′ E
CG-HK
09°56′ S, 25°28′ E
CG-SK
02°13′ S, 28°50′ E
1480 m
CG-SK
02°13′ S, 28°50′ E
1480 m
CG-IT
01°16′ N, 29°55′ E
1067 m
CG-HK
11°52′ S, 27°27′ E
1279 m
CG
09°56′ S, 25°58′ E
1450 m
CG-SK
ca 02°19′ S, 28°31′ E
CG-LB
09°56′ S, 25°58′ E
CG-LB
10°47′ S, 26°38′ E
CG-MN
04°35′ S, 27°08′ E
CG-HK
10°58′ S, 26°39′ E
CG-HK
10°58′ S, 26°47′ E
CG-NK
00°15′ S, 29°01′ E
1500 m
BY-MA
04°16′ S, 29°35′ E
997 m
CG-IT
01°19′ N, 29°52′ E
1160 m
CG-LB
11°02′ S, 26°29′ E
1334 m
CG-HK
11°46′ S, 26°30′ E
CG-IT
01°13′ N, 29°49′ E
CG-LB
11°02′ S, 26°29′ E
1334 m
CG-LB
11°02′ S, 26°29′ E
1334 m
CG-LB
10°50′ S, 26°39′ E
ca 1350 m
1480 m
827 m
CG
CG
271
1366 m
European Journal of Taxonomy 382: 1–327 (2017)
Grottes de Banjan
(see: Banjan Caves)
Grottes de Bushokwe
(see: Bushokwe Caves)
Grottes de Dethiox
(see: Kambove)
Grottes Kiwakishi
Grottoes de Bushokwe
(see: Bushokwe Caves)
Gumba
(see also: Gala)
Gundu, mount
(see also: Mount Gundu)
Haut Katanga Province
(see also: Upper Katanga Province)
Haut Lomami Province
(see also: Upper Lomani Province)
Haut Tshuapa
(see: Tshuapa River, upper)
Haut Uele Province
(see also: Upper Uele Province)
Haute Congo
Haute Zaire
Homa, mont
(see: Hoyo, mount)
Homa, Mount
(see also: Mount Homa)
How…
Hoyo, mount
(see also: Homa, mont, Mount Homa, Mount Hoyo, Mont Hoyo)
Ibambi
(see also: Dambi, Idambi)
Ibebero
Ibembo
Idambi
(see: Ibambi)
Idi Amin Dada, Lake
(see: Edward, Lake)
Idjwi, Island
(see also: Ïle Idjwi)
Ihema, Lake
(see also: Lac Ihema, Lake Ihema)
Ihunga-Semliki conluence
Ijenda Mission
Ikange
(see: Ikengo)
Ikela
Ikela Territory
Ikenge
(see: Ikengo)
Ikengo
(see also: Ikange, Ikenge)
Île de Kongolo
(see: Kongolo, Island)
Île Idjwi
(see: Idjwi, Island)
Île Mai
(see: Mai, Island)
Île Mbiye
(see: Mbiye, Island)
Ilimba
Imbo
Imbonga
Inakiluba
(see also: Nakiluba, Nakilubo)
Inangongo
Inimvua
Inkongo
(see: Inkongo-Kakese)
Inkongo-Kakese
(see also: Inkongo, Inkongu)
Inkongu
(see: Inkongo-Kakese)
Inongo
Irangi
(see also: Irangi, slightly E of)
272
CG-BC
05°16′ S, 14°53′ E
CG-SK
ca 02°59′ S, 28°55′ E
CG-LB
10°50′ S, 26°39′ E
CG-HK
CG-SK
09°00′ S, 27°12′ E
ca 02°59′ S, 28°55′ E
CG-MO
02°57′ N, 21°26′ E
412 m
CG-BU
CG
CG
CG-TP
01°00′ S, 23°02′ E
CG
CG
CG
CG-IT
01°13′ N, 29°49′ E
CG-IT
01°13′ N, 29°49′ E
CG
CG-IT
01°13′ N, 29°49′ E
1200 m
CG-HU
02°22′ N, 27°37′ E
687 m
CG-SK
CG-BU
CG-HU
02°38′ N, 23°37′ E
02°22′ N, 27°37′ E
420 m
687 m
CG-NK
ca 00°08′ S, 29°35′ E
CG-SK
02°09′ S, 29°04′ E
RW-ES
01°52′ S, 30°47′ E
CG-NK
BY-BI
CG-ET
00°10′ N, 29°35′ E
03°39′ S, 29°34′ E
00°08′ S, 18°08′ E
CG-TP
CG-TP
CG-ET
01°11′ S, 23°16′ E
01°11′ S, 23°16′ E
00°08′ S, 18°08′ E
301 m
CG-ET
00°08′ S, 18°08′ E
301 m
CG-TO
00°03′ N, 25°12′ E
CG-SK
02°09′ S, 29°04′ E
CG-TO
00°28′ N, 25°17′ E
CG-TO
00°28′ N, 25°17′ E
413 m
CG-NK
BY-BB
CG-ET
CG-HK
00°50′ N, 29°58′ E
03°13′ S, 29°20′ E
00°43′ S, 19°46′ E
11°34′ S, 27°19′ E
750 m
364 m
1280 m
CG-NK
CG-HU
CG-SN
00°33′ N, 29°41′ E
04°35′ N, 29°43′ E
04°53′ S, 23°18′ E
800 m
1061 m
400 m
CG-SN
04°53′ S, 23°18′ E
400 m
CG-SN
04°53′ S, 23°18′ E
400 m
CG-MA
CG-SK
01°57′ S, 18°16′ E
01°54′ S, 28°27′ E
335 m
1200 m
1600 m
301 m
425 m
1600 m
VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
Irangi Bos
(see: Irangi Forest)
Irangi Forest
(see also: Bukavu, 100 km W, Forêt d’Irangi, Irangi Bos)
Irangi, 10 km from
(see: Tshambutschu)
Irangi, 2 mi [=3.2 km] from
(see: Kasiba)
Irangi, 3 mi [=4.8 km] from
Irangi, slightly E of
(see: Irangi)
Irebu
Iringi
Iringite
(see: Eringite)
Irsac Labo
(see: Luiro)
Irumu
Irumu
Irumu, 20 mi [=32 km] S
(see: Mount Hoyo Caves)
Irumu, 241 km N
(see: Faradje)
Irumu, 25 km W
(see: Ituri Forest)
Irumu, 40 mi [=64 km] SW
(see: Ituri Forest)
Irumu, 40 mi SW
Irumu, 50 mi SW
Isai River
Isato
Ishangi
(see: Shangi)
Ishango River
Ishango-Nyakazi
Isirio
(see: Isiro)
Isiro
(see also: Isirio, Paulis)
Iso
(see also: Isoberlinia, Iso.4)
Iso.4
(see: Iso)
Isoberlinia
(see: Iso)
Isoga
Issehe River
Itombwe
(see also: Lemera Great Sink, Grand Ebouli de Lemera)
Ituri Forest
(see also: Irumu, 40 mi [=64 km] SW, Sturi Forest)
Ituri Forest
(see also: Irumu, 25 km W)
Ituri ht
Ituri Province
Ituri River
Ituri River, Bridge
Ituri, south
Iyonda
Jadotstad
(see: Likasi)
Jadotstad, 15 km S
(see: Mwanga Cave)
Jadotsville
(see: Likasi)
Jadotville
(see: Likasi)
Jadotville, 15 km S
(see: Mwanga Cave)
Jadotville, ca 8 km S
(see: Kakontwe)
Jadoville
(see: Likasi)
Jaima
Jiba
CG-SK
01°54′ S, 28°27′ E
CG-SK
01°54′ S, 28°27′ E
CG-SK
ca 01°58′ S, 28°30′ E
CG-SK
ca 01°54′ S, 28°27′ E
CG-SK
CG-SK
ca 01°54′ S, 28°27′ E
ca 01°54′ S, 28°27′ E
CG-ET
CG
CG-NK
00°37′ S, 17°45′ E
01°54′ S, 28°27′ E
00°51′ N, 29°37′ E
1270 m
CG-SK
02°14′ S, 28°48′ E
1646 m
CG-IT
01°27′ N, 29°52′ E
CG
CG-IT
01°27′ N, 29°52′ E
01°13′ N, 29°49′ E
841–1280
m
841 m
1200 m
CG-HU
03°44′ N, 29°43′ E
775 m
CG-IT
01°23′ N, 28°38′ E
CG-IT
01°00′ N, 29°16′ E
CG-IT
CG-IT
CG-IT
CG-SU
RW-OU
01°00′ N, 29°16′ E
01°24′ N, 28°04′ E
03°18′ N, 18°39′ E
02°24′ S, 29°00′ E
CG-NK
CG-NK
CG-HU
00°08′ S, 29°36′ E
00°08′ S, 29°35′ E
02°46′ N, 27°37′ E
950 m
950 m
730 m
CG-HU
02°46′ N, 27°37′ E
730 m
CG-HU
04°18′ N, 29°56′ E
CG-HU
04°18′ N, 29°56′ E
CG-HU
04°18′ N, 29°56′ E
CG-SK
CG-IT
CG-SK
03°20′ S, 28°12′ E
01°05′ N, 29°49′ E
04°04′ S, 28°40′ E
CG-IT
01°00′ N, 29°16′ E
CG-IT
01°23′ N, 29°38′ E
CG-IT
CG
CG-IT
CG-IT
CG-IT
CG-ET
CG-HK
ca 01°20′ N, 27°35′ E
01°20′ N, 27°35′ E
01°00′ N, 29°16′ E
00°02′ S, 18°12′ E
10°59′ S, 26°44′ E
1318 m
CG-HK
10°59′ S, 26°44′ E
1318 m
CG-HK
10°59′ S, 26°44′ E
1318 m
CG-HK
10°59′ S, 26°40′ E
1253 m
CG-HK
10°59′ S, 26°44′ E
1318 m
CG
CG-IT
01°52′ N, 30°42′ E
2123 m
5000–5100
ft
5000–5100
ft
323 m
1266 m
1200 m
CG-HK
CG-HK
273
European Journal of Taxonomy 382: 1–327 (2017)
Jiba Risasi
CG-IT
01°52′ N, 30°42′ E
1800 m
CG-IT
01°52′ N, 30°42′ E
1800 m
CG-IT
01°20′ N, 29°50′ E
1030 m
CG-NK
CG-NK
CG-NK
01°17′ S, 29°33′ E
00°27′ N, 29°44′ E
00°38′ S, 29°07′ E
1500 m
1791 m
CG-SK
CG-TG
CG-TG
CG-KS
CG-NK
CG-HL
02°30′ S, 28°47′ E
06°03′ S, 26°55′ E
06°03′ S, 26°55′ E
05°44′ S, 20°49′ E
00°51′ N, 29°39′ E
09°01′ S, 26°45′ E
CG-LM
CG-SK
CG-NK
06°40′ S, 23°40′ E
02°28′ S, 28°49′ E
00°38′ S, 29°07′ E
2044 m
1791 m
CG-IT
ca 01°13′ N, 29°49′ E
1200 m
CG-HL
09°01′ S, 26°45′ E
788 m
CG-HL
CG-KO
CG
09°08′ S, 26°16′ E
06°08′ S, 24°29′ E
732 m
842 m
CG-KO
CG-KO
06°08′ S, 24°29′ E
05°11′ S, 23°51′ E
CG-SK
CG
CG-SK
02°03′ S, 28°55′ E
1519 m
04°59′ S, 29°05′ E
1950 m
CG-TG
05°04′ S, 29°00′ E
2440 m
CG-HL
CG-LB
07°19′ S, 25°35′ E
09°56′ S, 25°58′ E
918 m
CG-LB
09°56′ S, 25°58′ E
CG-LL
06°12′ S, 22°23′ E
610 m
CG-NK
CG-HL
CG-SK
01°09′ S, 29°29′ E
08°16′ S, 26°36′ E
02°09′ S, 28°54′ E
1250 m
556 m
1455 m
CG-SK
02°09′ S, 28°54′ E
1455 m
CG-TG
CG-MO
RW-ES
08°22′ S, 29°15′ E
03°11′ S, 28°20′ E
01°30′ S, 30°30′ E
1800 m
CG-NK
01°13′ S, 29°28′ E
1250 m
CG
CG
CG-SK
CG-MN
CG-SK
02°00′ S, 28°45′ E
02°38′ S, 26°06′ E
02°09′ S, 28°54′ E
602 m
1455 m
CG-SK
02°09′ S, 28°54′ E
1455 m
CG-SK
CG-LB
03°10′ S, 28°21′ E
10°47′ S, 26°36′ E
1525 m
1438 m
CG-LB
10°47′ S, 26°36′ E
1438 m
CG-LB
CG-SK
10°44′ S, 26°23′ E
02°16′ S, 28°51′ E
CG-SK
02°16′ S, 28°51′ E
RW-ES
01°29′ S, 30°17′ E
(see also: Jiba Trisasi)
Jiba Trisasi
Jolohairi
(see: Jiba Risasi)
(see: Yolokairi)
Jomba
Kabakuli River
Kabale
(see also: Kabare)
Kabale
Kabalo
Kabalo Territory
Kabambaie [River]
Kabambi
Kabanga
(see: Kabenge)
Kabanga
Kabare
Kabare
(see: Kabale)
Kabatibi Cave
(see: Mount Hoyo Caves)
Kabenge
(see also: Kabanga)
Kabengere
Kabinda
Kabinda (former) district
(see: Lomami Province)
Kabinda Territory
Kabinda, 129 km NNW
(see: Kaseu)
Kabira
Kabive, 50 km SW
Kabobo Forest
(see also: Talama, 4 km SW)
Kabobo, mount
(see also: Mount Kabobo)
Kabongo
Kaboyaboya Cave
(see also: Grotte de Kaboyaboya, Kaboyaboya Grot)
Kaboyaboya Grot
(see: Kaboyaboya Cave)
Kabwe
(see also: Kananga, 50 km SW, Luluaburg, 50 km SW)
Kadehe River
Kadia
Kadjidji
(see: Kajuju)
Kadjudju
(see: Kajuju)
Kafwe
Kagada
Kagera National Park
(see: Akagera Park)
Kahama
(see: Gahama)
Kahitumba (Mutaka)
Kahuzi
Kahuzi-Biega National Park
Kailo
Kajuchu
(see: Kajuju)
Kajuju
(see also: Kadjudju, Kajuchu, Kadjidji)
Kakanda
Kakanda Cave
(see also: Grotte de Kakanda, Grotte de Kondo, Kakanda Grot, Kondo Cave,
Kondo Grot)
Kakanda Grot
(see: Kakanda Cave)
Kakanda Mutaka
Kakindo Cave
(see: Kakondo Cave)
Kakindo Grotte
(see: Kakondo Cave)
Kakitumba
274
2044 m
530 m
1260 m
788 m
1400 m
VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
Kakodo
Kakoi
Kakonda
Kakonde
Kakondo
Kakondo Cave
(see also: Grotte de Kakondo, Kakondo Grot, Kakindo Cave, Kakindo
Grotte, Grotte de Kakindo)
Kakondo Grot
(see: Kakondo Cave)
Kakontwa
(see: Kakontwe)
Kakontwe
(see also: Grotte de Kalomoni, Kakontwa, Kakontwe Camp, Kalomoni Cave,
Kalomoni Grot, Likasi, ca 8 km S, Jadotville, ca 8 km S)
Kakontwe Camp
(see: Kakontwe)
Kakontwe Cave
(see also: Grotte de Kakontwe, Kakontwe Grot)
Kakontwe Grot
(see: Kakontwe Cave)
Kakungu
Kakyelo
Kalanda
Kalehe Territory
Kalemi
(see: Kalemie)
Kalemie
(see also: Albertstad, Albertville, Kalemi)
Kalemie Territory
Kalemie-Moba (Road?)
(see also: Albertville-Moba (Road?))
Kalenda
Kalenda [Kasai]
Kalikimvua
Kalimabenge River
Kalina
Kaliza [River]
(see: Kiliza [River])
Kaloba, 5 km from
Kalomoni Cave
(see: Kakontwe)
Kalomoni Grot
(see: Kakontwe)
Kalondji
(see: Kalongi)
Kalonge
Kalongi
(see also: Kalonji, Kalondji)
Kalonji
(see: Kalongi)
Kalulako
Kalumbu Cave
(see also: Grotte de Kalumbu, Kalumbu Grot)
Kalumbu Grot
(see: Kalumbu Cave)
Kalutenge
Kamande
Kamatanda
Kambemba
Kambove
(see also: Dethiox Caves, Dethiox Grotten, Grottes de Dethiox, Detioux
Cave)
Kambukabakali
Kamiama
(see: Kaniama)
Kamikoni
Kamina Territory
Kamituga
Kamituga and Kampene, road
Kamohohora
Kamp u.m.H(aut).K(atanga).
(see also: Kamp U.M.H.K.)
Kamp U.M.H.K.
(see: Kamp u.m.H(aut).K(atanga).)
