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WIKTROP - Weed Identification and Knowledge in the Tropical and Mediterranean areas
WIKTROP - Weed Identification and Knowledge in the Tropical and Mediterranean areas
SpeciesMapsDocumentsIDAO

Achyranthes aspera L.

Accepted
Achyranthes aspera L.
Achyranthes aspera L.
Achyranthes aspera L.
Achyranthes aspera L.
Achyranthes aspera L.
Achyranthes aspera L.
Achyranthes aspera L.
Achyranthes aspera L.
Achyranthes aspera L.
Achyranthes aspera L.
Achyranthes aspera L.
Achyranthes aspera L.
Achyranthes aspera L.
Achyranthes aspera L.
Achyranthes aspera L.
Achyranthes aspera L.
Achyranthes aspera L.
Achyranthes aspera L.
Achyranthes aspera L.
Achyranthes aspera L.
Achyranthes aspera L.
Achyranthes aspera L.
Achyranthes aspera L.
Achyranthes aspera L.
Achyranthes aspera L.
Achyranthes aspera L.
Achyranthes aspera L.
Achyranthes aspera L.
Achyranthes aspera L.
Achyranthes aspera L.
Achyranthes aspera L.
Achyranthes aspera L.
Achyranthes aspera L.
Achyranthes aspera L.
Achyranthes aspera L.
Achyranthes aspera L.
Achyranthes aspera L.
Achyranthes aspera L.
Achyranthes aspera L.
Achyranthes aspera L.
Achyranthes aspera L.
Achyranthes aspera L.
Achyranthes aspera L.
Achyranthes aspera L.
/Achyranthes aspera/762.jpg
🗒 Synonyms
synonymAchyranthes acuminata E. Mey. ex Cooke & Wright
synonymAchyranthes acuminata E.Mey. ex Cooke & Wright
synonymAchyranthes aspera f. subgrandifolia Suess.
synonymAchyranthes aspera var. aspera
synonymAchyranthes aspera var. australis (R.Br.) Domin
synonymAchyranthes aspera var. canescens (R.Br.) Drake
synonymAchyranthes aspera var. obtusifolia Suess.
synonymAchyranthes aspera var. simplex Millsp.
synonymAchyranthes asperoides Pires de Lima
synonymAchyranthes australis R.Br.
synonymAchyranthes canescens R. Br.
synonymAchyranthes canescens R.Br.
synonymAchyranthes daito-insularis Tawada
synonymAchyranthes ellipticifolia Stokes
synonymAchyranthes fruticosa Desf.
synonymAchyranthes grandifolia Moq.
synonymAchyranthes obovata Peter [Illegitimate]
synonymAchyranthes obovatifolia Stokes
synonymAchyranthes okinawensis Tawada
synonymAchyranthes robusta C.H.Wright
synonymAchyranthes sicula Roth
synonymAchyranthes tenuifolia Steud. [Invalid]
synonymCadelaria punctata Raf.
synonymCentrostachys aspera (L.) Standl.
synonymCentrostachys australis (R.Br.) Standl.
synonymCentrostachys canescens (R.Br.) Standl.
synonymCentrostachys grandifolia (Moq.) Standl.
synonymCentrostachys indica (L.) Standl.
synonymStachyarpagophora aspera (L.) M.Gómez
🗒 Common Names
Afrikaans
  • Langklits
  • Haak-en-steek-bossie
  • Grootklits
  • Haak en steek klitsbossie
  • Knapsekêrel, Langklitskafboom
Chinese
  • Tǔ niú xī, 土牛膝
Comorian
  • Soho
  • Ntsohomaele,Ntsohoho Mayele
Creoles and pidgins; French-based
  • Ké rat, Jandam, Kolan (Antilles)
Créole Maurice
  • Herbe sergent
Créole Réunion
  • Herbe d'Eugène
  • Herbe zinde
  • Queue de rat
  • Herbe d'Inde
  • La zinde
  • Herbe des jeunes
Créole Seychelles
  • Herbe sergent
English
  • Chaff flower
  • Burweed, Hug me close, Man better man, Prickly chafflower, Rattail, Rough chaff flower, Rough chaff tree
  • Devil's horsewhip
French
  • Herbe à bengali, Queue de rat (Nouvelle-Calédonie), Achyranthe rude, Cadelari, Marie pourrie
Hindi
  • Chirchita, Latjira
Italian
  • Achirante aspra, Scimitro
Malgache
  • Tsipotiky
Ndebele
  • Udombo
Other
  • Tsipotiky beTsipotiky lahy, Tsipotiky vavy, Sary tsipotiky, Tsipotiky kely (Kibushi, Mayotte)
  • Tsohoho m'ba bole, Tsohoho n'drume, Tsohoho m'ba n'titi, Tsohoho n'dzishe (Shimaore, Mayotte)
Portuguese
  • Folhas galo, Malpica, Pupalia
Sanskrit
  • Apamarga
Sotho
  • Bohomane
Spanish; Castilian
  • Anamú, Chile de perro, Venven, Rabo de gato (Cuba), Abrojo (Mexico)
Swazi
  • Lemanamana
Tamil
  • Naaveri, Nayuruvi
Tswana
  • Moxato
Zulu
  • IsiNama
  • IBundlubundlu
  • ULimilwengwe
  • USibambangubo
📚 Overview
Overview
Brief