Kampala
Kamponde
275
CG-HU
CG-KU
CG
CG-TG
CG-SK
CG-SK
04°40′ N, 29°30′ E
05°06′ S, 18°56′ E
05°50′ S, 28°42′ E
07°52′ S, 29°43′ E
02°15′ S, 28°42′ E
02°16′ S, 28°51′ E
CG-SK
02°16′ S, 28°51′ E
CG-HK
10°59′ S, 26°40′ E
1253 m
CG-HK
10°59′ S, 26°40′ E
1253 m
CG-HK
10°59′ S, 26°40′ E
1253 m
CG-HK
10°59′ S, 26°38′ E
1294 m
CG-HK
10°59′ S, 26°38′ E
1294 m
CG-NK
CG-HK
CG
CG-SK
CG-TG
00°07′ S, 29°37′ E
12°21′ S, 29°35′ E
925 m
ca 02°06′ S, 28°55′ E
05°56′ S, 29°12′ E
763 m
CG-TG
05°56′ S, 29°12′ E
763 m
CG-TG
CG-TG
05°56′ S, 29°12′ E
05°57′ S, 29°01′ E
CG-KO
CG-LM
CG-HU
CG-SK
CG-KN
CG-SK
07°08′ S, 23°32′ E
07°08′ S, 23°32′ E
04°15′ N, 29°45′ E
03°25′ S, 29°09′ E
04°22′ S, 15°17′ E
03°22′ S, 28°01′ E
CG-HK
CG-HK
ca 10°16′ S, 27°37′ E
10°59′ S, 26°40′ E
1294 m
CG-HK
10°59′ S, 26°40′ E
1294 m
CG-KO
06°14′ S, 23°23′ E
799 m
CG-NK
CG-KO
00°20′ N, 29°48′ E
06°14′ S, 23°23′ E
2455 m
799 m
CG-KO
06°14′ S, 23°23′ E
799 m
CG-HK
CG-HK
11°42′ S, 27°34′ E
10°51′ S, 26°39′ E
CG-HK
10°51′ S, 26°39′ E
CG-LB
CG-NK
CG-LB
CG-NK
CG-LB
10°30′ S, 25°37′ E
00°35′ S, 29°18′ E
10°57′ S, 26°46′ E
00°34′ N, 29°41′ E
10°50′ S, 26°39′ E
CG-NK
CG-HL
00°01′ S, 29°36′ E
07°31′ S, 24°11′ E
930 m
887 m
CG-NK
CG-HL
CG-SK
CG-SK
CG-NK
CG-HK
01°16′ S, 29°19′ E
08°44′ S, 25°00′ E
03°03′ S, 28°11′ E
ca 03°38′ S, 27°16′ E
00°40′ S, 29°33′ E
11°41′ S, 27°29′ E
1207 m
CG-HK
11°41′ S, 27°29′ E
CG-SK
CG-LL
02°12′ S, 27°45′ E
06°42′ S, 22°56′ E
451 m
800 m
3051 m
821 m
821 m
128 m
925 m
800 m
1068 m
829 m
European Journal of Taxonomy 382: 1–327 (2017)
Kamuesha
CG-TG
06°12′ S, 21°26′ E
608 m
CG-TG
06°12′ S, 21°26′ E
608 m
CG-SK
CG-TG
03°01′ S, 28°13′ E
06°12′ S, 21°26′ E
608 m
CG-LL
05°54′ S, 22°25′ E
608 m
CG-LL
06°12′ S, 22°23′ E
610 m
RW-ES
02°09′ S, 30°05′ E
1432 m
CG-HK
10°49′ S, 26°07′ E
CG-HK
10°49′ S, 26°07′ E
CG-LB
CG-HL
09°56′ S, 25°58′ E
07°31′ S, 24°11′ E
887 m
CG-HL
CG-KO
07°31′ S, 24°11′ E
06°45′ S, 23°31′ E
805 m
CG-KO
06°45′ S, 23°31′ E
805 m
BY-BM
03°25′ S, 29°22′ E
844 m
CG-NK
01°10′ S, 29°28′ E
1150 m
RW-NO
CG-NK
CG-NK
BY-KI
BY-BM
01°26′ S, 29°35′ E
00°08′ S, 29°37′ E
01°10′ S, 29°34′ E
02°35′ S, 30°05′ E
03°25′ S, 29°22′ E
940 m
1300 m
1508 m
844 m
CG-NK
CG-NK
CG-LB
01°09′ S, 29°24′ E
01°23′ S, 29°13′ E
10°31′ S, 25°12′ E
1150 m
2343 m
1157 m
CG-LB
CG-LB
CG-MN
08°21′ S, 22°35′ E
11°40′ S, 27°29′ E
04°27′ S, 26°40′ E
881 m
625 m
CG-HK
11°02′ S, 26°57′ E
1135 m
CG-HK
11°02′ S, 26°57′ E
1135 m
CG-HK
11°45′ S, 27°20′ E
1274 m
CG-HK
11°45′ S, 27°20′ E
1274 m
CG-HK
11°45′ S, 27°20′ E
1274 m
CG-LB
RW-ES
CG-NK
CG-SK
CG-HK
CG-MA
CG-IT
10°41′ S, 22°52′ E
02°16′ S, 30°16′ E
00°39′ S, 29°21′ E
02°45′ S, 28°38′ E
11°39′ S, 27°17′ E
03°20′ N, 20°18′ E
02°42′ N, 30°33′ E
965 m
453 m
1158 m
CG-NK
BY-BB
BY
CG
CG
00°22′ N, 29°22′ E
03°03′ S, 29°20′ E
1412 m
1226 m
10°22′ S, 23°27′ E
10°27′ S, 23°27′ E
1087 m
(see also: Kamueshi, Kamwesha)
Kamueshi
(see: Kamuesha)
Kamusimbi
Kamwesha
(see: Kamuesha)
Kananga
(see also: Lulua, Luluabourg, Luluaburg, Mission St Joseph)
Kananga, 50 km SW
(see: Kabwe)
Kanazi
(see: Bugesera)
Kando Cave
(see also: Grotte de Kando, Kando Grot)
Kando Grot
(see: Kando Cave)
Kandu (Cave)
Kaniama
(see also: Kamiama)
Kaniama Territory
Kaniki
(see: Kaniki Kapangu)
Kaniki Kapangu
(see also: Bena-Kaniki, Kaniki, Mutembu Kaniki Kapangu)
Kaniosha
(see: Kanyosha)
Kantundwe
(see also: Kontundwe)
Kanyamiketo
Kanyatsi
Kanyawamukunzi River
Kanyinya
Kanyosha
(see also: Kaniosha)
Kanzarue River
Kanzenze
Kanzenze Mission
(see also: Liège Sacre-Coeur Mission, Mission de Liège Sacre-Coeur)
Kapanga
Kapaso
Kapaya
(see: Kasongo)
Kapolowe
(see also: Mission St Gerard, St Gerard Mission, Kapolowe Mission)
Kapolowe Mission
(see: Kapolowe)
Kaponda
(see also: Kaponde, Kapondo)
Kaponde
(see: Kaponda)
Kapondo
(see: Kaponda)
Kapushia
Karama
Karambi
Karambi
Karavia [mine]
Karawa
Karekere
(see: Kerekere)
Karibumba
Karonge
Karuzi Province
Kasai
Kasai Central Province
(see also: Lulua Province)
Kasai o
Kasai Occidental (former) Province
(see also: West Kasai (former) Province)
Kasai Oriental (former) Province
(see also: East Kasai (former) Province)
Kasai Oriental Province
(see also: East Kasai Province, Tshilinge (former) district)
Kasai Province
Kasaji
Kasaji Malonga
276
896 m
CG
CG
CG
CG
CG
CG-LB
CG-LB
VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
Kasaka
Kasaka
Kasambongo
(see also: Cape Cassa Mabongo)
Kasapa
Kasebaya River
(see: Kasebeya River)
Kasebere
Kasebeya River
(see also: Kasebaya River)
Kasende
Kasenga
(see also: Kasenga Mission, Mission Kasenga)
Kasenga Mission
(see: Kasenga)
Kasenga Road, km 106
Kasenga Road, km 36
(see also: Bulaga)
Kasenga Road, km 67
Kasenga Territory
Kasengi
(see: Kasengu)
Kasengu
(see also: Kasengi)
Kasenye
(see also: Kasenyi, Keseny, Kiseny)
Kasenye, 10 km N
Kasenyi
(see: Kasenye)
Kaseu
(see also: Kabinda, 129 km NNW)
Kashiobwe
(see also: Kashiobwe Mission, Mission Kashiobwe)
Kashiobwe Mission
(see: Kashiobwe)
Kashwa
Kasiba
(see also: Irangi, 2 mi [=3.2 km] from)
Kasindi
(see also: Kasindi Vieux, Kasindi-port, Old Kasindi)
Kasindi Vieux
(see: Kasindi)
Kasindi-port
(see: Kasindi)
Kasoma
Kasoma Cave
(see also: Grotte de Kasoma, Kasoma Grot)
Kasoma Grot
(see: Kasoma Cave)
Kasombo
Kasonga-Banze
(see also: Kasongo Baza, Kasongo-Banze)
Kasongo
(see also: Kapaya, Kazongo, Tongoni)
Kasongo Baza
(see: Kasonga-Banze)
Kasongo Mwano
Kasongo Samba
Kasongo Territory
Kasongo-Banze
(see: Kasonga-Banze)
Kasowena Cave
(see also: Grotte de Kasowena, Kasowena Grot)
Kasowena Grot
(see: Kasowena Cave)
Kassi River
Kasuo Cave Kabwe ka
Kaswabilenga
Kaswabilenga
Katabayi
Katana
(see also: Liège Saint Lambert)
Katana Cave
(see: Mahyusha Cave)
Katana Grot
(see: Mahyusha Cave)
277
CG-LM
CG-NK
CG-SK
05°07′ S, 26°09′ E
00°05′ N, 29°36′ E
03°24′ S, 29°08′ E
618 m
900 m
CG-HK
CG-NK
11°34′ S, 27°25′ E
01°10′ S, 29°28′ E
1243 m
1350 m
CG-NK
CG-NK
01°49′ S, 28°45′ E
01°10′ S, 29°28′ E
1350 m
CG-LL
CG-HK
05°57′ S, 22°32′ E
10°22′ S, 28°38′ E
619 m
938 m
CG-HK
10°22′ S, 28°38′ E
938 m
CG-HK
CG-HK
10°44′ S, 28°26′ E
11°37′ S, 27°40′ E
CG-HK
CG-HK
CG-NK
11°00′ S, 28°07′ E
10°22′ S, 28°38′ E
02°08′ N, 31°05′ E
1532 m
CG-NK
02°08′ N, 31°05′ E
1532 m
CG-HU
01°24′ N, 30°26′ E
671 m
CG-IT
CG-HU
ca 01°25′ N, 30°25′ E
01°24′ N, 30°26′ E
671 m
CG-KO
05°11′ S, 23°51′ E
CG-HK
09°39′ S, 28°37′ E
918 m
CG-HK
09°39′ S, 28°37′ E
918 m
CG-NK
CG-SK
01°18′ S, 29°06′ E
ca 01°54′ S, 28°27′ E
2000 m
CG-NK
00°02′ S, 29°43′ E
1049 m
CG-NK
00°02′ S, 29°43′ E
1049 m
CG-NK
00°02′ S, 29°43′ E
1049 m
CG-HL
CG-HL
09°25′ S, 26°37′ E
09°25′ S, 26°37′ E
CG-HL
09°25′ S, 26°37′ E
CG-HK
CG-LL
05°36′ S, 23°24′ E
532 m
CG-MN
04°27′ S, 26°40′ E
625 m
CG-LL
05°36′ S, 23°24′ E
532 m
CG-HK
CG-MN
CG-MN
CG-LL
08°44′ S, 27°51′ E
04°30′ S, 26°40′ E
04°27′ S, 26°40′ E
05°36′ S, 23°24′ E
532 m
CG-HK
10°58′ S, 26°39′ E
CG-HK
10°58′ S, 26°39′ E
CG-HU
CG-LM
CG
CG-HL
CG-KS
CG-SK
04°15′ N, 29°35′ E
06°37′ S, 24°58′ E
08°48′ S, 26°41′ E
06°11′ S, 21°20′ E
02°14′ S, 28°50′ E
1595 m
CG-SK
02°13′ S, 28°50′ E
1480 m
CG-SK
02°13′ S, 28°50′ E
1480 m
700 m
European Journal of Taxonomy 382: 1–327 (2017)
Katanda
(see also: Katenda)
Katanda Road, km 411
Katanga
Katanga (former) Province
(see also: Shaba (former) Province)
Katanga s
Katanga, near
Katapena
Kataye Sukunga
Kateke [River]
(see also: Kateke)
Katenda
(see: Katanda)
Katende chute
(see: Katende Falls)
Katende Falls
(see also: Katende chute, Katende Waterval )
Katende Waterval
(see: Katende Falls)
Katentania
Katjumbura
Katobwe
(see also: Bukama, 40 mi [=64 km] below)
Katshungu
Katupila
Katwé
Kavuaya
Kawa (Forest)
Kawezi-Uvira Road
Kayanza
Kayanza Province
Kayanza, 15 km W
(see: Rwegura)
Kayombe
(see: Dilolo)
Kayonza
Kayove
Kazongo
(see: Kasongo)
Kele Cave 1
(see also: Grotte de Kele 1, Kele Grot 1)
Kele Grot 1
(see: Kele Cave 1)
Kenya
Kere Kere
(see: Kerekere)
Kerekere
(see also: Karekere, Kere Kere)
Keri River
(see also: Kiri River)
Keroma
Keseki
(see also: Kesiki)
Keseny
(see: Kasenye)
Kesiki
(see: Keseki)
Keyberg
Kiabo, 14 km upstream
(see: Sombe)
Kiamanwa
Kiambi
Kiamokoto
(see also: Grotte de Kiwakishi, Kiwakishi Cave, Kiwakishi Grot)
Kiantapo Cave
(see: Kyantapo Cave)
Kiasala Cave
(see also: Grotte de Kiasala, Grotte de Kyasala, Kiasala Grot, Kyasala Cave,
Kyasala Grot, Kyasala grande Grotte)
Kiasala Grot
(see: Kiasala Cave)
Kiavikere
(see also: Kiavikire)
Kiavikire
(see: Kiavikere)
278
CG-NK
00°51′ S, 29°22′ E
CG-NK
CG
CG
01°00′ S, 29°00′ E
975 m
CG
CG
CG-HK
CG-KS
CG-HL
ca 08°35′ S, 26°25′ E
CG-NK
00°51′ S, 29°22′ E
975 m
CG-KS
05°46′ S, 21°40′ E
647 m
CG-KS
05°46′ S, 21°40′ E
647 m
CG-KS
05°46′ S, 21°40′ E
647 m
CG-LB
RW-ES
CG-HL
10°19′ S, 25°54′ E
01°33′ S, 30°44′ E
08°54′ S, 26°04′ E
1597 m
CG-SK
CG-HL
CG-HK
CG-BC
CG-IT
CG-SK
BY-KY
BY
BY-KY
02°27′ S, 27°23′ E
08°50′ S, 26°44′ E
10°28′ S, 27°51′ E
05°20′ S, 14°52′ E
01°34′ N, 30°32′ E
ca 03°19′ S, 29°10′ E
02°55′ S, 29°37′ E
1013 m
1957 m
02°55′ S, 29°31′ E
2311 m
CG-LB
10°28′ S, 22°28′ E
1069 m
RW-ES
RW-OU
CG-MN
01°57′ S, 30°31′ E
01°53′ S, 29°21′ E
04°27′ S, 26°40′ E
2400 m
1841 m
625 m
CG-BC
05°31′ S, 14°36′ E
CG-BC
05°31′ S, 14°36′ E
CG-HK
CG-IT
11°44′ S, 27°30′ E
02°42′ N, 30°33′ E
1119 m
1158 m
CG-IT
02°42′ N, 30°33′ E
1158 m
CG-MA
01°29′ S, 18°58′ E
CG-HU
CG-MA
04°35′ N, 29°32′ E
02°07′ S, 16°32′ E
308 m
CG-HU
01°24′ N, 30°26′ E
671 m
CG-MA
02°07′ S, 16°32′ E
308 m
CG-HK
CG-HL
11°43′ S, 27°25′ E
08°50′ S, 26°02′ E
CG-HK
CG-TG
CG-HL
09°30′ S, 27°55′ E
07°20′ S, 28°01′ E
09°10′ S, 27°04′ E
607 m
900 m
CG-LB
09°56′ S, 25°58′ E
1525 m
CG-LB
09°56′ S, 25°58′ E
1300 m
CG-LB
09°56′ S, 25°58′ E
1300 m
CG-NK
00°24′ N, 29°46′ E
1553 m
CG-NK
00°24′ N, 29°46′ E
1553 m
09°04′ S, 26°43′ E
610 m
753 m
VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
Kiavinionge
(see: Kyavinionge)
Kibanda
Kibanda
Kibanga
Kibimba Mission
Kibira National Park
(see also: Teza, 2.3 km N, 0.7 km W)
Kibumba
Kibumbu crater
Kibungo (former) Prefecture
(see: Eastern Prefecture)
Kibuye
Kibwe Lya Mikako Cave
(see also: Ribue Lya Mikako Cave, Grotte de Ribue Lya Mikako, Grotte de
Kibwe Lya Mikako)
Kidada
Kidaho
(see also: Colline Kidaho)
Kifumwanshi
Kifuruzi
(see also: Gifuruzi)
Kigale
(see: Kigali)
Kigali
(see also: Kigale)
Kigali Prefecture
Kigamba
Kigwena Hill
Kihanga
(see: Gihanga)
Kihumba
(see: Kiumba)
Kikanda
Kikango
(see also: Moba - Mplala Road, km 35)
Kikole-Pecheur
Kikondja
(see also: Kikondji)
Kikondji
(see: Kikondja)
Kikura
Kikwit
(see also: Kikwit ITPK)
Kikwit ITPK
(see: Kikwit)
Kilembe, 52 mi [=84 km] S
Kilemue
Kilia Kaparata
Kilibozi
Kiligi
Kilinga
Kilitza [River]
(see: Kiliza [River])
Kiliza [River]
(see also: Kaliza [River], Kilitza [River])
Kilo mines
(see: Kilomines)
Kilobelobe
(see: Kilobilobe)
Kilobilobe
(see also: Kilobelobe)
Kilobono
(see also: Kilobore, Kiloboze)
Kilobore
(see: Kilobono)
Kiloboze
(see: Kilobono)
Kilomamensa
Kilomines
(see also: Bambu, Mines de Kilo, Kilo mines)
Kilubulushi
(see: Kitubulushi)
Kilubwezi
Kilwa
Kilwezi
Kimbemba Cataracts
279
CG-TO
00°07′ S, 26°36′ E
935 m
CG-HK
CG-SK
CG-HL
BY-GI
BY-MV
08°57′ S, 27°38′ E
03°20′ S, 27°12′ E
08°50′ S, 26°44′ E
03°18′ S, 29°49′ E
03°13′ S, 29°34′ E
CG-NK
RW-NO
RW
01°28′ S, 29°20′ E
01°25′ S, 29°33′ E
ca 02°00′ S, 30°35′ E
1223 m
688 m
1300 m
1730 m
2200–2400
m
RW-OU
CG-NK
02°03′ S, 29°21′ E
00°15′ S, 29°01′ E
1457 m
1500 m
CG-BC
RW-NO
05°22′ S, 14°32′ E
01°22′ S, 29°45′ E
1944 m
CG-HK
BY-MA
11°27′ S, 27°51′ E
04°19′ S, 29°35′ E
RW-KV
01°57′ S, 30°04′ E
1567 m
RW-KV
01°57′ S, 30°04′ E
1567 m
RW
BY-CA
BY-RU
BY-BB
03°04′ S, 30°31′ E
04°09′ S, 29°32′ E
03°12′ S, 29°18′ E
1532 m
918 m
814 m
CG-SK
01°56′ S, 29°05′ E
1602 m
CG-HK
CG-TG
11°33′ S, 27°37′ E
06°37′ S, 29°30′ E
CG
CG-HL
08°11′ S, 26°26′ E
607 m
CG-HL
08°11′ S, 26°26′ E
607 m
CG-NK
CG-KU
00°33′ N, 29°57′ E
05°04′ S, 18°53′ E
1900 m
438 m
CG-KU
05°04′ S, 18°53′ E
438 m
CG-KU
CG-SK
CG-NK
CG-SK
CG-SK
CG-NK
CG-SK
05°47′ S, 19°54′ E
02°53′ S, 28°07′ E
00°23′ N, 29°36′ E
03°03′ S, 28°09′ E
02°53′ S, 28°18′ E
01°10′ S, 28°29′ E
03°22′ S, 28°01′ E
CG-SK
03°22′ S, 28°01′ E
CG-IT
01°48′ N, 30°14′ E
CG-HK
11°42′ S, 27°34′ E
CG-HK
11°42′ S, 27°34′ E
CG-SK
03°03′ S, 28°11′ E
CG-SK
03°03′ S, 28°11′ E
CG-SK
03°03′ S, 28°11′ E
CG
CG-IT
01°48′ N, 30°14′ E
1466 m
CG-HK
12°44′ S, 28°56′ E
1257 m
CG
CG-HK
CG-HL
CG-BC
09°18′ S, 28°20′ E
09°06′ S, 26°46′ E
ca 04°51′ S, 14°06′ E
918 m
854 m
980 m
1275 m
1466 m
European Journal of Taxonomy 382: 1–327 (2017)
Kimbinga Market
Kimilolo
Kimpoko
(see: Kimpoko-Nsele)
Kimpoko-Nsele
(see also: Kimpoko, Nsele)
Kimuenza Cave
(see also: Grotte de Kimuenza, Kimuenza Grot, Kimwenza Cave)
Kimuenza Grot
(see: Kimuenza Cave)
Kimuna
Kimvuadu
Kimwenza Cave
(see: Kimuenza Cave)
Kinawa Passage
Kinazi
Kinda
Kindu
(see also: Kindu-Port-Empain, Port-Empain, Port de Kindu)
Kindu, 30 km SW
(see: Scierie Forest)
Kindu-Port-Empain
(see: Kindu)
Kingabwa
Kingulube
Kinigi
Kinke
Kinkole
Kinshasa
(see also: Leopoldstad, Léopoldville, Lovanium University, Leopoldville)
Kinshasa City Province
Kinshasa, 10 km W
(see also: TASOK (The American School of Kinshasa))
Kinshasa, near Leopoldville
(see also: Léo-Est)
Kintobongo
(see also: Kitombongo, Kitobongo, Kintobongo-Bunge)
Kintobongo-Bunge
(see: Kintobongo)
Kinyamahura
Kinzambi Mission
(see also: Mission de Kinzambi)
Kionja
Kipangaribwe River
Kipopo
Kipushi
Kipushi Territory
Kipushi, 15 km from
(see: Wasseige Farm)
Kipushi, Road to
Kiremba Mission
Kiri River
(see: Keri River)
Kirumura, 1.1 - 2.1 km N, 0.8 - 1.1 km E
(see: Giserama, Colline)
Kirundo Province
Kisala
(see: Kisala-Kiama)
Kisala Nkiama
(see: Kisala-Kiama)
Kisala-Kiama
(see also: Kisala Nkiama, Kisala)
Kisanga
Kisanga
Kisangani
(see also: Singitini, Stanleystad, Stanleyville, Zookis, Université de
Kisangani, Faculté des Sciences)
Kisangani - Mambasa Road, km 122
(see: Maganga)
Kisangani, 45 km E
(see: Tshakala)
Kisangani, Zone
Kisantu
Kiseny
(see: Kasenye)
280
CG-KU
CG-HK
CG-KN
05°08′ S, 19°03′ E
11°42′ S, 27°25′ E
04°14′ S, 15°33′ E
294 m
CG-KN
04°14′ S, 15°33′ E
294 m
CG-BC
04°28′ S, 15°17′ E
CG-BC
04°28′ S, 15°17′ E
RW-SU
CG-BC
CG-BC
02°33′ S, 29°44′ E
05°15′ S, 13°18′ E
04°28′ S, 15°17′ E
CG-NK
BY-KI
CG-KS
CG-MN
00°40′ N, 29°50′ E
02°23′ S, 29°58′ E
04°47′ S, 21°48′ E
02°57′ S, 25°57′ E
CG-MN
03°10′ S, 25°46′ E
CG-MN
02°57′ S, 25°57′ E
504 m
CG-KN
CG-SK
RW-NO
CG-HK
CG-ET
CG-KN
04°18′ S, 15°18′ E
02°39′ S, 28°01′ E
01°27′ S, 29°34′ E
11°27′ S, 27°29′ E
03°57′ S, 16°07′ E
04°20′ S, 15°19′ E
1238 m
2179 m
1287 m
474 m
240 m
CG
CG-KN
ca 04°20′ S, 15°15′ E
CG-KN
04°18′ S, 15°18′ E
177 m
CG-HL
08°54′ S, 26°23′ E
608 m
CG-HL
08°54′ S, 26°23′ E
608 m
CG-NK
CG-KU
01°18′ S, 29°33′ E
04°59′ S, 18°46′ E
1905 m
312 m
RW-ES
CG-HL
01°50′ S, 30°43′ E
08°56′ S, 27°12′ E
CG-HK
CG-HK
CG-HK
CG-HK
11°34′ S, 27°21′ E
11°46′ S, 27°14′ E
11°46′ S, 27°14′ E
11°50′ S, 27°20′ E
1677 m
1513–1760
m
CG-HK
BY-BI
CG-MA
ca 11°45′ S, 27°15′ E
03°56′ S, 29°41′ E
01°29′ S, 18°58′ E
BY-CI
02°50′ S, 29°25′ E
2045 m
BY
CG-BC
04°48′ S, 13°10′ E
433 m
CG-BC
04°48′ S, 13°10′ E
433 m
CG-BC
04°48′ S, 13°10′ E
433 m
CG-SK
CG-SK
CG-TO
02°25′ S, 28°18′ E
00°16′ N, 29°43′ E
00°31′ N, 25°12′ E
960 m
447 m
CG-TO
00°51′ N, 26°22′ E
CG-TO
00°27′ N, 25°32′ E
CG-TO
CG-BC
CG-HU
585 m
1788 m
470 m
504 m
1329 m
453 m
ca 00°30′ N, 25°12′ E
05°07′ S, 15°05′ E 421–571 m
01°24′ N, 30°26′ E
671 m
VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
Kisenyi
(see: Gisenyi)
Kisenyi, Road to Sake
(see: Gisenyi)
Kishero
Kisiguru (Mission)
(see: Kiziguro (Mission))
Kisiki
(see also: Kisiki Kalume, Beni, 36 km N)
Kisiki Kalume
(see: Kisiki)
Kisingani
Kisisile
Kisiushi
Kissenji
(see: Gisenyi)
Kiswishi [River]
Kitabi
Kitangua
Kitega
(see: Gitega)
Kitimi
Kitobongo
(see: Kintobongo)
Kitombongo
(see: Kintobongo)
Kitonga
Kitongo
Kitubulushi
(see also: Kilubulushi)
Kitundu Kinda
Kitutu
Kitwabaluzi
Kiulu Mgemgo
Kiumba
(see also: Kihumba)
Kivu
(see: Kivu region)
Kivu Meer
(see: Lake Kivu)
Kivu mountains
Kivu region
(see also: Kivu)
Kivu Volcano
(see: Nyirogongo Volcano)
Kivu, Lac
(see: Lake Kivu)
Kivu, Lake
(see: Lake Kivu)
Kivu, near
Kivulo
Kiwakishi Cave
(see: Kiamokoto)
Kiwakishi Grot
(see: Kiamokoto)
Kiyite
Kiziguro (Mission)
(see also: Kisiguru (Mission), Kiziguru (Mission))
Kiziguru (Mission)
(see: Kiziguro (Mission))
Ko[g]enga
Kodja Hill
Kodoko River
(see: Kokodo River)
Kokodo River
(see also: Kodoko River)
Kokodu
Kokonde
Koloka
Kolwezi
Kolwezi Province
Komanda
(see also: Komande)
Komande
(see: Komanda)
Kona
RW-OU
01°41′ S, 29°15′ E
1450 m
RW-OU
01°41′ S, 29°15′ E
1450 m
CG-NK
RW-ES
00°11′ N, 29°39′ E
01°46′ S, 30°24′ E
925 m
CG-NK
00°45′ N, 29°34′ E
1177 m
CG-NK
00°45′ N, 29°34′ E
1177 m
CG
CG-NK
CG-HK
RW-OU
00°32′ N, 25°11′ E
01°25′ S, 29°30′ E
11°30′ S, 27°30′ E
01°41′ S, 29°15′ E
1250 m
1450 m
CG-HK
RW-SU
CG-KS
BY-GI
12°51′ S, 29°12′ E
02°31′ S, 29°25′ E
06°17′ S, 20°20′ E
03°26′ S, 29°56′ E
649 m
1845 m
CG-MA
CG-HL
03°58′ S, 15°53′ E
08°54′ S, 26°23′ E
608 m
CG-HL
08°54′ S, 26°23′ E
608 m
CG-NK
CG-SK
CG-HK
00°30′ S, 29°18′ E
03°46′ S, 28°11′ E
12°44′ S, 28°56′ E
920 m
1400 m
1257 m
CG-HL
CG-SK
CG-SK
CG-KU
CG-SK
09°30′ S, 24°55′ E
03°17′ S, 28°05′ E
02°53′ S, 28°35′ E
04°24′ S, 17°42′ E
01°56′ S, 29°05′ E
1040 m
1543 m
1602 m
CG
281
CG-SK
ca 02°00′ S, 28°55′ E
CG-SK
CG
ca 02°00′ S, 28°45′ E
CG-NK
01°31′ S, 29°15′ E
CG-SK
ca 02°00′ S, 28°55′ E
CG-SK
ca 02°00′ S, 28°55′ E
CG-SK
CG-HK
CG-HL
ca 02°00′ S, 28°45′ E
09°10′ S, 27°04′ E
900 m
CG-HL
09°10′ S, 27°04′ E
900 m
CG-NK
RW-ES
01°46′ S, 30°24′ E
RW-ES
01°46′ S, 30°24′ E
CG
CG-HU
CG-HU
03°09′ N, 29°29′ E
04°38′ N, 29°30′ E
CG-HU
04°38′ N, 29°30′ E
CG-HU
CG
CG-BU
CG-LB
CG
CG-IT
04°35′ N, 29°32′ E
03°23′ N, 24°30′ E
10°43′ S, 25°28′ E
1448 m
01°23′ N, 29°46′ E
928 m
CG-IT
01°23′ N, 29°46′ E
928 m
CG-MO
02°02′ N, 22°47′ E
European Journal of Taxonomy 382: 1–327 (2017)
Kondo Cave
(see: Kakanda Cave)
Kondo Grot
(see: Kakanda Cave)
Kondue [River]
Kongo Central Province
(see also: Bas Congo Province, Bas Congo (former) Province)
Kongolo
Kongolo, Island
(see also: Ile de Kongolo)
Kontundwe
(see: Kantundwe)
Konyola
Korovi, mount
(see also: Mount Korovi)
Koteli
(see: Kotili)
Kotili
(see also: Koteli)
Kouilou River
Kuango
Kuili
(see also: Kuili Ngongo)
Kuili Ngongo
(see: Kuili)
Kuleponge
(see: Kulepongi)
Kulepongi
(see also: Kuleponge, Kulepongo)
Kulepongo
(see: Kulepongi)
Kumbi
Kumi
Kundelungu National Park
(see also: Park National de Kundelungo)
Kunungu
Kwama Kwama Cave
(see also: Grotte de Kwama Kwama, Kwama Kwama Grot)
Kwama Kwama Grot
(see: Kwama Kwama Cave)
Kwamouth
Kwango
Kwango Province
Kwilu Province
Kwogofe, Colline
(see also: Colline Kwogofe)
Kya irtumbi
(see: Mutwanga)
Kyamakonde Cave
Kyantapo Cave
(see also: Grotte de Kyantapo, Kiantapo Cave, Kyantapo Grot, Lubudi, 30
km SW)
Kyantapo Grot
(see: Kyantapo Cave)
Kyasala Cave
(see: Kiasala Cave)
Kyasala grande Grotte
(see: Kiasala Cave)
Kyasala Grot
(see: Kiasala Cave)
Kyavinionge
(see also: Kiavinionge)
Kyniamahura
Labo Irsac
(see: Luiro)
Lac Albert
(see: Albert, Lake)
Lac Cohoha
(see: Cohoha, Lake)
Lac Edouard
(see: Edward, Lake)
Lac Fwa
(see: Fwa, Lake)
Lac Idi Amine
(see: Edward, Lake)
282
CG-LB
10°47′ S, 26°36′ E
1438 m
CG-LB
10°47′ S, 26°36′ E
1438 m
CG-SN
CG
04°57′ S, 23°21′ E
ca 05°00′ S, 15°00′ E
CG-TG
CG-TO
05°25′ S, 27°00′ E
00°03′ N, 25°12′ E
CG-NK
01°10′ S, 29°28′ E
1150 m
CG-TO
CG-IT
00°03′ N, 25°12′ E
01°39′ N, 30°36′ E
2010 m
CG-BU
02°52′ N, 24°33′ E
432 m
CG-BU
02°52′ N, 24°33′ E
432 m
CG
CG-BC
CG-KU
04°45′ S, 13°36′ E
04°24′ S, 17°42′ E
CG-KU
04°24′ S, 17°42′ E
CG-MN
03°22′ N, 25°41′ E
604 m
CG-MN
03°22′ N, 25°41′ E
604 m
CG-MN
03°22′ N, 25°41′ E
604 m
CG
CG-MN
CG-HK
04°32′ S, 26°33′ E
ca 10°30′ S, 27°30′ E
CG-MA
CG-NK
02°06′ S, 16°26′ E
01°19′ N, 29°52′ E
1200 m
CG-NK
01°19′ N, 29°52′ E
1200 m
CG-MA
CG-KG
CG
CG
BY-CI
03°10′ S, 16°12′ E
06°28′ S, 17°27′ E
ca 06°30′ S, 17°46′ E
146 m
1020 m
02°50′ S, 29°25′ E
CG-NK
00°20′ N, 29°45′ E
2180–2240
m
1338 m
CG-LB
CG-LB
ca 09°56′ S, 25°58′ E
09°56′ S, 25°58′ E
1525 m
CG-LB
09°56′ S, 25°58′ E
1525 m
CG-LB
09°56′ S, 25°58′ E
1300 m
CG-LB
09°56′ S, 25°58′ E
1300 m
CG-LB
09°56′ S, 25°58′ E
1300 m
CG-TO
00°07′ S, 26°36′ E
935 m
CG
CG-SK
02°14′ S, 28°48′ E
1646 m
CG-IT
ca 01°40′ N, 30°40′ E
BY-KI
02°30′ S, 30°07′ E
CG-NK
ca 00°08′ S, 29°35′ E
CG-LL
05°43′ S, 23°25′ E
CG-NK
ca 00°08′ S, 29°35′ E
VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
Lac Ihema
RW-ES
01°52′ S, 30°47′ E
CG-SK
ca 02°00′ S, 28°55′ E
CG-MA
02°05′ S, 18°20′ E
(see: Ihema, Lake)
Lac Kivu
(see: Lake Kivu)
Lac Léopold II
(see: Mai-Ndombe, Lake)
Lac Marion
(see: Marion, Lake)
Lac Mihindi
(see: Mihindi, Lake)
Lac Moero
(see: Moero, Lake)
Lac Mugunga
(see: Mugunga, Lake)
Lac Ondo
(see: Ondo, Lake)
Lac Tanganyika, bord Nord-Ouest
(see: Lake Tanganyika, NW bank)
Lac Tumba
(see: Tumba, Lake)
Lai Lay
(see: Lailai)
Lailai
(see also: Lai Lay)
Lake Albert
(see: Albert, Lake)
Lake Cohoha
(see: Cohoha, Lake)
Lake Edward
(see: Edward, Lake)
Lake Edward, S end
Lake Fwa
(see: Fwa, Lake)
Lake Idi Amin Dada
(see: Edward, Lake)
Lake Ihema
(see: Ihema, Lake)
Lake Kivu
(see also: Kivu, Lake, Kivu, Lac, Kivu Meer, Lac Kivu)
Lake Kivu, near
Lake Leopold II
(see: Mai-Ndombe, Lake)
Lake Mai-Ndombe
(see: Mai-Ndombe, Lake)
Lake Marion
(see: Marion, Lake)
Lake Mihindi
(see: Mihindi, Lake)
Lake Mobutu Sese Seko
(see: Albert, Lake)
Lake Moero