Code

ACYAS

Growth form

broadleaf

Biological cycle

annual

Habitat

terrestrial

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ravi luckhun
StatusUNDER_CREATION
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References
    Diagnostic Keys
    Description
    Global description
    Achyranthes aspera is an erect plant, reaching 30 to 80 cm in height, abundantly branched and with hairy stem. The leaves are opposite, ovate, with acute or rounded apex. They are hairy and green on the upper surface, whilst smooth on the lower surface. The young leaves are silvery in colour. The spikes are very long and curved at the top. The flowers are very small, numerous, closely spaced, and green in colour. They are erect facing upward before flowering and bend down at fruiting. The fruits are small, dry, and brownish in colour. It is one-seeded, and can readily adhere to animals and clothing with their spines.Several varieties are present in Reunion, one of them being the variety sicula, characterised by a usually acuminate leaf at the top, and the variety aspersa with rounded end, apiculate or abruptly reduced into a short and narrow acumen.


    Cotyledons

    Cotyledons are elliptical to linear, 20 mm long by 3 mm wide, and the petiole is 5 mm long. Cotyledon attenuates to an acute angle at the leaf base and tip.

    First leaves

    The first leaves are simple, opposite, and ovate in shape. Margin is entire. The upper face is green and pubescent while the lower side is silvery and pubescent.

    Growth habit

    This is a slender, erect, and multi-branched herb, reaching up to 2 m high.

    Underground system

    It has a taproot.

    Stem

    Stems are full, squarish with swollen nodes. The surfaces of the stem have longitudinal grooves. Finely hairy to hairless when older.

    Leaf

    Leaves are simple, opposite and petiolate. Ovate to elliptic lamina (sometimes almost circular). They are 4 to 9 cm long by 2 - 4 cm wide. The margin is entire. Both surfaces are covered in short hairs although older leaves can be smooth. Each leaf has 4 to 9 arching veins.

    Inflorescence

    Long upright, terminal spikes. 10 to 50 cm in length.

    Flower

    Flowers are scaly, very small and numerous. They are mostly green, but can contain purples and pinks. They have no petal.  Calyx with 3 to 5 scarious sepals lanceolate, acute at the top. Outside, 2 spiny bracts, arched at the top and the bottom third of which is expanded in membranous wings. Flowers are pitched upwards when in bud, spread whilst flowering and then fold down to form fruits. The flowers start opening from the base of the inflorescence.

    Fruit

    Straw-coloured utricule, surrounded by the spiky perianth, 2.5-2.8 mm long, 1-1.5 mm wide, rounded at the base, with truncate or depressed apex; pointed downwards and pressed against the flowering stalk; indehiscent, 1-seeded, thin-walled; enclosed by persistent perianth and bracts,detaching easily from rachis.


    Seed

    Seed 2 to 3 mm long, 1 to 1.5 mm wide, truncate above, reddish to dark brown and shiny, enclosed in chaffy calyx parts that remain attached.
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      No Data
      📚 Natural History
      Life Cycle

      Life cycle

      Annual
      Annual

      China: Achyranthes aspera flowers from June to August and fruits in October.
      Nicaragua: Achyranthes aspera flowers and fruits from July to March.

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        Cyclicity

        Northern Cameroon: Achyranthes aspera is a weed growing at the middle and especially at the end of the cropping season. Germination occurs in June, regardless of plowing, when rains are abundant and regular. This phase of germination is very short (1 to 2 weeks). In unweeded fields, flowering starts late August and fruiting in October, a full cycle of four months. In contrast, in crops regularly weeded and mounded, a new short germination phase can occur in September, after the last cultural operation. The development cycle length is reduced to one month, from emergence to fruiting. The flowering of this species is induced by the reduction of the length of day. The plant disappears after beginning of the dry season (November-December).
        Mayotte: It is an annual plant which grows all year round.
        New Caledonia: Germination occurs after the first rains, growth is then fast enough to flowering that occurs in 1 or 2 months. Fruiting occurs until cool season before the plant dies.
        West Indies: Achyranthes aspera flowers and fruits all year round.

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          Reproduction
          This annual weed reproduces by seed. The seeds are dispersed by hooking into the fur of animals or by falling naturally.
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            Morphology

            Type of prefoliation

            Leaf ratio medium
            Leaf ratio medium
            Broad leaves
            Broad leaves

            Equality of opposite leaves

            Opposite leaves equal
            Opposite leaves equal

            Latex

            Without latex
            Without latex

            Stem section

            Square
            Square
            Ridged or grooved
            Ridged or grooved

            Root type

            Taproot
            Taproot

            Stipule type

            No stipule
            No stipule

            Fruit type

            Grain of grasses
            Grain of grasses

            Lamina base

            attenuate
            attenuate

            Lamina margin

            hairy
            hairy
            entire
            entire

            Lamina apex

            apiculate
            apiculate
            attenuate
            attenuate
            acuminate
            acuminate
            obtuse
            obtuse
            rounded
            rounded

            Upperface pilosity

            Less hairy
            Less hairy
            Dense hairy
            Dense hairy

            Upperface hair type

            Pubescent
            Pubescent
            Appressed
            Appressed

            Simple leaf type

            Lamina elliptic
            Lamina elliptic

            Lamina Veination

            in arc
            in arc
            pennate
            pennate

            Stem pilosity

            Less hairy
            Less hairy

            Life form

            Broadleaf plant
            Broadleaf plant
            Ecology

            Commonly found in disturbed areas. Prefers moist soil but can grow well in dry areas.

            Northern Cameroon: Achyranthes aspera develops from the Sudanese to Sahelo-Sudanese regions. It is present in 20-30% of cultivated fields. In Sudano-Sahelian region, this species develops only on soils with high water retention (fersialitic soils) or with a water table near the surface (alluvial, waterlogged soils). In the Sudanese region, it develops over a wider range of soils, although it is very rare on ferruginous soils with degraded sandy surface horizon.
            Central Africa: Achyranthes aspera grows in humid and shady places, frequent in swamps, mud of the water's edge, but also ubiquitous (savannah, sclerophyllous forest), thanks to a remarkable adaptability and to the process of dissemination (passive zoochory) of its fruits; it constitutes an invasive weed of the edge of roads, inhabited places, crops, pastures, fallow lands, following man even in dry stations, up to 1,600 m of altitude in the mountains of Kivu.
            Comoros
            : The species is present in all the islands, both in humid and dry environments, up to 400 m altitude.
            Madagascar: A. aspera is a fairly common species of rainfed crops in warm areas (low and medium altitudes), or on relatively rich soils  in recently cleared land plots and poorly maintained.
            Mauritius: A. aspera is a ruderal species in disturbed areas and along roadsides, and also a weed of crops. It is more common in the northern part of the island.
            Mayotte: Achyranthes aspera is a weed that is very present in ylang-tree plots and in forage crops. It can be found in fruit and food crops. It is mainly present in the dry zone of the south of the island, but it can also be found in the north.
            New Caledonia: A. aspera is a weed of fallow land, disturbed areas and pastures. It particularly enjoys moist soil but grows well in dry soils of the coast and back over the islets.
            Reunion: The species is common in Reunion island, occurring evenly from 0-900 m altitude. It grows particularly along rivers, in wet places, in disturbed habitats and, in general, as ruderal species and weed in irrigated plots of the West Coast.
            Seychelles: This species is observed from coastal areas to higher altitudes.
            West Indies: Achyranthes aspera is typically a very ubiquitous ruderal species, present from 0 to 1000 m altitude.