(see: Moero, Lake)
Lake Mugunga
(see: Mugunga, Lake)
Lake Ondo
(see: Ondo, Lake)
Lake Rutanzige
(see: Edward, Lake)
Lake Tanganyika, NW bank
(see also: Lac Tanganyika, bord Nord-Ouest, Tanganyika Meer, NW oever,
Tanganyika, Lake, NW bank)
Lake Tumba
(see: Tumba, Lake)
Lake Tumba, Island in
Lamya Valley
(see also: Lanuga Valley)
Lanuga Valley
(see: Lamya Valley)
Lemba Plain
(see also: Lemba Vlakte)
Lemba Vlakte
(see: Lemba Plain)
Lemera
Lemera Great Sink
(see: Itombwe)
Lemera, SW of
283
CG
RW-ES
01°32′ S, 30°42′ E
CG-HK
09°00′ S, 28°45′ E
CG-NK
01°37′ S, 29°08′ E
CG-NK
01°08′ S, 29°19′ E
CG-SK
ca 03°20′ S, 29°12′ E
CG-ET
ca 00°48′ S, 18°03′ E
CG-SK
01°57′ S, 28°34′ E
1276 m
CG-SK
01°57′ S, 28°34′ E
1276 m
CG-IT
ca 01°40′ N, 30°40′ E
BY-KI
02°30′ S, 30°07′ E
CG-NK
ca 00°08′ S, 29°35′ E
CG-NK
CG-LL
ca 00°41′ S, 29°23′ E
05°43′ S, 23°25′ E
CG-NK
ca 00°08′ S, 29°35′ E
RW-ES
01°52′ S, 30°47′ E
CG-SK
ca 02°00′ S, 28°55′ E
CG-SK
CG-MA
ca 02°00′ S, 29°10′ E
02°05′ S, 18°20′ E
CG-MA
02°05′ S, 18°20′ E
1500 m
CG
RW-ES
01°32′ S, 30°42′ E
CG-IT
ca 01°40′ N, 30°40′ E
CG-HK
09°00′ S, 28°45′ E
CG-NK
01°37′ S, 29°08′ E
CG-NK
01°08′ S, 29°19′ E
CG-NK
ca 00°08′ S, 29°35′ E
CG-SK
ca 03°20′ S, 29°12′ E
CG-ET
ca 00°48′ S, 18°03′ E
CG-ET
CG-IT
ca 00°48′ S, 18°03′ E
00°50′ N, 29°59′ E
CG-IT
00°50′ N, 29°59′ E
CG-KN
04°18′ S, 15°18′ E
CG-KN
04°18′ S, 15°18′ E
CG-SK
CG-SK
02°08′ S, 28°50′ E
04°04′ S, 28°40′ E
2281 m
CG-SK
ca 02°10′ S, 28°50′ E
2682 m
1500 m
European Journal of Taxonomy 382: 1–327 (2017)
Léo-Est
(see: Kinshasa, near Leopoldville)
Leopold 2nd, Lake
(see: Mai-Ndombe, Lake)
Leopold II Meer
(see: Mai-Ndombe, Lake)
Leopold II, Lake
(see: Mai-Ndombe, Lake)
Leopoldstad
(see: Kinshasa)
Leopoldville
(see: Kinshasa)
Léopoldville
(see: Kinshasa)
Lesse
Leverstad
(see: Lusanga)
Leverville
(see: Lusanga)
Libenge
(see also: Libengué)
Libengué
(see: Libenge)
Liège Sacre-Coeur Mission
(see: Kanzenze Mission)
Liège Saint Lambert
(see: Katana)
Lieki
Likandi River
(see: Litaki River)
Likasi
(see also: Farm ‘des Grands Bois’, Jadotstad, Jadotsville, Jadotville, Mission
Sainte Thérèse, Panda Camp, St Therese Mission, Jadoville, St. Theresa
Mission)
Likasi, 130 km N
(see: Tenke)
Likasi, 15 km S
(see: Mwanga Cave)
Likasi, ca 8 km S
(see: Kakontwe)
Likati River
Liki
Lima
Liondo
(see also: Loondo)
Liondo
Lisala
(see also: Lizala)
Lisala Territory
Litaki River
(see also: Likandi River)
Lizala
(see: Lisala)
Loanza Mission
(see: Luanza Mission)
Lobango
(see also: Lubango)
Lobi
Lodja
(see also: Loja)
Lodja Territory
Lofoi [River]
Loja
(see: Lodja)
Lomami Province
(see also: Kabinda (former) district)
Lomera
Lonzo
(see: Loso)
Loondo
(see: Liondo)
Loondo Mugunga
Losho
(see: Loso)
Loso
(see also: Losho, Lonzo)
284
CG-KN
04°18′ S, 15°18′ E
CG-MA
02°05′ S, 18°20′ E
CG-MA
02°05′ S, 18°20′ E
CG-MA
02°05′ S, 18°20′ E
CG-KN
04°20′ S, 15°19′ E
240 m
CG-KN
04°18′ S, 15°18′ E
240 m
CG-KN
04°20′ S, 15°19′ E
240 m
CG-NK
CG-KU
00°45′ N, 29°48′ E
04°50′ S, 18°44′ E
800 m
307 m
CG-KU
04°50′ S, 18°44′ E
307 m
CG-MA
03°39′ N, 18°38′ E
326 m
CG-MA
03°39′ N, 18°38′ E
326 m
CG-LB
10°31′ S, 25°12′ E
1157 m
CG-SK
02°14′ S, 28°50′ E
1595 m
CG-TO
CG-BU
00°41′ N, 24°11′ E
03°11′ N, 26°11′ E
508 m
CG-HK
10°59′ S, 26°44′ E
1318 m
CG-HK
10°35′ S, 26°07′ E
1512 m
CG-HK
10°59′ S, 26°40′ E
1253 m
CG-BU
CG-NK
CG-NK
CG-TP
02°54′ N, 24°03′ E
00°07′ S, 29°34′ E
00°12′ N, 29°18′ E
00°55′ S, 22°41′ E
416 m
CG-TP
CG-MO
00°48′ S, 22°39′ E
02°09′ N, 21°31′ E
508 m
409 m
CG-MO
CG-BU
02°09′ N, 21°31′ E
03°11′ N, 26°11′ E
CG-MO
02°09′ N, 21°31′ E
409 m
CG-HK
08°42′ S, 28°42′ E
1132 m
CG-NK
00°19′ S, 29°13′ E
2197 m
CG-BU
CG-SN
04°10′ N, 23°06′ E
03°29′ S, 23°26′ E
450 m
CG-SN
CG-LB
CG-SN
03°29′ S, 23°26′ E
10°13′ S, 27°25′ E
03°29′ S, 23°26′ E
450 m
CG-NK
CG-NK
00°07′ N, 29°35′ E
00°08′ S, 29°36′ E
915 m
950 m
CG-TP
00°55′ S, 22°41′ E
416 m
CG-TP
CG-NK
00°42′ S, 22°41′ E
00°08′ S, 29°36′ E
950 m
CG-NK
00°08′ S, 29°36′ E
950 m
177 m
CG-HK
CG
VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
Lotakamela
Lovanium University
(see: Kinshasa)
Lovoi
Lower Uele Province
(see: Bas Uele Province)
Lualaba Province
Lualaba River
(see: Mulongo)
Luanu
Luanza missie
(see: Luanza Mission)
Luanza Mission
(see also: Loanza Mission, Luanza missie, Mission de Luanza)
Lubango
(see: Lobango)
Lubenge
Lubengera (Mission)
(see: Rubengera (Mission))
Luberezi
(see: Luberizi)
Luberizi
(see also: Luberezi)
Lubondaie-Tshimbulu
(see also: Lubondai-Tshimbulu)
Lubondai-Tshimbulu
(see: Lubondaie-Tshimbulu)
Lubudi
(see also: Lubudi, large Cave, Lubudi, small Cave, Lusolo Cave)
Lubudi, 30 km SW
(see: Kyantapo Cave)
Lubudi, large Cave
(see: Lubudi)
Lubudi, small Cave
(see: Lubudi)
Lubudi-Lusolo
Lubuka Cave
(see also: Grotte de Lubuka, Lubuka Grot)
Lubuka Grot
(see: Lubuka Cave)
Lubumbashi
(see also: Elisabethstad, Elisabethville, E’ville, Lumumbashi, Elizabethville)
Lubumbashi, 30 km S
(see: Baya quarry)
Lubumbashi, 68 km E
Lubumbashi, 69 km E
(see: Musonge)
Lubumbashi, 70 km E
(see: Mumena)
Lubutu
Lubutu Territory
Lubwe
Luebo
Luebo Territory
Luena
Lufa River
Luira River, upper
(see also: Lupufa River, near)
Lufu
Lugarama
Luhohov (River)
Luiko
(see: Lwiko)
Luiro
(see also: Irsac Labo, Labo Irsac, Lwiro, Mwiru, Lwiro Katana)
Luiza Territory
Lukafu
Lukatu Cave
(see: Ntoto Cave)
Lukatu Grot
(see: Ntoto Cave)
Luki Biosphere Reserve
Luki Biosphere Reserve, Forest NE of
Lukobwe
Lukoi River
Lukolela
Lukolela, 4 m. from?
285
CG-TP
CG-KN
00°14′ S, 21°57′ E
04°20′ S, 15°19′ E
508 m
240 m
CG-HL
CG
08°48′ S, 24°50′ E
1126 m
CG
CG-TG
07°50′ S, 27°00′ E
578 m
CG-KU
CG-HK
05°13′ S, 19°08′ E
08°42′ S, 28°42′ E
1132 m
CG-HK
08°42′ S, 28°42′ E
1132 m
CG-NK
00°19′ S, 29°13′ E
2197 m
CG-TG
RW-SU
07°30′ S, 29°59′ E
02°03′ S, 29°26′ E
1722 m
CG-SK
02°59′ S, 29°06′ E
957 m
CG-SK
02°59′ S, 29°06′ E
957 m
CG-LL
06°34′ S, 22°39′ E
794 m
CG-LL
06°34′ S, 22°39′ E
794 m
CG-LB
09°57′ S, 25°58′ E
1307 m
CG-LB
09°56′ S, 25°58′ E
1525 m
CG-LB
09°57′ S, 25°58′ E
1307 m
CG-LB
09°57′ S, 25°58′ E
1307 m
CG-LB
CG-SK
09°56′ S, 25°58′ E
03°12′ S, 28°51′ E
CG-SK
03°12′ S, 28°51′ E
CG-HK
11°40′ S, 27°28′ E
1208 m
CG-HK
11°52′ S, 27°27′ E
1279 m
CG-HK
CG-HK
11°07′ S, 28°08′ E
11°28′ S, 27°52′ E
1183 m
CG-HK
11°04′ S, 28°08′ E
CG-MN
CG-MN
CG-HK
CG-KS
CG-KS
CG-HL
CG-HU
CG-HK
00°44′ S, 26°35′ E
00°44′ S, 26°35′ E
10°43′ S, 27°23′ E
05°21′ S, 21°25′ E
05°36′ S, 21°25′ E
09°26′ S, 25°47′ E
04°02′ N, 29°46′ E
ca 10°54′ S, 26°60′ E
CG-BC
RW-ES
CG-NK
CG-SK
05°40′ S, 13°53′ E
01°26′ S, 30°32′ E
01°30′ S, 28°06′ E
03°05′ S, 28°47′ E
CG-SK
02°14′ S, 28°48′ E
1646 m
CG-LL
CG-HK
CG-TG
07°12′ S, 22°25′ E
10°31′ S, 27°33′ E
06°59′ S, 29°43′ E
930 m
827 m
CG-TG
06°59′ S, 29°43′ E
827 m
CG-BC
CG-BC
CG-HL
CG
CG-MA
CG-MA
05°38′ S, 13°13′ E
ca 05°38′ S, 13°13′ E
09°06′ S, 27°13′ E
01°03′ S, 17°12′ E
ca 01°03′ S, 17°12′ E
669 m
478 m
285 m
European Journal of Taxonomy 382: 1–327 (2017)
Lukonga
Lukonzolwa
Lukula
Lukula, 15 km W
(see: Bondo)
Lukulu
Lula
Lulenga
(see: Rugari)
Lulimba
Lulimbi
Lulola-Kasanga
(see also: Mulolo-Kisanga)
Lulua
(see: Kananga)
Lulua Province
(see: Kasai Central Province)
Luluabourg
(see: Kananga)
Luluabourg ravine
Luluaburg
(see: Kananga)
Luluaburg, 50 km SW
(see: Kabwe)
Lume River, near
Lumumbashi
(see: Lubumbashi)
Lundjulu
Luofu
Lupufa River, near
Luira River, upper
Luputa
Lurhogole Swamp area
Lusambo
(see also: Luzambo)
Lusanga
(see also: Leverstad, Leverville)
Lusanga Mwanza
Lusano and KKK Zone
Lushala
Lusilubi Valley
Lusinga
CG-LL
CG-HK
CG-BC
CG-BC
05°48′ S, 22°37′ E
08°47′ S, 28°39′ E
05°23′ S, 12°57′ E
05°24′ S, 12°54′ E
632 m
918 m
196 m
CG-TG
CG-LL
CG-NK
07°08′ S, 27°07′ E
ca 07°25′ S, 22°33′ E
01°25′ S, 29°22′ E
573 m
CG-SK
CG-NK
CG-MN
04°42′ S, 28°38′ E
00°28′ S, 29°39′ E
03°06′ S, 26°59′ E
877 m
CG-LL
05°54′ S, 22°25′ E
608 m
CG-LL
05°54′ S, 22°25′ E
608 m
CG-LL
CG-LL
ca 05°53′ S, 22°26′ E
05°54′ S, 22°25′ E
608 m
CG-LL
06°12′ S, 22°23′ E
610 m
CG-NK
CG-HK
ca 00°15′ N, 29°35′ E
11°40′ S, 27°26′ E
1209 m
CG-NK
CG-NK
CG-HK
00°20′ S, 28°36′ E
00°37′ S, 29°07′ E
ca 10°54′ S, 26°60′ E
1300 m
1809 m
CG-LM
CG-SK
CG-SN
07°10′ S, 23°42′ E
ca 02°00′ S, 28°45′ E
04°58′ S, 23°27′ E
841 m
397 m
CG-KU
04°50′ S, 18°44′ E
307 m
CG-BC
CG
CG-SK
CG-NK
CG-HL
05°50′ S, 13°03′ E
2089 m
CG
Lusolo Cave
(see: Lubudi)
Lusolo Grot
(see also: Grotte de Lusolo)
Luta Nzige
(see: Albert, Lake)
Lutunguru
Luvungi
Luzambo
(see: Lusambo)
Lwana River
Lwiko
(see also: Luiko)
Lwiro
(see: Luiro)
Lwiro Katana
(see: Luiro)
Mabali
Mabali
Mabanga
Mabanza
Mabaye
(see: Mabayi)
Mabaye, 10 km W
(see: Buhoro)
Mabayi
(see also: Mabaye)
Mabereni River
Maboya
(see: Maboyo)
Maboyo
(see also: Maboya)
Mabwe
Mabwe River
CG-LB
286
02°13′ S, 28°49′ E
ca 00°25′ N, 29°46′ E
08°56′ S, 27°12′ E 1513–1760
m
09°57′ S, 25°58′ E
1307 m
CG-HK
09°56′ S, 25°28′ E
CG-IT
ca 01°40′ N, 30°40′ E
CG-NK
CG-SK
CG-SN
00°28′ S, 28°49′ E
02°52′ S, 29°02′ E
04°58′ S, 23°27′ E
CG-SK
CG-SK
ca 01°56′ S, 28°28′ E
03°05′ S, 28°47′ E
CG-SK
02°14′ S, 28°48′ E
1646 m
CG-SK
02°14′ S, 28°48′ E
1646 m
CG-SK
CG-ET
CG-HU
RW-OU
BY-CI
02°20′ S, 28°50′ E
00°55′ S, 18°09′ E
04°22′ N, 29°47′ E
02°03′ S, 29°25′ E
02°43′ S, 29°15′ E
1612 m
1436 m
BY-CI
02°42′ S, 29°11′ E
1453 m
BY-CI
02°43′ S, 29°15′ E
1436 m
CG-HU
CG-NK
03°05′ N, 29°22′ E
00°19′ N, 29°20′ E
1374 m
CG-NK
00°19′ N, 29°20′ E
1374 m
CG-HL
CG-HL
08°39′ S, 26°31′ E
08°42′ S, 26°29′ E
574 m
590 m
1774 m
887 m
397 m
VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
Macaco
Madimao
Mai, Island
(see also: Île Mai)
Magachi
Maganga
(see also: Kisangani - Mambasa Road, km 122, Mangangu)
Maginda
Mahagi
(see also: Mahaji)
Mahagi Port
(see also: Mahaji Port)
Mahagi Territory
Mahaji
(see: Mahagi)
Mahaji Port
(see: Mahagi Port)
Mahysha Cave
(see: Mahyusha Cave)
Mahysha Grot
(see: Mahyusha Cave)
Mahyusa Cave
(see: Mahyusha Cave)
Mahyusha Cave
(see also: Grotte de Katana, Grotte de Mahysha, Grotte de Mahyusha,
Katana Cave, Katana Grot, Mahysha Cave, Mahysha Grot, Mahyusha Grot,
Mahyusa Cave)
Mahyusha Grot
(see: Mahyusha Cave)
Mai-Ndombe Province
Mai-Ndombe, Lake
(see also: Lake Mai-Ndombe, Lake Leopold II, Leopold II, Lake, Lac
Léopold II, Leopold II Meer, Leopold 2nd, Lake)
Mai-ya-Moto
(see: May ya Moto)
Maka
Makaia N’Tete
Makala
Makamba
Makamba Province
Makaw
Makayova River
Makayova River, lower
Make
Makengo
Makisabo
(see also: Makisabu)
Makisabu
(see: Makisabo)
Makpe
(see also: Makpe River, upper)
Makpe River, upper
(see: Makpe)
Makulakulu
(see: Mukulakulu)
Malandje
(see: Malandji)
Malandji
(see also: Malandje, Malanji)
Malanji
(see: Malandji)
Malela
Maleli
Malembe
Malili
Malingongo
Malulu
Mambabwanga Hill
(see: Batala’s Village)
Mambaka