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              No Data
              📚 Habitat and Distribution
              Description

              Geographical distibution

              Madagascar
              Madagascar
              Reunion Island
              Reunion Island
              Comoros
              Comoros
              Mauritius
              Mauritius
              Seychelles
              Seychelles
              Origin

              Native to southern Asia, Australia, and some Pacific Islands.

              World Distribution

              Widespread in the tropics.

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                No Data
                📚 Occurrence
                No Data
                📚 Demography and Conservation
                Risk Statement

                Local weediness


                Northern Cameroon: A. aspera is a species characteristic of non-intensified or less intensified crops including traditional food crops (sorghum, groundnut). It is common in the recently cleared land in crops where pre-emergence herbicides are not applied and fertilizers less used. It is absent from older, intensively cultivated plots, receiving herbicides regularly. But this type of cropping system is important in the Sudanese region, which explains the low frequency of A. aspera. In addition, this species is easily removed by repeated weeding.

                Comoros: A. aspera is a weed present in cassava plantations and at the border of all crop fields.

                Madagascar: A. aspera is a common species in field edges and abandoned plots. It is less intrusive and rarely interfering. In the crop, it is relatively less frequent but sometimes locally abundant and harmful to crops that have a rather long cycle such as cotton, cassava. Due to its late vegetative development, it hinders harvesting operations (cotton, corn, peanuts, cassava).

                Mauritius: A. aspera is an occasional weed in cultivated fields,  with a low competitveness.

                New Caledonia: A. aspera is an introduced species: it was observed from the late 18th century. This is now a common species throughout the territory at rather low to medium altitudes. A. aspera is not an invasive species in pastures but its development is often related to overgrazing, it constitutes with other minor weeds a set of non or less patatable species whose abundance is involved in the degradation of pastures and decreases the production of forage species.

                Reunion: A. aspera is present in 13% of surveys with an average cover of 7-15%. Present in 17% of surveys in sugarcane and 3% in vegetable crops, it does not constitute any particular problem when the weeds is well controlled. Conversely,  its cover can reach from 30 to 50% due to poor weed control.

                Seychelles: A. aspera is a weed of crops, cultivated areas and fallow lands.

                Arouna Seechurn
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                  No Data
                  📚 Uses and Management
                  Uses

                  Food: Leaves of Achyranthes aspera can be eaten.
                  Medicinal: Achyranthes aspera is traditionally used to treat chest pain and stomach ache. In Chad, the boiled leaves are used against scabies (Yambata) and provide a remedy for headaches (Dundusana); incinerated after boiling, they yield cooking salt (Abok). In India, it is used to prepare potash and is said to be a remedy for scorpion stings.
                  Other: Fruits are eaten by birds.

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                    Management

                    Local Management

                    Madagascar: A. aspera is easily controlled by uprooting. Little information is available on its sensitivity to herbicides: A. aspera is well managed by postemergence 2,4-D or glyphosate. A. aspera is well controlled by a cover crop.

                    New Caledonia: In pastures, it is appropriate to prevent germination and propagation of A. aspera , an annual species, to a maxmum by maintaining a dense grassy cover. Livestock having stayed in infested areas must be quarantined. Isolated plants can be easily uprooted by hand with gloves. For established on stands, overgrazing of infested areas at an early stage of growth can contribute to weed out this species. Slashing only causes regrowth of the plant and can contribute to its spread, if carried to fruition. So we complement it with a herbicide spraying in targeted on isolated plots or in full if more diffuse infestation. Preference will be given to conventional herbicides such as 2,4-D, selective to grasses.