Mambasa Territory
Mamudioma
Mandima
Mangangu
(see: Maganga)
Mangbwelu River
287
CG-KS
CG-HK
CG-TO
05°28′ S, 21°10′ E
11°40′ S, 27°29′ E
00°28′ N, 25°17′ E
RW-ES
CG-TO
01°20′ S, 30°34′ E
00°51′ N, 26°22′ E
CG-NK
CG-IT
00°49′ N, 29°54′ E
02°18′ N, 30°59′ E
850 m
1671 m
CG-IT
02°09′ N, 31°14′ E
679 m
CG-IT
CG-IT
02°18′ N, 30°59′ E
02°18′ N, 30°59′ E
1671 m
CG-IT
02°09′ N, 31°14′ E
679 m
CG-SK
02°13′ S, 28°50′ E
1480 m
CG-SK
02°13′ S, 28°50′ E
1480 m
CG-SK
02°13′ S, 28°50′ E
1480 m
CG-SK
02°13′ S, 28°50′ E
1480 m
CG-SK
02°13′ S, 28°50′ E
1480 m
CG
CG-MA
02°05′ S, 18°20′ E
CG-NK
00°14′ N, 29°43′ E
950 m
CG
CG-BC
CG-TO
BY-MA
BY
CG-MA
CG-NK
CG-NK
CG-BC
CG-KN
CG-NK
08°56′ S, 26°04′ E
05°33′ S, 13°02′ E
00°33′ N, 27°45′ E
04°08′ S, 29°48′ E
634 m
1472 m
03°29′ S, 18°19′ E
00°43′ N, 29°43′ E
00°43′ N, 29°43′ E
04°36′ S, 15°10′ E
04°46′ S, 16°19′ E
00°24′ N, 29°36′ E
315 m
750 m
750 m
CG-NK
00°24′ N, 29°36′ E
950 m
CG-HU
04°20′ N, 29°34′ E
CG-HU
04°20′ N, 29°34′ E
CG-HL
09°33′ S, 25°47′ E
828 m
CG-LL
05°56′ S, 22°18′ E
608 m
CG-LL
05°56′ S, 22°18′ E
608 m
CG-LL
05°56′ S, 22°18′ E
608 m
CG-BC
CG-HU
CG-NK
CG-HU
CG-NK
CG-NK
CG-NK
05°59′ S, 12°37′ E
04°20′ N, 29°37′ E
ca 01°26′ S, 28°04′ E
04°13′ N, 29°32′ E
00°44′ N, 29°40′ E
00°32′ N, 29°35′ E
00°30′ N, 29°15′ E
2m
CG-TO
CG-IT
CG-NK
CG-IT
CG-TO
00°51′ N, 27°33′ E
01°21′ N, 29°03′ E
00°40′ N, 29°38′ E
ca 01°21′ N, 29°03′ E
00°51′ N, 26°22′ E
1000 m
CG-NK
00°21′ N, 29°44′ E
1300 m
611 m
950 m
800 m
1000 m
European Journal of Taxonomy 382: 1–327 (2017)
Mani Cave or Nkela Cave or Mbumba-Nsi
(see also: Grotte de Mani ou Grotte de Nkela de Mbumba-Nsi, Mani Grot of
Nkela Grot van Mbumba-Nsi)
Mani Grot of Nkela Grot van Mbumba-Nsi
(see: Mani Cave or Nkela Cave or Mbumba-Nsi)
Maniema Province
Manono
(see also: Manono Katanga)
Manono Katanga
(see: Manono)
Marari
Marion, Lake
(see also: Lake Marion, Lac Marion)
Marumbi
Marunga mountains
Marungu
Masaba
Masabu
Masaka - Kigali
Masako
Masako km 15 Buta
Masha
Masibingi
Masimengo
Masombwe
Matadi
(see also: Matidi)
Matale
Matamba
Matata Cave
Matidi
(see: Matadi)
Matupi Cave
(see also: Grotte de Matupi, Matupi Grot)
Matupi Grot
(see: Matupi Cave)
Mauda
Maunzi
(see also: Mavunzi, Mawunzi, Mvuazi)
Mavunzi
(see: Maunzi)
Mawambi
Mawunzi
(see: Maunzi)
May ya Moto
(see also: Moto, Mai ya, Moto, May a, Moto, May ya, May ya Moto, sources
chaudes, Mai-ya-Moto)
May ya Moto, sources chaudes
(see: May ya Moto)
Mayumbe
Mayumbe area
Mayumbe, near
Mbalaka (Plantation)
Mbale
Mbandaka
Mbanza-Ngungu
(see also: Thysstad, Thysville)
Mbawu
M’baya Cave
(see: Baya Cave)
M’baya Grot
(see: Baya Cave)
Mbelekelo
Mbiye, Island
(see also: Île Mbiye)
Mboga
(see: Boga)
Mbuji-Mayi
(see also: Mbuy Mayi, Bakwanga)
Mbuji-Mayi, 32 km SE
Mbuy Mayi
(see: Mbuji-Mayi)
Mbuye-Bala River
Mbwambala
Medja
(see: Medje)
288
CG
CG
CG
CG-TG
07°18′ S, 27°25′ E
664 m
CG-TG
07°18′ S, 27°25′ E
664 m
CG-HU
CG
04°20′ N, 29°40′ E
CG
CG-TG
CG-TG
CG-KG
CG-KU
RW-KV
CG-TO
CG-TO
RW-OU
CG
CG-NK
CG-HK
CG-BC
07°30′ S, 29°59′ E
06°43′ S, 29°31′ E
04°33′ S, 16°54′ E
04°33′ S, 16°54′ E
01°60′ S, 30°11′ E
00°36′ N, 25°13′ E
00°40′ N, 25°14′ E
01°36′ S, 29°20′ E
1360 m
1722 m
597 m
1448 m
00°34′ N, 29°42′ E
09°05′ S, 27°12′ E
05°49′ S, 13°27′ E
800 m
1163 m
18 m
CG-SK
CG-LL
CG-IT
CG-BC
02°39′ S, 28°22′ E
05°57′ S, 22°16′ E
00°58′ N, 29°40′ E
05°49′ S, 13°27′ E
1217 m
619 m
1160 m
18 m
CG-IT
01°16′ N, 29°55′ E
1067 m
CG-IT
01°16′ N, 29°55′ E
1067 m
CG-HU
CG-BC
04°05′ N, 27°41′ E
05°19′ S, 15°07′ E
547 m
CG-BC
05°19′ S, 15°07′ E
547 m
CG-IT
CG-BC
01°04′ N, 28°34′ E
05°19′ S, 15°07′ E
653 m
547 m
CG-NK
00°14′ N, 29°43′ E
950 m
CG-NK
00°14′ N, 29°43′ E
950 m
CG-HU
CG-BC
CG-HU
CG-KU
CG
CG-ET
CG-BC
02°30′ N, 27°37′ E
ca 04°30′ S, 12°30′ E
ca 02°30′ N, 27°37′ E
05°00′ S, 18°53′ E
678 m
00°04′ N, 18°16′ E
05°15′ S, 14°52′ E
306 m
604 m
CG-LL
CG-HK
05°54′ S, 22°25′ E
11°52′ S, 27°27′ E
1279 m
CG-HK
11°52′ S, 27°27′ E
1279 m
CG-SK
CG-TO
03°20′ S, 28°04′ E
00°28′ N, 25°17′ E
1100 m
413 m
CG-IT
01°03′ N, 29°56′ E
1562 m
CG-KO
06°09′ S, 23°36′ E
549 m
CG-KO
CG-KO
ca 06°09′ S, 23°36′ E
06°09′ S, 23°36′ E
549 m
CG-HL
CG-KU
CG-HU
ca 09°10′ S, 27°04′ E
05°03′ S, 18°55′ E
02°25′ N, 27°18′ E
617 m
2085 m
VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
Medje
(see also: Medja)
Medje, about 30 mi [=48 km] ENE
(see: Pawa)
Menga
Meridi River
M’gayu
Mihindi, Lake
(see also: Lake Mihindi, Lac Mihindi)
Mikalaie (Mission)
(see: Mikalaya)
Mikalaya
(see also: St Joseph Mission, Mikalaya missie, Mikalaya Mission, Mission
de Mikalaya, Mikalaie (Mission), St. Jos. (Mission), Saint-Joseph (Mission),
Saint Joseph de Luluabourg (Mission), St. Joseph de Luluabourg Mission)
Mikalaya missie
(see: Mikalaya)
Mikalaya Mission
(see: Mikalaya)
Miki
Mimbulu
Mimuli
Mines de Kilo
(see: Kilomines)
Misotschi-Kabogo Highlands
Mission de Kinzambi
(see: Kinzambi Mission)
Mission de Liège Sacre-Coeur
(see: Kanzenze Mission)
Mission de Luanza
(see: Luanza Mission)
Mission de Mikalaya
(see: Mikalaya)
Mission Kasenga
(see: Kasenga)
Mission Kashiobwe
(see: Kashiobwe)
Mission Sacre Coeur
(see: Sacred Heart Mission)
Mission Sainte Thérèse
(see: Likasi)
Mission St Gerard
(see: Kapolowe)
Mission St Joseph
(see: Kananga)
Mistandunga
Misu
Mitwaba Territory
Moanda
(see: Muanda)
Moba
(see also: Baudhuinville, Baudouinville, Boudewijnstad)
Moba - Mplala Road, km 35
(see: Kikango)
Moba Territory
Mobutu Sese Seko, Lake
(see: Albert, Lake)
Moenda
Moenge
Moera
Moera’s village
(see also: Beni, 25 km NW)
Moero Meer
(see: Moero, Lake)
Moero, Lake
(see also: Moero Meer, Lake Moero, Lac Moero)
Mogbwamu River
Moka River
(see: Moko River)
Moko River
(see also: Moka River)
Molindi River
Moliro
Mondombe
Mondunga II Chefferie
Mongala Province
289
CG-HU
02°25′ N, 27°18′ E
CG-HU
02°31′ N, 27°04′ E
CG-KU
CG-HU
CG-TO
RW-ES
05°03′ S, 18°51′ E
04°40′ N, 29°33′ E
01°35′ N, 27°13′ E
01°32′ S, 30°42′ E
CG-LL
06°01′ S, 22°19′ E
608 m
CG-LL
06°01′ S, 22°19′ E
608 m
CG-LL
06°01′ S, 22°19′ E
608 m
CG-LL
06°01′ S, 22°19′ E
608 m
CG-SK
CG-HK
RW-ES
CG-IT
03°24′ S, 28°42′ E
11°43′ S, 27°19′ E
01°27′ S, 30°15′ E
01°48′ N, 30°14′ E
2060 m
1466 m
CG-SK
CG-KU
05°06′ S, 29°04′ E
04°59′ S, 18°46′ E
1880 m
312 m
CG-LB
10°31′ S, 25°12′ E
1157 m
CG-HK
08°42′ S, 28°42′ E
1132 m
CG-LL
06°01′ S, 22°19′ E
CG-HK
10°22′ S, 28°38′ E
938 m
CG-HK
09°39′ S, 28°37′ E
918 m
CG-KU
05°02′ S, 18°51′ E
CG-HK
10°59′ S, 26°44′ E
1318 m
CG-HK
11°02′ S, 26°57′ E
1135 m
CG-LL
05°54′ S, 22°25′ E
608 m
CG-MA
CG-IT
CG-HK
CG-BC
02°05′ S, 16°22′ E
01°45′ N, 30°18′ E
08°39′ S, 27°20′ E
05°56′ S, 12°21′ E
228 m
CG-TG
07°03′ S, 29°47′ E
763 m
CG-TG
06°37′ S, 29°30′ E
CG-TG
CG-IT
04°18′ N, 21°11′ E
ca 01°40′ N, 30°40′ E
CG-NK
CG-TO
CG-NK
CG-NK
00°25′ N, 29°46′ E
02°03′ N, 22°52′ E
00°39′ N, 29°30′ E
ca 00°35′ N, 29°25′ E
CG-HK
09°00′ S, 28°45′ E
CG-HK
09°00′ S, 28°45′ E
CG-HU
CG-HU
04°35′ N, 29°44′ E
04°09′ N, 29°43′ E
CG-HU
04°09′ N, 29°43′ E
CG-NK
CG-TG
CG-TP
CG-SU
CG
01°09′ S, 29°19′ E
08°13′ S, 30°34′ E
00°53′ S, 22°45′ E
617 m
0m
508 m
1000 m
914 m
1000 m
764 m
414 m
European Journal of Taxonomy 382: 1–327 (2017)
Mongbwala
(see: Mongbwalu)
Mongbwalu
(see also: Mongbwala)
Mongende
Mont Hoyo
(see: Hoyo, mount)
Mont Hoyo Cave
(see: Mount Hoyo Caves)
Mont Hoyo, Grottes
(see: Mount Hoyo Caves)
Montembessa Cave
(see also: Grotte de Montembessa)
Morubia
Moseda
(see: Mosenda)
Mosenda
(see also: Moseda, Mosendwa, Museya)
Mosendwa
(see: Mosenda)
Mosenge
Moto, Mai ya
(see: May ya Moto)
Moto, May a
(see: May ya Moto)
Moto, May ya
(see: May ya Moto)
Mount Gundu
(see: Gundu, mount)
Mount Homa
(see: Homa, Mount)
Mount Homa
(see: Hoyo, mount)
Mount Hoyo
(see: Hoyo, mount)
Mount Hoyo Caverns
(see: Mount Hoyo Caves)
Mount Hoyo Caves
(see also: Mount Hoyo Caverns, Mont Hoyo, Grottes, Irumu, 20 mi [=32
km] S, Mont Hoyo Cave, Kabatibi Cave)
Mount Kabobo
(see: Kabobo, mount)
Mount Kahuzi, W slope
Mount Korovi
(see: Korovi, mount)
Mount Mugusa
(see: Mugusa, mount)
Mount Muvo
(see: Muvo, mount)
Mount Ne
(see: Né, mount)
Mount Sombwe
Mount Tshiaberimu
(see: Tshiabirimu, mount)
Mount Tshiabirimu
(see: Tshiabirimu, mount)
Mount Wago
(see: Wago, mount)
Mouth of Congo
Mpala
M’Pala, Kazenze
Mpaza
Mpe
Mpinga Cave
(see: Mpinga Mission)
Mpinga Mission
(see also: Rutana, 46 km from, Mpinga Cave)
Mpopola Cave
(see also: Grotte de Mpopola, Mpopola Grot)
Mpopola Grot
(see: Mpopola Cave)
Muanda
(see also: Moanda)
Mudina
(see: Ndemba)
290
CG-IT
01°57′ N, 30°02′ E
1374 m
CG-IT
01°57′ N, 30°02′ E
1374 m
CG-MA
CG-IT
02°06′ S, 16°20′ E
01°13′ N, 29°49′ E
1200 m
CG-IT
01°13′ N, 29°49′ E
1200 m
CG-IT
01°13′ N, 29°49′ E
1200 m
CG-HU
CG-NK
04°11′ N, 29°29′ E
00°04′ N, 29°22′ E
1737 m
CG-NK
00°04′ N, 29°22′ E
1737 m
CG-NK
00°04′ N, 29°22′ E
1737 m
CG-KU
CG-NK
04°39′ S, 18°58′ E
00°14′ N, 29°43′ E
508 m
950 m
CG-NK
00°14′ N, 29°43′ E
950 m
CG-NK
00°14′ N, 29°43′ E
950 m
CG
CG-BU
CG-IT
01°13′ N, 29°49′ E
CG-IT
01°13′ N, 29°49′ E
1200 m
CG-IT
01°13′ N, 29°49′ E
1200 m
CG-IT
01°13′ N, 29°49′ E
1200 m
CG-IT
01°13′ N, 29°49′ E
1200 m
CG-TG
05°04′ S, 29°00′ E
2440 m
CG-SK
CG-IT
02°12′ S, 28°40′ E
01°39′ N, 30°36′ E
2560 m
2010 m
RW-SU
02°29′ S, 29°51′ E
1575 m
CG-NK
01°14′ S, 29°18′ E
CG-IT
01°42′ N, 30°39′ E
CG-HL
CG-NK
09°10′ S, 26°43′ E
ca 00°05′ S, 29°26′ E
CG-NK
ca 00°05′ S, 29°26′ E
CG-IT
01°45′ N, 30°40′ E
CG-BC
CG
CG-LB
CG-HU
CG-MA
BY-RT
ca 06°00′ S, 12°30′ E
06°45′ S, 29°31′ E
10°29′ S, 25°13′ E
04°14′ N, 29°37′ E
02°12′ S, 17°38′ E
03°44′ S, 30°11′ E
878 m
399 m
1930 m
BY-RT
03°44′ S, 30°11′ E
1930 m
CG-HL
09°56′ S, 25°58′ E
1450 m
CG
09°56′ S, 25°58′ E
1450 m
CG-BC
05°56′ S, 12°21′ E
0m
CG-LL
05°30′ S, 22°16′ E
2240 m
774 m
VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
Mudina Ndemba
(see: Ndemba)
Mueka
(see: Mweka)
Muezi Lupungu
Mufunga
Mugaba Cave
(see also: Grotte de Mugaba)
Mugarura
Mugeri Mission
(see also: Petit Seminaire de Mugeri)
Mugogwe
(see also: Colline Mugogwe)
Mugongo
Mugunga Meer
(see: Mugunga, Lake)
Mugunga, Lake
(see also: Lac Mugunga, Lake Mugunga, Mugunga Meer)
Mugusa, mount
(see also: Mount Mugusa)
Muhabura (Volcano)
(see also: Muhavura (Volcano))
Muhasi
(see: Muhazi)
Muhavura (Volcano)
(see: Muhabura (Volcano))
Muhazi
(see also: Colline Murunda, Muhasi)
Muhulu
Muite Nzige
(see: Albert, Lake)
Mukishi [Station]
Mukuen
(see also: Mukween, Mukwen, Mukwena)
Mukulakula
(see: Mukulakulu)
Mukulakulu
(see also: Makulakulu, Mukulakula)
Mukuta
(see: Mututa)
Mukwande River
Mukwaya
Mukween
(see: Mukuen)
Mukwen
(see: Mukuen)
Mukwena
(see: Mukuen)
Mulala
Mule
Mulima
Mulolo
Mulolo
Mulolo-Kisanga
(see: Lulola-Kasanga)
Mulombo
(see: Mulongo)
Mulong Cave
(see: Mulonga Cave)
Mulonga Cave
(see also: Grotte de Mulonga, Mulong Cave, Mulonga Grot)
Mulonga Grot
(see: Mulonga Cave)
Mulongo
(see also: Mulombo, Lualaba River)
Muluchu
Mulungu
(see also: Mulungu Miti)
Mulungu Kivu
Mulungu Miti
(see: Mulungu)
Mulungu-Tshibinda
(see: Tshibinda)
Mulungwishi
Mulungwishi, Cave
(see also: Grotte de Mulungwishi)
291
CG-LL
05°30′ S, 22°16′ E
CG-KS
04°51′ S, 21°34′ E
CG-HL
CG-HK
CG-SK
09°25′ S, 24°55′ E
10°58′ S, 26°47′ E
ca 02°19′ S, 28°31′ E
RW-OU
CG-SK
ca 01°42′ S, 30°45′ E
02°12′ S, 28°42′ E
RW-SU
02°31′ S, 29°49′ E
CG-NK
CG-NK
01°30′ S, 29°20′ E
01°37′ S, 29°08′ E
1500 m
CG-NK
01°37′ S, 29°08′ E
1500 m
RW-SU
02°29′ S, 29°51′ E
1575 m
RW-NO
01°23′ S, 29°43′ E
RW-ES
01°54′ S, 30°25′ E
RW-NO
01°23′ S, 29°43′ E
RW-ES
01°54′ S, 30°25′ E
1529 m
CG-NK
CG-IT
01°03′ S, 27°17′ E
ca 01°40′ N, 30°40′ E
671 m
CG-HL
CG-HK
08°30′ S, 24°44′ E
11°46′ S, 27°27′ E
CG-HL
09°33′ S, 25°47′ E
828 m
CG-HL
09°33′ S, 25°47′ E
828 m
CG-HK
11°21′ S, 27°47′ E
1229 m
CG-NK
CG-LM
CG-HK
00°19′ N, 29°42′ E
06°32′ S, 23°26′ E
11°46′ S, 27°27′ E
1300 m
CG-HK
11°46′ S, 27°27′ E
CG-HK
11°46′ S, 27°27′ E
CG-NK
CG-SK
CG
CG-LB
CG-MN
CG-MN
00°19′ N, 29°44′ E
02°24′ S, 28°33′ E
CG-TG
07°50′ S, 27°00′ E
CG-LB
09°56′ S, 25°58′ E
CG-LB
09°56′ S, 25°58′ E
CG-LB
09°56′ S, 25°58′ E
CG-TG
07°50′ S, 27°00′ E
578 m
RW-ES
CG-SK
01°32′ S, 30°37′ E
02°19′ S, 28°48′ E
1159 m
CG-SK
CG-SK
02°54′ S, 27°56′ E
02°19′ S, 28°48′ E
1159 m
1159 m
CG-SK
02°19′ S, 28°46′ E
1918 m
CG-LB
CG-LB
10°47′ S, 26°38′ E
10°47′ S, 26°38′ E
1366 m
466 m
1493 m
1529 m
1200 m
1936 m
09°56′ S, 26°58′ E
03°06′ S, 26°58′ E
03°06′ S, 26°59′ E
578 m
European Journal of Taxonomy 382: 1–327 (2017)
Mumba
Mumena
CG-NK
CG-HK
01°27′ S, 28°53′ E
11°04′ S, 28°08′ E
1765 m
1460 m
CG-HK
CG-HK
11°46′ S, 26°31′ E
11°04′ S, 28°08′ E
1460 m
BY-BI
CG-SK
CG-KU
CG-HL
03°55′ S, 29°35′ E
03°35′ S, 28°13′ E
05°28′ S, 18°35′ E
08°45′ S, 26°46′ E
1880 m
CG-HL
08°45′ S, 26°46′ E
890 m
CG-HK
CG-HK
10°58′ S, 26°32′ E
10°45′ S, 27°15′ E
1158 m
CG-NK
BY
BY-CA
RW-OU
RW-NO
01°08′ S, 29°27′ E
1275 m
03°11′ S, 30°40′ E
01°54′ S, 29°22′ E
01°29′ S, 29°36′ E
1826 m
1711 m
RW-NO
01°29′ S, 29°36′ E
CG-NK
CG-SK
CG-SK
CG-NK
CG-HL
01°20′ S, 29°34′ E
02°42′ S, 27°02′ E
01°37′ S, 28°18′ E
01°40′ S, 28°07′ E
07°36′ S, 24°33′ E
1500 m
CG-NK
00°04′ N, 29°22′ E
1737 m
RW-SU
02°32′ S, 29°52′ E
CG-MA
CG-HL
03°01′ S, 16°54′ E
07°36′ S, 24°33′ E
180 m
894 m
CG-HL
07°36′ S, 24°33′ E
894 m
CG-SK
CG-HK
ca 02°00′ S, 28°45′ E
11°28′ S, 27°52′ E
1183 m
CG
CG-NK
CG-KO
08°22′ S, 29°38′ E
01°09′ S, 29°27′ E
06°45′ S, 23°31′ E
1150 m
805 m
CG-NK
CG-NK
BY-BL
RW-OU
CG-HK
00°19′ N, 29°44′ E
00°19′ N, 29°45′ E
03°35′ S, 29°22′ E
01°36′ S, 29°23′ E
11°21′ S, 27°47′ E
2370 m
1229 m
CG-NK
00°20′ N, 29°45′ E
1338 m
CG-NK
01°14′ S, 29°18′ E
CG-HK
BY-MW
BY
CG-NK
CG-BC
ca 09°11′ S, 27°13′ E
03°30′ S, 29°47′ E
1764 m
00°32′ S, 29°17′ E
05°19′ S, 15°07′ E
547 m
CG-HK
CG-HK
10°45′ S, 27°15′ E
10°45′ S, 27°15′ E
1158 m
1158 m
CG-MN
04°26′ S, 26°48′ E
612 m
CG-MN
04°26′ S, 26°48′ E
612 m
CG-MN
04°35′ S, 27°08′ E
CG-MN
04°35′ S, 27°08′ E
CG-SK
02°13′ S, 28°51′ E
(see also: Lubumbashi, 70 km E, Mumene)
Mumena
Mumene
(see: Mumena)
Mumushwizi Valley
Munga
Mungamba
Munoi
(see: Munoi conluence)
Munoi conluence
(see also: Munoi)
Mura pumping Station
Muradingusha
(see: Mwadingusha)
Murambi
Muramvya Province
Murore
Murunda
Musanse
(see: Musanze)
Musanze
(see also: Colline Musanze, Musanse)
Musanze-Djomba
Musenge
Musenge
Musenge
Musenji
(see: Musindji)
Museya
(see: Mosenda)
Musha
(see also: Colline Musha)
Mushie
Musindji
(see also: Musingi, Musenji)
Musingi
(see: Musindji)
Musisi swamp area
Musonge
(see also: Lubumbashi, 69 km E )
Musosa, near
Musugeriza River
Mutembu Kaniki Kapangu
(see: Kaniki Kapangu)
Mutsora
Mutsora Station
Mutumba Mission
Mutura
Mututa
(see also: Mukuta)
Mutwanga
(see also: Kya irtumbi)
Muvo, mount
(see also: Mount Muvo)
Muvule, Cave 4 km from
Muyebe
Muyinga Province
Muyirimbo
Mvuazi
(see: Maunzi)
Mwadingusha
Mwadingusha
(see also: Muradingusha)
Mwanakusu
(see: Mwana-Kusu)
Mwana-Kusu
(see also: Mwanakusu)
Mwanakusu Cave
(see also: Grotte de Mwanakusu, Mwanakusu Grot)
Mwanakusu Grot
(see: Mwanakusu Cave)
Mwanda
292
586 m
894 m
1200 m
1394 m
VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
Mwanga Cave
(see also: Defrenne Cave, Defrenne Grot, Grotte de Defrenne, Grotte de
Mwanga, Jadotstad, 15 km S, Jadotville, 15 km S, Likasi, 15 km S, Mwanga
Grot)
Mwanga Grot
(see: Mwanga Cave)
Mwaro Province
Mwebe Bridge
Mweka
(see also: Mueka, Mweka Kasai)
Mweka Kasai
(see: Mweka)
Mweka Territory
Mwelampande
(see: Tenke)
Mwela-Pande Cave
(see: Tenke)
Mwenda-Katuka Piste
Mwenga
Mwenga Territory
Mwenge
Mwera
Mweya
Mwiga
Mwiru
(see: Luiro)
Mwittanzige
(see: Albert, Lake)
Myabikere
(see: Nyabikere)
Nadegbe River
Nagbarama
Nagera
(see: Nagero)
Nagero
(see also: Nagera)
Naimniam
Nakiluba
(see: Inakiluba)
Nakilubo
(see: Inakiluba)
Nakpanga
Nakpanga River
Naleza
Nambiliki / Dungu
Nampume
(see also: Nampune)
Nampune
(see: Nampume)
Nasosurro River
Ndama
Ndelele
Ndemba
(see also: Mudina, Mudina Ndemba)
Ndjili
Ndomo
Ndote
Ndotwe
Ndua
(see: Ndwa)
Ndwa
(see also: Ndua)
Né, mount
(see also: Mount Ne)
Nepoko Bomili
Netonna
New Beni
(see: Beni)
Ngamaye
Nganza
Nganza River
Ngayu [River]
(see also: Gayu [River])
N’Gayu area
Ngerere Lepi
Ngeze Kidoko
293
CG-HK
10°58′ S, 26°39′ E
CG-HK
10°58′ S, 26°39′ E
BY
CG-KU
CG-KS
05°03′ S, 18°50′ E
04°51′ S, 21°34′ E
466 m
CG-KS
04°51′ S, 21°34′ E
466 m
CG-KS
CG-HK
04°51′ S, 21°34′ E
10°35′ S, 26°07′ E
1512 m
CG-HK
10°35′ S, 26°07′ E
1512 m
CG-NK
CG-SK
CG-SK
CG-SK
CG-HK
BY-GI
CG-NK
CG-SK
ca 00°25′ N, 29°46′ E
03°02′ S, 28°26′ E
03°02′ S, 28°26′ E
03°42′ S, 28°10′ E
11°19′ S, 27°18′ E
03°29′ S, 29°56′ E
00°40′ S, 29°27′ E
02°14′ S, 28°48′ E
CG-IT
ca 01°40′ N, 30°40′ E
CG-NK
01°10′ S, 29°27′ E
1160 m
CG-HU
CG-HU
CG-HU
04°29′ N, 29°39′ E
04°17′ N, 29°32′ E
03°45′ N, 29°31′ E
753 m
CG-HU
03°45′ N, 29°31′ E
753 m
CG-BU
CG-HK
11°34′ S, 27°19′ E
1280 m
CG-HK
11°34′ S, 27°19′ E
1280 m
CG-HU
CG-HU
CG
CG-HU
CG-HU
04°36′ N, 29°34′ E
04°39′ N, 29°32′ E
15°27′ S, 27°20′ E
03°40′ N, 29°10′ E
04°21′ N, 29°49′ E
CG-HU
04°21′ N, 29°49′ E
CG
CG
CG-HU
CG-LL
04°22′ N, 29°47′ E
05°30′ S, 22°16′ E
900 m
CG-KN
CG-NK
CG-NK
CG-NK
CG-MA
04°28′ S, 15°21′ E
00°19′ N, 29°20′ E
00°11′ N, 29°39′ E
00°12′ S, 29°28′ E
02°09′ S, 16°25′ E
327 m
CG-MA
02°09′ S, 16°25′ E
CG-IT
01°42′ N, 30°39′ E
CG-TO
CG-BC
CG-NK
01°40′ N, 27°01′ E
05°58′ S, 12°27′ E
00°30′ N, 29°28′ E
CG-NK
CG-TG
CG-LL
CG-HU
00°44′ N, 29°38′ E
07°07′ S, 29°26′ E
ca 05°57′ S, 22°22′ E
01°35′ N, 27°13′ E
CG-HU
CG-NK
CG-IT
ca 01°35′ N, 27°13′ E
00°39′ N, 29°53′ E
01°16′ N, 30°25′ E
1643 m
1762 m
1646 m
940 m
1238 m
815 m
1200 m
650 m
European Journal of Taxonomy 382: 1–327 (2017)
Ngioni
CG-IT
01°34′ N, 30°15′ E
1332 m
CG-NK
CG-NK
00°19′ N, 29°45′ E
00°20′ N, 28°44′ E
905 m
CG-NK
01°41′ S, 29°13′ E
1499 m
CG-NK
00°20′ N, 28°44′ E
905 m
CG-KS
06°35′ S, 20°42′ E
528 m
CG-KS
06°35′ S, 20°42′ E
528 m
BY
CG-KU
CG-NK
CG-NK
04°45′ S, 18°11′ E
00°43′ S, 29°26′ E
01°10′ S, 29°27′ E
549 m
1160 m
CG-NK
01°15′ S, 28°45′ E
1297 m
CG-SK
CG-HU
CG-HU
CG-HU
CG-BU
CG-IT
02°07′ S, 28°54′ E
05°03′ N, 27°07′ E
03°42′ N, 27°52′ E
ca 03°42′ N, 27°52′ E
02°25′ N, 26°28′ E
02°15′ N, 31°07′ E
644 m
619 m
1382 m
CG-IT
02°15′ N, 31°07′ E
1382 m
CG-IT
CG-HL
CG-NK
02°10′ N, 30°39′ E
08°35′ S, 26°18′ E
01°31′ S, 29°15′ E
1793 m
CG-SK
CG-KN
CG
02°16′ S, 28°47′ E
04°10′ S, 15°55′ E
(see: Bunia)
Ngokai
Ngoma
(see also: N’goma)
Ngoma
(see: Goma)
N’goma
(see: Ngoma)
Ngombe
(see also: Gombe, N’gombe)
N’gombe
(see: Ngombe)
Ngozi Province
Ngula
Ngwiro
Niabikere
(see: Nyabikere)
Niabiondo
(see: Nyabiondo)
Niambasha
Niam-Niam
Niangara
Niangara Zone
Niapu
Niarembe
(see also: Niarombe)
Niarombe
(see: Niarembe)
Nioka
Nionga
Niragongo Volcano
(see: Nyirogongo Volcano)
Nkene
Nkiene
Nord-Kivu Province
(see also: North Kivu Province)
Nord Ubangui Province
(see also: North Ubangui Province)
North Kivu Province
(see: Nord-Kivu Province)
North Loango
North Ubangui Province
(see: Nord Ubangui Province)
Northern Prefecture
Nsele
(see: Kimpoko-Nsele)
Nshamba
Nsiamfumu
(see also: Vista)
Nsiana
N’soro
Ntango
Ntoto Cave
(see also: Grotte de Lukatu, Lukatu Cave, Lukatu Grot)
Nyabikere
(see also: Myabikere, Niabikere)
Nyabiondo
(see also: Niabiondo)
Nyabitembe River
Nyagatarugwa River Valley
Nyakaliba
Nyakariba River
Nyakibanda
(see also: Colline Nyakibanda)
Nyakichozi
Nyakinama
Nyamasheke
Nyamata
(see also: Colline Nyamata)
Nyambere
(see: Nyarusambo)
Nyambumba
Nyamugari Hill
Nyamunyunye
Nyangwe
(see also: Pene Bwebwe)
CG
CG
CG
CG
294
RW
CG-KN
04°14′ S, 15°33′ E
294 m
CG-LL
CG-BC
05°53′ S, 22°26′ E
05°52′ S, 12°17′ E
2m
CG
CG-HU
RW-OU
CG-TG
03°15′ N, 29°25′ E
02°36′ S, 29°13′ E
06°59′ S, 29°43′ E
827 m
CG-NK
01°10′ S, 29°27′ E
1160 m
CG-NK
01°15′ S, 28°45′ E
1297 m
CG-NK
BY-BI
CG
CG-NK
RW-SU
01°13′ S, 29°24′ E
03°55′ S, 29°35′ E
1225 m
1785 m
01°13′ S, 29°24′ E
02°38′ S, 29°41′ E
1325 m
2067 m
RW-ES
RW-OU
RW-OU
RW-ES
01°23′ S, 30°37′ E
01°34′ S, 29°38′ E
02°20′ S, 29°05′ E
02°10′ S, 30°06′ E
1250 m
1661 m
1581 m
CG-NK
01°29′ S, 29°21′ E
RW-OU
BY-MV
CG-SK
CG-MN
01°42′ S, 29°17′ E
03°18′ S, 29°45′ E
02°21′ S, 28°49′ E
04°13′ S, 26°11′ E
2590 m
507 m
VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
Nyankanda
Nyankende
Nyankokome
Nyankunde
Nyanza
Nyanza
(see: Nyanza-Lac)
Nyanza Territory
Nyanza-Lac
(see also: Nyanza)
Nyanza-Lac, 10 km N
Nyarusambo
(see also: Nyambere)
Nyarutovu
Nyiragongo Volcano
(see: Nyirogongo Volcano)
Nyirogongo Volcano
(see also: Volcan Nyirogongo, Niragongo Volcano, Volcan Niragong,
Nyiragongo Volcano, Volcan Nyiragongo, Tshamiagongo Volcano, Volcan
Tshamiagongo, Tshaninagongo, Kivu Volcano, Volcan du Kivu)
Nyumba Cave
Nyundo
Nyungwe Forest Reserve
(see also: Forêt Naturelle de Nyungwe)
Nzoro
(see also: Vankerckhovenville, Vankerckhovenstad)
Nzulu
(see also: N’Zulu)
N’Zulu
(see: Nzulu)
Obenge
Ogondjo
Old Kalonge
(see: Ugongo)
Old Kasindi
(see: Kasindi)
Olongo-Nsongo [River]
(see also: Batoko (River))
Omaniumdu
Ondo, Lake
(see also: Lake Ondo, Lac Ondo)
Oriental (former) Province
Oshwe
Oso River
Pahoni
(see: Paoni)
Panda
(see: Pande)
Panda Camp
(see: Likasi)
Pande
(see also: Panda)
Pandé, Forest N of
Panga
Pania-Mutombo
Pania-Mutombo
(see also: Pania-Mutombo)
(see: Pania-Mutombo)
Paoni
(see also: Pahoni)
Parc National Albert
(see: Virunga National Park)
Parc National de Garamba
(see: Garamba National Park)
Parc National de la Kagera
(see: Akagera Park)
Park National de Kundelungo
(see: Kundelungu National Park)
Paulis
(see: Isiro)
Pawa
(see also: Medje, about 30 mi [=48 km] ENE)
Pelenge, Gorges
Pempere Cave