                    Reunion: Regular use of herbicides or repeated weeding limit its development in crops.
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                      📚 Information Listing
                      References
                      1. Husson, O., H. Charpentier, F.-X. Chabaud, K. Naudin, Rakotondramanana et L. Séguy (2010). Flore des jachères et adventices des cultures. Annexe 1 : les principales plantes de jachères et adventices des cultures à Madagascar. In : Manuel pratique du semis direct à Madagascar. Annexe 1 - Antananarivo : GSDM/CIRAD, 2010 : 64 p.
                      1. Akobundu I.O. & Agyakwa C.W., 1989. Guide des adventices d'Afrique de l'Ouest. Institut international d'agriculture tropicale. Ibadan, Nigeria, 521 p.
                      1. Berhaut J., 1967. Flore du Sénégal. 2ème éd. Clairafrique éd., Dakar, Sénégal, 485p.
                      1. Le Bourgeois, T., A. Carrara, M. Dodet, W. Dogley, A. Gaungoo, P. Grard, Y. Ibrahim, E. Jeuffrault, G. Lebreton, P. Poilecot, J. Prosperi, J. A. Randriamampianina, A. P. Andrianaivo and F. Théveny (2008). Advent-OI : Principales adventices des îles du sud-ouest de l'Océan Indien. Cédérom. Montpellier, France, Cirad ed.
                      1. Grard, P., T. Le Bourgeois, J. Rodenburg, P. Marnotte, A. Carrara, R. Irakiza, D. Makokha, G. kyalo, K. Aloys, K. Iswaria, N. Nguyen and G. Tzelepoglou (2012). AFROweeds V.1.0: African weeds of rice. Cédérom. Montpellier, France & Cotonou, Bénin, Cirad-AfricaRice eds.
                      1. Berhaut J., 1971. Flore illustrée du Sénégal. Tome 1. Clairafrique éd., Dakar, Sénégal, 626 p.
                      1. Donfack P., 1993. Etude de la dynamique de la végétation après abandon de la culture au Nord-Cameroun. Thèse Dc. 3ème cycle , Faculté des sciences, Univ. de Yaoundé, Cameroun, 192 p.
                      1. Hutchinson J., Dalziel J. M., Keay R. W. J. & Hepper F. N., 1954. Flora of West Tropical Africa. Vol. I part. 1. 2ème éd. The Whitefriars Press ed., London & Tonbridge, 295 p.
                      1. Ivens G. W., 1989. East African Weeds and Their Control. Oxford University Press, Nairobi, Kenya, 289p.
                      1. Ivens G. W., Moody K. & Egunjobi J. K., 1978. West African Weeds. Oxford University Press, Ibadan, Nigeria, 255 p.
                      1. le Bourgeois Th., 1993. Les mauvaises herbes dans la rotation cotonnière au Nord-Cameroun (Afrique) - Amplitude d'habitat et degré d'infestation - Cycle de développement.. Thèse USTL Montpellier II, Montpellier, France, 241p.
                      1. Blanfort, V., F. Desmoulins, J. Prosperi, T. Le Bourgeois, R. Guiglion and P. Grard (2010). AdvenPaC V.1.0 : Adventices et plantes à conflit d'intérêt des Pâturages de Nouvelle-Calédonie. Montpellier, France, IAC, Cirad.
                      1. Traoré H., 1991. Influence des facteurs agro-écologiques sur la constitution des communautés adventices des principales cultures céréalières (sorgho, mil, maïs) du Burkina-Faso.
                      2. Barthelat, F. 2019. La Flore illustrée de Mayotte. Meze, Paris, France, Collection Inventaires et Biodiversité, Biotope – Muséum national d’Histoire naturelle. 487 p.
                      3. Huat, J., Nagy, M., Carpente, A., Schwartz, M., Le Bourgeois, T. & Marnotte, P. 2021. Guide de la flore spontannée des agrosystèmes de Mayotte. Montpellier, Cirad. 150 p.
                      4. Fournet, J. 2002. Flore illustrée des phanérogames de Guadeloupe et de Martinique. Montpellier, France, Cirad, Gondwana éditions.
                      1. Swaziland Alien Plants Database
                      2. Blanfort, V., F. Desmoulins, J. Prosperi, T. Le Bourgeois, R. Guiglion and P. Grard (2010). AdvenPaC V.1.0 : Adventices et plantes à conflit d'intérêt des Pâturages de Nouvelle-Calédonie. Montpellier, France, IAC, Cirad
                      3. Le Bourgeois, T., A. Carrara, M. Dodet, W. Dogley, A. Gaungoo, P. Grard, Y. Ibrahim, E. Jeuffrault, G. Lebreton, P. Poilecot, J. Prosperi, J. A. Randriamampianina, A. P. Andrianaivo and F. Théveny (2008). Advent-OI : Principales adventices des îles du sud-ouest de l'Océan Indien. Cirad. Montpellier, France, Cirad.
                      4. US Forest Service, Pacific Island Ecosystems at Risk (PIER)
                      5. Bio-NET EAFRINET
                      6. Australian Tropical Rainforest Plants
                      7. Le Bourgeois, T., P. Grard, L. C. Foxcroft, D. Thompson, A. Carrara, A. Guézou, R. W. Taylor and T. Marshall (2013). Pl@ntInvasive-Kruger V.1.0 : Alien plants of the Kruger National Park. Cdrom. Montpellier, France, Skukuza, South Africa, Cirad-SANparks-SAEON eds.
                      Information Listing > References
                      1. Husson, O., H. Charpentier, F.-X. Chabaud, K. Naudin, Rakotondramanana et L. Séguy (2010). Flore des jachères et adventices des cultures. Annexe 1 : les principales plantes de jachères et adventices des cultures à Madagascar. In : Manuel pratique du semis direct à Madagascar. Annexe 1 - Antananarivo : GSDM/CIRAD, 2010 : 64 p.
                      2. Agis http://www.agis.agric.za/wip/
                      3. Adventilles http://data.plantnet-project.org/adventilles/_design/datamanager/index.html#/viewdoc/mm_9c11b4afef852eb758710127d08d6399##270f412b7ed9aa59543fced652006b04/0
                      4. Lucid http://keys.lucidcentral.org/keys/v3/eafrinet/weeds/key/weeds/Media/Html/Achyranthes_aspera_(Devils_Horsewhip).htm
                      5. Sntc http://www.sntc.org.sz/alienplants/speciesinfo.asp?spid=36
                      6. Flora of Zimbabwe http://www.zimbabweflora.co.zw/speciesdata/species.php?species_id=122450
                      7. Akobundu I.O. & Agyakwa C.W., 1989. Guide des adventices d'Afrique de l'Ouest. Institut international d'agriculture tropicale. Ibadan, Nigeria, 521 p.
                      8. Berhaut J., 1967. Flore du Sénégal. 2ème éd. Clairafrique éd., Dakar, Sénégal, 485p.
                      9. Le Bourgeois, T., A. Carrara, M. Dodet, W. Dogley, A. Gaungoo, P. Grard, Y. Ibrahim, E. Jeuffrault, G. Lebreton, P. Poilecot, J. Prosperi, J. A. Randriamampianina, A. P. Andrianaivo and F. Théveny (2008). Advent-OI : Principales adventices des îles du sud-ouest de l'Océan Indien. Cédérom. Montpellier, France, Cirad ed.
                      10. Grard, P., T. Le Bourgeois, J. Rodenburg, P. Marnotte, A. Carrara, R. Irakiza, D. Makokha, G. kyalo, K. Aloys, K. Iswaria, N. Nguyen and G. Tzelepoglou (2012). AFROweeds V.1.0: African weeds of rice. Cédérom. Montpellier, France & Cotonou, Bénin, Cirad-AfricaRice eds.
                      11. Berhaut J., 1971. Flore illustrée du Sénégal. Tome 1. Clairafrique éd., Dakar, Sénégal, 626 p.
                      12. Donfack P., 1993. Etude de la dynamique de la végétation après abandon de la culture au Nord-Cameroun. Thèse Dc. 3ème cycle , Faculté des sciences, Univ. de Yaoundé, Cameroun, 192 p.
                      13. Hutchinson J., Dalziel J. M., Keay R. W. J. & Hepper F. N., 1954. Flora of West Tropical Africa. Vol. I part. 1. 2ème éd. The Whitefriars Press ed., London & Tonbridge, 295 p.
                      14. Ivens G. W., 1989. East African Weeds and Their Control. Oxford University Press, Nairobi, Kenya, 289p.