(see also: Grotte de Pempere, Pimpin Cave)
Pempéré mountains
295
03°17′ S, 30°19′ E
03°55′ S, 30°00′ E
BY-RY
BY-MA
RW
CG-IT
RW-SU
BY-MA
01°34′ N, 30°15′ E
02°27′ S, 29°36′ E
04°21′ S, 29°36′ E
772 m
BY-MA
BY-MA
ca 04°20′ S, 29°36′ E
04°21′ S, 29°36′ E
772 m
BY-MA
CG-NK
ca 04°20′ S, 29°36′ E
01°29′ S, 29°21′ E
RW-SU
CG-NK
02°37′ S, 29°36′ E
01°31′ S, 29°15′ E
CG-NK
01°31′ S, 29°15′ E
RW-SU
RW-OU
RW-OU
02°39′ S, 29°42′ E
01°42′ S, 30°02′ E
02°31′ S, 29°18′ E
1594 m
2500 m
CG-HU
03°20′ N, 29°30′ E
791 m
CG-NK
00°36′ S, 29°05′ E
1500 m
CG-NK
00°36′ S, 29°05′ E
1500 m
CG-TO
CG-IT
CG-NK
01°23′ S, 25°02′ E
02°11′ N, 30°56′ E
00°20′ N, 29°49′ E
1700 m
2285 m
CG-NK
00°02′ S, 29°43′ E
1049 m
CG-MA
01°39′ S, 18°05′ E
CG-SN
CG-NK
03°21′ S, 23°16′ E
01°08′ S, 29°19′ E
464 m
CG
CG-MA
CG-NK
CG-IT
03°24′ S, 19°30′ E
01°04′ S, 27°25′ E
01°59′ N, 28°52′ E
368 m
913 m
CG-HK
10°37′ S, 26°50′ E
1140 m
CG-HK
10°59′ S, 26°44′ E
1318 m
CG-HK
10°37′ S, 26°50′ E
1140 m
CG-HK
CG-TO
CG-SN
CG-SN
ca 10°37′ S, 26°50′ E
01°51′ N, 26°25′ E
05°11′ S, 23°51′ E
05°11′ S, 23°51′ E
ca 1100 m
434 m
407 m
407 m
CG-IT
01°59′ N, 28°52′ E
913 m
CG-NK
01°00′ S, 29°15′ E
CG-HU
04°10′ N, 29°30′ E
RW-ES
01°30′ S, 30°30′ E
CG-HK
ca 10°30′ S, 27°30′ E
CG-HU
02°46′ N, 27°37′ E
730 m
CG-HU
02°32′ N, 27°42′ E
696 m
CG-HL
CG-HK
08°40′ S, 26°50′ E
10°58′ S, 26°47′ E
CG-HK
1553 m
2000 m
European Journal of Taxonomy 382: 1–327 (2017)
Pene Bwebwe
(see: Nyangwe)
Penge
Petit Seminaire de Mugeri
(see: Mugeri Mission)
PFNK
PFNK 15 and 17
PFNK.15
(see also: Piste Frontière Nord, km 15)
PFNK.16
(see also: Piste Frontière Nord, km 16)
PFSK 17 and 20
PFSK 8
PFSK.17
(see also: Piste Frontière Sud, km 17)
PFSK.20
(see also: Piste Frontière Sud, km 20)
PFSK.8
(see also: Piste Frontière Sud, km 8)
Pidigala Nord River
Pidigala River
Pierkat quarry
(see: Baya quarry)
Pili Pili
(see: Pilipili)
Pilipili
(see also: Pili Pili)
Pimpin Cave
(see: Pempere Cave)
Piste Frontière Nord, km 15
(see: PFNK.15)
Piste Frontière Nord, km 16
(see: PFNK.16)
Piste Frontière Sud, km 17
(see: PFSK.17)
Piste Frontière Sud, km 20
(see: PFSK.20)
Piste Frontière Sud, km 8
(see: PFSK.8)
PNA
(see: Virunga National Park)
Poko
Poko Territory
(see also: Poko Zone)
Poko Zone
(see: Poko Territory)
Popokabaka
Popokabaka Territory
Port Chaltin
(see: Aketi)
Port de Kindu
(see: Kindu)
Port-Empain
(see: Kindu)
PPK 51 and 56
PPK 72
PPK.10
PPK.51
Putnam, Camp
(see: Camp Putnam)
Pweto
Pweto Territory
Quatorze
(see also: Boende, 14 km N by road)
Ramangabo
(see: Rumangabo)
Rambira
Rambura
Ratabo Kinazi Gitorams
Resha
(see also: Bujumbura, 60 km S)
Rethy
(see also: Reti, Rety, Retsi, Retshi)
Reti
(see: Rethy)
Retshi
(see: Rethy)
296
CG-MN
04°13′ S, 26°11′ E
507 m
CG-HU
CG-SK
02°48′ N, 27°55′ E
02°12′ S, 28°42′ E
828 m
1493 m
CG-HU
CG-HU
CG-HU
04°02′ N, 29°05′ E
04°25′ N, 29°47′ E
04°25′ N, 29°47′ E
CG-HU
04°25′ N, 29°47′ E
CG-HU
CG-HU
CG-HU
04°18′ N, 29°55′ E
04°20′ N, 29°51′ E
04°18′ N, 29°55′ E
CG-HU
04°18′ N, 29°55′ E
CG-HU
04°20′ N, 29°51′ E
CG-HU
CG-HU
CG-HK
04°37′ N, 29°42′ E
04°35′ N, 29°32′ E
11°52′ S, 27°27′ E
CG-TO
00°33′ N, 27°44′ E
CG-TO
00°33′ N, 27°44′ E
CG-HK
10°58′ S, 26°47′ E
CG-HU
04°25′ N, 29°47′ E
CG-HU
04°25′ N, 29°47′ E
CG-HU
04°18′ N, 29°55′ E
CG-HU
04°18′ N, 29°55′ E
CG-HU
04°20′ N, 29°51′ E
CG-NK
01°00′ S, 29°15′ E
CG-BU
CG-BU
03°09′ N, 26°53′ E
ca 03°09′ N, 26°53′ E
CG-BU
ca 03°09′ N, 26°53′ E
CG-KG
CG-KG
CG-BU
05°42′ S, 16°35′ E
05°42′ S, 16°35′ E
02°50′ N, 23°56′ E
ca 500 m
CG-MN
02°57′ S, 25°57′ E
504 m
CG-MN
02°57′ S, 25°57′ E
504 m
CG-HU
CG-HU
CG-HU
CG-HU
CG-IT
04°06′ N, 29°25′ E
04°14′ N, 29°31′ E
03°46′ N, 29°29′ E
04°06′ N, 29°25′ E
01°24′ N, 28°35′ E
675 m
CG-HK
CG-HK
CG-TP
08°28′ S, 28°54′ E
08°28′ S, 28°54′ E
00°10′ N, 20°56′ E
326 m
CG-NK
01°21′ S, 29°22′ E
1587 m
CG-SK
CG-SK
RW-SU
BY-RU
02°03′ S, 28°54′ E
01°40′ S, 29°35′ E
02°13′ S, 29°58′ E
03°54′ S, 29°24′ E
CG-IT
02°05′ N, 30°54′ E
2116 m
CG-IT
02°05′ N, 30°54′ E
2116 m
CG-IT
02°05′ N, 30°54′ E
2116 m
1279 m
641 m
485 m
951 m
VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
Retsi
CG-IT
02°05′ N, 30°54′ E
2116 m
CG-IT
02°05′ N, 30°54′ E
2116 m
CG-NK
00°15′ S, 29°01′ E
1500 m
CG-TO
CG-NK
RW-SU
CG-BC
CG-NK
RW-SU
ca 01°00′ N, 26°45′ E
01°13′ S, 29°26′ E
02°25′ S, 29°10′ E
04°45′ S, 13°36′ E
00°40′ N, 29°43′ E
02°03′ S, 29°26′ E
BY-MA
04°16′ S, 29°35′ E
997 m
CG-TO
RW-SU
RW-SU
02°19′ N, 24°59′ E
02°30′ S, 29°44′ E
02°19′ S, 29°56′ E
1346 m
CG-NK
01°11′ S, 29°27′ E
1203 m
CG-NK
01°25′ S, 29°22′ E
2089 m
BY-MY
RW-NO
BY-BI
CG-NK
02°44′ S, 30°24′ E
01°30′ S, 29°38′ E
03°55′ S, 29°35′ E
00°47′ S, 29°17′ E
1651 m
1772 m
2170 m
1027 m
RW-ES
RW-ES
CG-NK
01°48′ S, 30°30′ E
02°09′ S, 30°37′ E
01°21′ S, 29°22′ E
1625 m
1709 m
1587 m
CG-NK
CG-NK
01°10′ S, 29°28′ E
01°21′ S, 29°22′ E
1200 m
1587 m
CG-IT
BY-RU
BY
CG-HU
CG-HU
CG-NK
BY-BL
BY-BB
BY-BL
01°34′ N, 29°07′ E
03°58′ S, 29°26′ E
809 m
03°11′ N, 27°52′ E
ca 03°11′ N, 27°52′ E
01°12′ S, 29°35′ E
03°21′ S, 29°29′ E
03°16′ S, 29°14′ E
03°21′ S, 29°16′ E
BY-BL
03°21′ S, 29°16′ E
BY-RT
BY
BY-RT
03°55′ S, 29°60′ E
1810 m
03°44′ S, 30°11′ E
1930 m
CG-NK
ca 00°08′ S, 29°35′ E
CG-NK
01°11′ S, 29°27′ E
CG-NK
ca 01°11′ N, 29°27′ E
CG-NK
01°11′ N, 29°27′ E
CG-NK
01°24′ S, 27°49′ E
(see: Rethy)
Rety
(see: Rethy)
Ribue Lya Mikako Cave
(see: Kibwe Lya Mikako Cave)
Risimu Zone
Rodahira River
Routabansougera
Ruango
Ruanoli-Mavea conluence
Rubengera (Mission)
(see also: Lubengera (Mission))
Rubindi
(see also: Grotte de Rubindi)
Rubi-Tele
Ruboma Ruhashya
Rubona
(see also: Colline Rubona)
Ruchuru
(see: Rutshuru)
Rugari
(see also: Lulenga)
Rugari Mission
Ruhengeri
Ruhinga Hill
Ruindi
(see: Rwindi)
Rukara
Rukira
Rumangabo
(see also: Ramangabo, Rumengabo)
Rumanura River
Rumengabo
(see: Rumangabo)
Rumoka
Rumonge
Rumonge Province
Rungu
Rungu Zone
Rusenyi River
Rushubi mine
Rusizi
Rusizi Delta
(see also: Russissi, bank)
Russissi, bank
(see: Rusizi Delta)
Rutana
Rutana Province
Rutana, 46 km from
(see: Mpinga Mission)
Rutanzige, Lake
(see: Edward, Lake)
Rutshuru
(see also: Ruchuru)
Rutshuru Territory
(see also: Rutshuru Zone)
Rutshuru Zone
(see: Rutshuru Territory)
Rutu [River]
(see: Utu [River])
Ruvara
Ruwenzori
Ruwenzori SE
Ruwenzori, Foothills
Ruwenzori, S side
Ruyigi
Ruyigi Province
Rwamagana
Rwanda
Rwankalo
Rwankwi
Rwegura
(see also: Kayanza, 15 km W)
Rwere Rwindi
Rwindi
(see also: Ruindi)
CG
CG-NK
CG-NK
CG-NK
CG-NK
BY-RY
BY
RW-ES
RW
RW
RW-NO
BY-KY
CG-NK
CG-NK
297
1200 m
1750 m
800 m
704 m
1300 m
ca 1743 m
1219 m
1341 m
00°25′ N, 29°50′ E
ca 00°30′ N, 29°50′ E
2286 m
ca 00°25′ N, 29°50′ E
03°29′ S, 30°15′ E
1842 m
01°57′ S, 30°26′ E
1528 m
01°32′ S, 29°32′ E
02°55′ S, 29°31′ E
2311 m
00°47′ S, 29°18′ E
00°47′ S, 29°17′ E
1027 m
European Journal of Taxonomy 382: 1–327 (2017)
Rwintare
(see also: Rwintare Mission)
Rwintare Mission
(see: Rwintare)
Sabe
Sacred Heart Mission
(see also: Mission Sacre Coeur)
Saga Saga Cave
(see also: Grotte de Saga Saga, Saga Saga Grot)
Saga Saga Grot
(see: Saga Saga Cave)
Saint Joseph de Luluabourg (Mission)
(see: Mikalaya)
Sainte Elisabeth
Saint-Joseph (Mission)
(see: Mikalaya)
Sakania
(see also: Sakania Mission)
Sakania Mission
(see: Sakania)
Sakania Territory
Saliboko
Salomoni Cave
Samba
Samboko River
Sange
Sankuru Province
Sciere Forest
Scierie Forest
(see also: Kindu, 30 km SW)
Semliki River
Semmio
(see: Zemio)
Semnio
(see: Zemio)
Senga
Shaba
Shaba (former) Province
(see: Katanga (former) Province)
Shabunda
Shabunda Territory
Shainkolobwe
(see: Shinkolobwe)
Shangi
(see also: Colline Shangi, Ishangi, Shangi Schad)
Shangi Schad
(see: Shangi)
Shangugu
(see also: Cyangugu)
Shibambo
(see also: Chibambo, Shibombo, Tshibambo)
Shibombo
(see: Shibambo)
Shindaika
Shinkolobwe
(see also: Chinkolobwe, Dethioux Farm, Cave I, Dethioux Farm, Cave II,
Dethioux Farm, Tantara Cave, Dethioux Hoeve, Grot I, Dethioux Hoeve,
Grot II, Dethioux Hoeve, Tantara Grot, Grotte de Tantara, Ferme Dethioux,
Grotte I, Ferme Dethioux, Grotte II, Ferme Dethioux, Shainkolobwe, Tantara
Cave, Trou aux Serpents)
Shyanda
(see also: Colline Shyanda)
Simbi Cave
Sinda River
Singitini
(see: Kisangani)
Sombe
(see also: Kiabo, 14 km upstream)
Soudan River
South Kivu Province
(see: Sud-Kivu Province)
South Ubangui Province
(see: Sud Ubangui Province)
Southern Prefecture
St Gerard Mission
(see: Kapolowe)
298
BY-MW
03°31′ S, 29°39′ E
2111 m
BY-MW
03°31′ S, 29°39′ E
2111 m
CG-IT
CG-KU
01°23′ N, 30°26′ E
05°02′ S, 18°51′ E
700 m
CG-IT
01°19′ N, 29°52′ E
1160 m
CG-IT
01°19′ N, 29°52′ E
1160 m
CG-LL
06°01′ S, 22°19′ E
608 m
CG-TO
CG-LL
01°33′ N, 25°20′ E
06°01′ S, 22°19′ E
432 m
608 m
CG-HK
12°45′ S, 28°34′ E
1278 m
CG-HK
12°45′ S, 28°34′ E
1278 m
CG-HK
CG-IT
CG-HK
CG-MN
CG-IT
CG-SK
CG
CG-MN
CG-MN
12°45′ S, 28°34′ E
01°42′ N, 30°29′ E
10°59′ S, 27°17′ E
04°38′ S, 26°22′ E
01°00′ N, 29°22′ E
03°06′ S, 29°15′ E
ca 04°58′ S, 23°27′ E
03°10′ S, 25°49′ E
03°10′ S, 25°49′ E
CG-NK
CG-BU
00°45′ N, 29°50′ E
05°03′ N, 27°07′ E
CG-BU
05°03′ N, 27°07′ E
CG-NK
CG
CG
01°40′ S, 28°07′ E
950 m
CG-SK
CG-SK
CG-LB
02°42′ S, 27°20′ E
02°42′ S, 27°20′ E
11°02′ S, 26°29′ E
596 m
RW-OU
02°24′ S, 29°00′ E
RW-OU
02°24′ S, 29°00′ E
RW-OU
02°29′ S, 28°54′ E
1537 m
CG-HK
10°18′ S, 28°37′ E
928 m
CG-HK
10°18′ S, 28°37′ E
928 m
CG-HK
CG-LB
11°39′ S, 27°41′ E
11°02′ S, 26°29′ E
1334 m
RW-SU
02°33′ S, 29°48′ E
1646 m
RW-SU
CG-IT
CG-TO
02°31′ S, 29°40′ E
01°02′ N, 30°04′ E
00°31′ N, 25°12′ E
830 m
447 m
CG-HL
08°50′ S, 26°02′ E
CG-HU
CG
04°35′ N, 29°35′ E
1874 m
610 m
850 m
1334 m
CG
RW
CG-HK
11°02′ S, 26°57′ E
1135 m
VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
St Joseph Mission
(see: Mikalaya)
St Therese Mission
(see: Likasi)
St. Jos. (Mission)
(see: Mikalaya)
St. Joseph de Luluabourg Mission
(see: Mikalaya)
St. Theresa Mission
(see: Likasi)
Stanleystad
(see: Kisangani)
Stanleyville
(see: Kisangani)
Sturi Forest
(see: Ituri Forest)
Sud-Kivu Province
(see also: South Kivu Province)
Sud Ubangui Province
(see also: South Ubangui Province)
Sukisa
Swanepoel I Cave
(see also: Grotte Swanepoel I, Swanepoel I Grot)
Swanepoel I Grot
(see: Swanepoel I Cave)
Swanepoel II Abyss
Taere River
(see also: Tayele (River))
Talama, 4 km SW
(see: Kabobo Forest)
Talatala
Tamba Tamba Cave b 12
(see also: Tamba Tamba Grot b 12)
Tamba Tamba Grot b 12
(see: Tamba Tamba Cave b 12)
Tamira
Tandala
Tanganika Province
(see: Tanganyika Province)
Tanganyika Meer, NW oever
(see: Lake Tanganyika, NW bank)
Tanganyika Province
(see also: Tanganika Province)
Tanganyika, Congo shore of Lake
Tanganyika, Lake, NW bank
(see: Lake Tanganyika, NW bank)
Tantara Cave
(see: Shinkolobwe)
TASOK (The American School of Kinshasa)