                      15. Ivens G. W., Moody K. & Egunjobi J. K., 1978. West African Weeds. Oxford University Press, Ibadan, Nigeria, 255 p.
                      16. Trin http://keys.trin.org.au/key-server/data/0e0f0504-0103-430d-8004-060d07080d04/media/Html/taxon/Achyranthes_aspera.htm
                      17. le Bourgeois Th., 1993. Les mauvaises herbes dans la rotation cotonnière au Nord-Cameroun (Afrique) - Amplitude d'habitat et degré d'infestation - Cycle de développement.. Thèse USTL Montpellier II, Montpellier, France, 241p.
                      18. HEAR http://www.hear.org/pier/species/achyranthes_aspera.htm
                      19. Flora of China http://www.efloras.org/florataxon.aspx?flora_id=2&taxon_id=200006961
                      20. Plants of the World Online https://powo.science.kew.org/taxon/urn:lsid:ipni.org:names:2468-2
                      21. The World Flora Online http://www.worldfloraonline.org/taxon/wfo-0000516177
                      22. Blanfort, V., F. Desmoulins, J. Prosperi, T. Le Bourgeois, R. Guiglion and P. Grard (2010). AdvenPaC V.1.0 : Adventices et plantes à conflit d'intérêt des Pâturages de Nouvelle-Calédonie. Montpellier, France, IAC, Cirad.
                      23. Traoré H., 1991. Influence des facteurs agro-écologiques sur la constitution des communautés adventices des principales cultures céréalières (sorgho, mil, maïs) du Burkina-Faso.
                      24. Barthelat, F. 2019. La Flore illustrée de Mayotte. Meze, Paris, France, Collection Inventaires et Biodiversité, Biotope – Muséum national d’Histoire naturelle. 487 p.
                      25. Huat, J., Nagy, M., Carpente, A., Schwartz, M., Le Bourgeois, T. & Marnotte, P. 2021. Guide de la flore spontannée des agrosystèmes de Mayotte. Montpellier, Cirad. 150 p.
                      26. Fournet, J. 2002. Flore illustrée des phanérogames de Guadeloupe et de Martinique. Montpellier, France, Cirad, Gondwana éditions.
                      27. Swaziland Alien Plants Database
                      28. Blanfort, V., F. Desmoulins, J. Prosperi, T. Le Bourgeois, R. Guiglion and P. Grard (2010). AdvenPaC V.1.0 : Adventices et plantes à conflit d'intérêt des Pâturages de Nouvelle-Calédonie. Montpellier, France, IAC, Cirad
                      29. Le Bourgeois, T., A. Carrara, M. Dodet, W. Dogley, A. Gaungoo, P. Grard, Y. Ibrahim, E. Jeuffrault, G. Lebreton, P. Poilecot, J. Prosperi, J. A. Randriamampianina, A. P. Andrianaivo and F. Théveny (2008). Advent-OI : Principales adventices des îles du sud-ouest de l'Océan Indien. Cirad. Montpellier, France, Cirad.
                      30. US Forest Service, Pacific Island Ecosystems at Risk (PIER)
                      31. Bio-NET EAFRINET
                      32. Australian Tropical Rainforest Plants
                      33. Le Bourgeois, T., P. Grard, L. C. Foxcroft, D. Thompson, A. Carrara, A. Guézou, R. W. Taylor and T. Marshall (2013). Pl@ntInvasive-Kruger V.1.0 : Alien plants of the Kruger National Park. Cdrom. Montpellier, France, Skukuza, South Africa, Cirad-SANparks-SAEON eds.

                      Plantes envahissantes et dégradation des pâturages et des espaces pastoraux en Nouvelle-Calédonie

                      Thomas Le Bourgeois
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                      Thomas Le Bourgeois
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                        WIKTROP - Weed Identification and Knowledge in the Tropical and Mediterranean areasWIKTROP - Weed Identification and Knowledge in the Tropical and Mediterranean areas
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