(see: Kinshasa, 10 km W)
Tayele (River)
(see: Taere River)
Tembwe
Temvo
Tenke
(see also: Likasi, 130 km N, Mwelampande, Mwela-Pande Cave)
Teturi
Teza, 2.3 km N, 0.7 km W
(see: Kibira National Park)
Thysstad
(see: Mbanza-Ngungu)
Thysville
(see: Mbanza-Ngungu)
Thysville, Cave
(see: Banjan Caves)
Thysville, Cave b 12 a
Thysville, Cave b 13 a
(see: Banjan Caves)
Tingasi
Tondu
Tongoni
(see: Kasongo)
Topetope
Tora
Tore
(see also: Tori)
299
CG-LL
06°01′ S, 22°19′ E
608 m
CG-HK
10°59′ S, 26°44′ E
1318 m
CG-LL
06°01′ S, 22°19′ E
608 m
CG-LL
06°01′ S, 22°19′ E
608 m
CG-HK
10°59′ S, 26°44′ E
1318 m
CG-TO
00°31′ N, 25°12′ E
447 m
CG-TO
00°31′ N, 25°12′ E
447 m
CG-IT
01°00′ N, 29°16′ E
CG
CG
CG-BU
CG-LB
02°19′ N, 24°59′ E
10°50′ S, 26°39′ E
470 m
ca 1350 m
CG-LB
10°50′ S, 26°39′ E
ca 1350 m
CG-LB
CG-HU
10°51′ S, 26°42′ E
04°33′ N, 29°47′ E
ca 1350 m
CG-SK
04°59′ S, 29°05′ E
CG-IT
CG-BC
01°15′ N, 29°50′ E
05°43′ S, 14°27′ E
CG-BC
05°43′ S, 14°27′ E
RW-OU
CG-SU
CG
01°34′ S, 29°22′ E
02°58′ N, 19°21′ E
CG-SK
ca 03°20′ S, 29°12′ E
1170 m
456 m
CG
CG
CG-SK
ca 06°00′ S, 29°11′ E
ca 03°20′ S, 29°12′ E
CG-LB
11°02′ S, 26°29′ E
CG-KN
ca 04°20′ S, 15°15′ E
CG-HU
04°33′ N, 29°47′ E
CG-TG
CG-BC
CG-HK
06°30′ S, 29°22′ E
05°30′ S, 13°01′ E
10°35′ S, 26°07′ E
1055 m
238 m
1512 m
CG-IT
BY-MV
01°04′ N, 29°08′ E
03°13′ S, 29°34′ E
CG-BC
05°15′ S, 14°52′ E
766 m
2200–2400
m
604 m
CG-BC
05°15′ S, 14°52′ E
604 m
CG-BC
05°16′ S, 14°53′ E
CG-BC
CG-BC
05°16′ S, 14°52′ E
05°16′ S, 14°53′ E
CG-HU
CG-ET
CG-MN
03°24′ N, 27°55′ E
00°53′ S, 18°09′ E
04°27′ S, 26°40′ E
CG-SK
BY-BI
CG-HU
02°16′ S, 28°03′ E
03°41′ S, 29°34′ E
04°20′ N, 29°50′ E
1334 m
625 m
2186 m
European Journal of Taxonomy 382: 1–327 (2017)
Tori
(see: Tore)
Trappist Mission
Trou aux Serpents
(see: Shinkolobwe)
Tschakala
(see: Tshakala)
Tschambutscha
(see: Tshambutschu)
Tschinputu
(see: Cinputu)
Tschuapa
(see: Tshuapa)
Tsebahu
Tshakala
(see also: Kisangani, 45 km E, Tschakala)
Tshamalenge Cave
(see also: Grotte de Tshamalenge)
Tshambi
Tshambutschu
(see also: Irangi, 10 km from, Tschambutscha)
Tshamiagongo Volcano
(see: Nyirogongo Volcano)
Tshaninagongo
(see: Nyirogongo Volcano)
Tshapona
(see: Tshiapona)
Tshebahu River
Tshiabirimu, mount
(see also: Mount Tshiabirimu, Mount Tshiaberimu)
Tshiapona
(see also: Tshapona)
Tshibaka
(see: Tshibaka-Kaji)
Tshibaka-Kaji
(see also: Tshibaka)
Tshibambo
(see: Shibambo)
Tshibasha
Tshibashi (River)
(see also: Tshibashie (River))
Tshibashie (River)
(see: Tshibashi (River))
Tshibati
Tshibinda
(see also: Mulungu-Tshibinda)
Tshibungu
Tshikapa
(see also: Chikapa)
Tshikapa Territory
Tshilinge (former) district
(see: Kasai Oriental Province)
Tshimputu
Tshisangwe
Tshopo
Tshopo Falls
(see also: Tshoppo Falls)
Tshopo Province
Tshoppo Falls
(see: Tshopo Falls)
Tshuapa
(see also: Tschuapa)
Tshuapa district
Tshuapa Province
Tshuapa River, upper
(see also: Haut Tshuapa, Tshuapa, Bovenloop)
Tshuapa, Bovenloop
(see: Tshuapa River, upper)
Tshyohaha, Lake
(see: Cyohaha, Lake)
Tumba, Lac
(see: Tumba, Lake)
Tumba, Lake
(see also: Lake Tumba, Tumba, Lac, Lac Tumba, Tumbe Meer)
Tumbe Meer
(see: Tumba, Lake)
300
CG-HU
04°20′ N, 29°50′ E
CG-KU
CG-LB
05°02′ S, 18°51′ E
11°02′ S, 26°29′ E
1334 m
CG-TO
00°27′ N, 25°32′ E
453 m
CG-SK
ca 01°58′ S, 28°30′ E
CG-LL
05°22′ S, 22°21′ E
CG-TP
00°14′ S, 20°42′ E
CG-IT
CG-TO
01°13′ N, 29°49′ E
00°27′ N, 25°32′ E
CG-HK
11°46′ S, 26°30′ E
CG-NK
CG-SK
00°44′ S, 29°14′ E
ca 01°58′ S, 28°30′ E
CG-NK
01°31′ S, 29°15′ E
CG-NK
01°31′ S, 29°15′ E
CG-HL
07°11′ S, 24°19′ E
826 m
CG-IT
CG-NK
01°19′ N, 29°52′ E
ca 00°05′ S, 29°26′ E
1200 m
CG-HL
07°11′ S, 24°19′ E
826 m
CG-LL
07°43′ S, 23°12′ E
882 m
CG-LL
07°43′ S, 23°12′ E
882 m
CG-HK
10°18′ S, 28°37′ E
928 m
CG-LL
CG-LL
05°50′ S, 22°25′ E
05°45′ S, 22°08′ E
CG-LL
05°45′ S, 22°08′ E
CG-SK
CG-SK
02°14′ S, 28°47′ E
02°19′ S, 28°46′ E
1918 m
CG-LL
CG-KS
05°38′ S, 22°16′ E
06°25′ S, 20°48′ E
608 m
485 m
CG-KS
CG
06°25′ S, 20°48′ E
CG-LL
CG-HK
CG-TO
CG-TO
05°48′ S, 22°19′ E
11°28′ S, 27°39′ E
00°28′ N, 25°33′ E
00°33′ N, 25°11′ E
CG
CG-TO
00°33′ N, 25°11′ E
CG-TP
00°14′ S, 20°42′ E
CG-TP
CG
CG-TP
00°13′ S, 20°52′ E
CG-TP
01°00′ S, 23°02′ E
RW-ES
02°15′ S, 30°00′ E
CG-ET
ca 00°48′ S, 18°03′ E
CG-ET
ca 00°48′ S, 18°03′ E
CG-ET
ca 00°48′ S, 18°03′ E
01°00′ S, 23°20′ E
453 m
1050 m
VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
Tumbwe
Tungudu
Ubuvumo lava hole
Uduku
(see also: Uduku Moke)
Uduku Moke
(see: Uduku)
Uele [River]
(see also: Welle [River])
Ugongo
(see also: Old Kalonge)
Ukaika
Uma
Umangi [Mission]
Université de Kisangani, Faculté des Sciences
(see: Kisangani)
Upemba National Park
Upper Congo
Upper Katanga Province
(see: Haut Katanga Province)
Upper Lomani Province
(see: Haut Lomami Province)
Upper Uele Province
(see: Haut Uele Province)
Usambiers
Usumbura
(see: Bujumbura)
Utu [River]
(see also: Rutu [River])
Utukuru
Uvinza
(see: Uvira)
Uvira
(see also: Uvinza)
Uvira Territory
Uwinka
Vaku
Vankerckhovenstad
(see: Nzoro)
Vankerckhovenville
(see: Nzoro)
Vieux Beni
(see also: Ancien Beni, Fort Beni)
Vinamont Farm
Virunga National Park
(see also: Albert National Park, Parc National Albert, PNA)
Vista
(see: Nsiamfumu)
Vitshumbi
(see also: Bitshumbi)
Volcan du Kivu
(see: Nyirogongo Volcano)
Volcan Niragong
(see: Nyirogongo Volcano)
Volcan Nyiragongo
(see: Nyirogongo Volcano)
Volcan Nyirogongo
(see: Nyirogongo Volcano)
Volcan Tshamiagongo
(see: Nyirogongo Volcano)
Vumbi
Wafania
(see also: Wafanya)
Wafanya
(see: Wafania)
Wagenia
Wago, mount
(see also: Mount Wago, Wasa Wago, mount, Blukwa, 10 km E)
Walikale
Walikale, 15 mi [=24 km] S
Wamba Territory
Wangata
(see: Bandaka)
Wasa
(see: Wasa Wago Forest)
Wasa Wago Forest
(see also: Wasa)
301
CG-NK
CG-NK
RW-NO
CG-HU
ca 01°00′ S, 29°15′ E
00°45′ N, 29°40′ E
840 m
04°32′ N, 29°33′ E
950 m
CG-HU
04°32′ N, 29°33′ E
950 m
CG
ca 03°41′ N, 26°03′ E
CG-NK
00°20′ N, 29°49′ E
2285 m
CG-IT
CG-TO
CG-TP
CG-TO
00°45′ N, 28°45′ E
00°32′ N, 25°54′ E
02°07′ S, 21°24′ E
00°31′ N, 25°12′ E
900 m
508 m
CG-HL
CG
CG
09°00′ S, 26°45′ E
447 m
CG
CG
CG
BY-BM
03°23′ S, 29°22′ E
CG-NK
01°24′ S, 27°49′ E
CG-HU
CG-SK
04°18′ N, 29°56′ E
03°24′ S, 29°09′ E
227 m
CG-SK
03°24′ S, 29°09′ E
227 m
CG-SK
RW-OU
CG-BC
CG-HU
03°24′ S, 29°08′ E
02°28′ S, 29°12′ E
05°15′ S, 13°10′ E
03°20′ N, 29°30′ E
351 m
791 m
CG-HU
03°20′ N, 29°30′ E
791 m
CG-NK
00°26′ N, 29°33′ E
585 m
CG-HK
CG-NK
11°41′ S, 27°29′ E
01°00′ S, 29°15′ E
CG-BC
05°52′ S, 12°17′ E
2m
CG-NK
00°41′ S, 29°23′ E
907 m
CG-NK
01°31′ S, 29°15′ E
CG-NK
01°31′ S, 29°15′ E
CG-NK
01°31′ S, 29°15′ E
CG-NK
01°31′ S, 29°15′ E
CG-NK
01°31′ S, 29°15′ E
CG-HL
CG-TP
08°38′ S, 25°58′ E
01°21′ S, 20°20′ E
932 m
396 m
CG-TP
01°21′ S, 20°20′ E
396 m
CG-TO
CG-IT
00°23′ N, 25°12′ E
01°45′ N, 30°40′ E
508 m
2240 m
CG-NK
CG-NK
CG-HU
CG-ET
01°25′ S, 28°03′ E
01°40′ S, 28°07′ E
703 m
00°04′ N, 18°16′ E
306 m
CG-IT
01°44′ N, 30°39′ E
1386 m
CG-IT
01°44′ N, 30°39′ E
1386 m
794 m
European Journal of Taxonomy 382: 1–327 (2017)
Wasa Wago, mount
(see: Wago, mount)
Washiha Forest
(see also: Bugarula, 6 km N)
Wasseige boerderij
(see: Wasseige Farm)
Wasseige Farm
(see also: Ferme Wasseige, Kipushi, 15 km from, Wasseige boerderij)
Weko Forest
Welle [River]
(see: Uele [River])
West Kasai (former) Province
(see: Kasai Occidental (former) Province)
Western Prefecture
Wilibadi River
Yaekela
Yakubuku
(see: Yakuluku)
Yakuluku
(see also: Yakubuku)
Yalokele
Yalosemba
(see also: Yalosembe, Yalosombo)
Yalosembe
(see: Yalosemba)
Yalosombo
(see: Yalosemba)
Yalusaka
Yamazo Yamono
Yambasa
(see: Yambata)
Yambata
(see also: Yambasa)
Yambombo
Yangambi
Yangambi Division Forest
Yaosuka
Yayama
Yema Mayombe
Yokamba Elongo
Yoko
Yolohotiri Cave
(see also: Grotte de Yolohotiri, Yolohotiri Grot)
Yolohotiri Grot
(see: Yolohotiri Cave)
Yolokairi
(see also: Jolohairi)
Zambi
Zega
(see: Zeya)
Zemio
(see also: Semmio, Semnio)
Zeya
(see also: Zega)
Zimet Quarter
Zobia
Zongo
Zookis
(see: Kisangani)
302
CG-IT
01°45′ N, 30°40′ E
2240 m
CG-SK
ca 02°00′ S, 29°04′ E
1500 m
CG-HK
11°50′ S, 27°20′ E
CG-HK
11°50′ S, 27°20′ E
CG-TO
CG
01°00′ N, 24°33′ E
04°07′ N, 22°26′ E
CG
RW
CG-HU
CG-TO
CG-HU
03°55′ N, 29°08′ E
00°48′ N, 24°18′ E
04°20′ N, 28°48′ E
508 m
697 m
CG-HU
04°20′ N, 28°48′ E
697 m
CG-ET
CG-MO
00°40′ S, 22°48′ E
02°35′ N, 21°47′ E
443 m
422 m
CG-MO
02°35′ N, 21°47′ E
422 m
CG-MO
02°35′ N, 21°47′ E
422 m
CG-TP
CG-MO
CG-MO
01°05′ S, 22°58′ E
02°35′ N, 21°47′ E
02°26′ N, 21°58′ E
420 m
414 m
CG-MO
02°26′ N, 21°58′ E
414 m
CG-MO
CG-TO
CG-TO
CG-TO
CG-TP
CG-BC
CG-TP
CG-TO
CG-IT
02°35′ N, 21°47′ E
00°47′ N, 24°28′ E
00°47′ N, 24°24′ E
00°45′ N, 24°29′ E
01°16′ S, 23°07′ E
05°45′ S, 12°17′ E
00°01′ N, 22°17′ E
00°17′ N, 25°17′ E
01°13′ N, 29°49′ E
CG-IT
01°13′ N, 29°49′ E
CG-IT
01°20′ N, 29°50′ E
1030 m
CG-BC
CG-IT
05°51′ S, 12°52′ E
01°45′ N, 30°44′ E
650 m
CG-BU
05°03′ N, 27°07′ E
CG-IT
01°45′ N, 30°44′ E
650 m
BY-BM
CG-BU
CG-SU
CG-TO
03°24′ S, 29°21′ E
02°58′ N, 25°56′ E
04°20′ N, 18°35′ E
00°31′ N, 25°12′ E
609 m
437 m
456 m
18 m
429 m
447 m
VAN CAKENBERGHE V. et al., The bats of Congo, Rwanda and Burundi revisited
Acknowledgements
We would like to extend our greatest thanks to our contact persons in the various museums (see “Summary
table of museum specimens” section), who provided the necessary information to enable us to make
this update, especially to Wim Wendelen (Royal Museum for Central Africa, Tervuren, Belgium) and
Tom Geerinckx (Royal Belgian Institute of Natural Sciences, Brussels, Belgium). The contribution of
Guy-Crispin Gembu Tungaluna and Prescott Musaba Akawa was inancially supported by the Flemish
Interuniversity Council (VLIR-UOS). Prescott Musaba was also supported by CEBioS, a programme
funded by the Directorate General for development Cooperation (DGD) and housed at the RBINS.
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Manuscript received: 31 August 2016
Manuscript accepted: 23 November 2016
Published on: 18 December 2017
Topic editor: Rudy Jocqué
Desk editor: Kristiaan Hoedemakers
Printed versions of all papers are also deposited in the libraries of the institutes that are members of the
EJT consortium: Muséum national d’Histoire naturelle, Paris, France; Botanic Garden Meise, Belgium;
Royal Museum for Central Africa, Tervuren, Belgium; Natural History Museum, London, United
Kingdom; Royal Belgian Institute of Natural Sciences, Brussels, Belgium; Natural History Museum
of Denmark, Copenhagen, Denmark; Naturalis Biodiversity Center, Leiden, the Netherlands; Museo
Nacional de Ciencias Naturales-CSIC, Madrid, Spain; Real Jardín Botánico de Madrid CSIC, Spain.
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